Sample records for sucker rod string

  1. Modeling and simulation performance of sucker rod beam pump

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Aditsania, Annisa, E-mail: annisaaditsania@gmail.com; Rahmawati, Silvy Dewi, E-mail: silvyarahmawati@gmail.com; Sukarno, Pudjo, E-mail: psukarno@gmail.com

    2015-09-30

    Artificial lift is a mechanism to lift hydrocarbon, generally petroleum, from a well to surface. This is used in the case that the natural pressure from the reservoir has significantly decreased. Sucker rod beam pumping is a method of artificial lift. Sucker rod beam pump is modeled in this research as a function of geometry of the surface part, the size of sucker rod string, and fluid properties. Besides its length, sucker rod string also classified into tapered and un-tapered. At the beginning of this research, for easy modeling, the sucker rod string was assumed as un-tapered. The assumption provedmore » non-realistic to use. Therefore, the tapered sucker rod string modeling needs building. The numerical solution of this sucker rod beam pump model is computed using finite difference method. The numerical result shows that the peak of polished rod load for sucker rod beam pump unit C-456-D-256-120, for non-tapered sucker rod string is 38504.2 lb, while for tapered rod string is 25723.3 lb. For that reason, to avoid the sucker rod string breaks due to the overload, the use of tapered sucker rod beam string is suggested in this research.« less

  2. Robust technology and system for management of sucker rod pumping units in oil wells

    NASA Astrophysics Data System (ADS)

    Aliev, T. A.; Rzayev, A. H.; Guluyev, G. A.; Alizada, T. A.; Rzayeva, N. E.

    2018-01-01

    We propose a technology for calculating the robust, normalized correlation functions of the signal from the force sensor on the rod string attached to the hanger of the sucker rod pumping unit. The robust normalized correlation functions are used to form sets of informative attribute combinations, each of which corresponds to a technical condition of the sucker rod pumping unit. We demonstrate how these sets can be used to solve identification and management problems in the oil production process in real time using inexpensive controllers. The results obtained from using the system on real objects are also presented in this paper. It was determined that the energy saved and prolonged overhaul period substantially increased the cost-effectiveness.

  3. 13. SOUTHEAST TO SUCKER ROD WORK BENCH AND WOODEN SUCKER ...

    Library of Congress Historic Buildings Survey, Historic Engineering Record, Historic Landscapes Survey

    13. SOUTHEAST TO SUCKER ROD WORK BENCH AND WOODEN SUCKER ROD STORAGE RACKS ALONG EAST WALL OF FACTORY INTERIOR. AT THIS BENCH WORKERS RIVETED THREADED WROUGHT IRON CONNECTORS TO THE ENDS OF 20' LONG WOODEN SUCKER RODS (THE RODS WHICH EXTEND DOWNWARD IN THE WELL FROM THE GROUND SURFACE TO PISTON DISPLACEMENT PUMPS WHICH ACTUALLY ELEVATE WATER TO THE SURFACE). ROZNOR HEATER AT THE FAR RIGHT WAS ADDED CIRCA 1960. - Kregel Windmill Company Factory, 1416 Central Avenue, Nebraska City, Otoe County, NE

  4. Deliquification (SIC) of gas wells. Liberal District-Amoco Production Company

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Smalley, R. Jr.

    Various solutions are presented to the problem of deliquefying gas wells to achieve maximum ultimate recovery and avoid premature abandonment. Advantages and disadvantages of each method of deliquefication are discussed. The methods described include blowing up the casing, siphon strings (gas and liquids up tubing, or gas up casing-tubing annulus, and liquids up tubing), gas cycling, compression, bottomhole separators, plunger lift, and sucker rod pumping.

  5. Study on dynamic characteristics of hydraulic pumping unit on offshore platform

    NASA Astrophysics Data System (ADS)

    Chang, Zong-yu; Yu, Yan-qun; Qi, Yao-guang

    2017-12-01

    A new technology of offshore oil rod pumping production is developed for offshore heavy oil recovery. A new type of miniature hydraulic pumping unit with long-stroke, low pumping speed and compact structure is designed based on the spatial characteristics of offshore platforms. By combining the strengths of sinusoidal velocity curve and trapezoidal velocity curve, a kinematical model of the acceleration, the velocity and displacement of the pumping unit's hanging point is established. The results show that the pumping unit has good kinematic characteristics of smooth motion and small dynamic load. The multi-degree-of-freedom dynamic model of the single-well pumping unit is established. The first and second order natural frequencies of the sucker rod string subsystem and the pumping unit subsystem are studied. The results show that the first and the second order natural frequencies among the pumping rod string, pumping unit-platform subsystem and the dynamic excitation have differences over 5 times from each other, indicating that resonance phenomenon will not appear during the operation and the dynamic requirements for field use are met in the system.

  6. Features of electric drive sucker rod pumps for oil production

    NASA Astrophysics Data System (ADS)

    Gizatullin, F. A.; Khakimyanov, M. I.; Khusainov, F. F.

    2018-01-01

    This article is about modes of operation of electric drives of downhole sucker rod pumps. Downhole oil production processes are very energy intensive. Oil fields contain many oil wells; many of them operate in inefficient modes with significant additional losses. Authors propose technical solutions to improve energy performance of a pump unit drives: counterweight balancing, reducing of electric motor power, replacing induction motors with permanent magnet motors, replacing balancer drives with chain drives, using of variable frequency drives.

  7. Production optimization of sucker rod pumping wells producing viscous oil in Boscan field, Venezuela

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Guirados, C.; Sandoval, J.; Rivas, O.

    1995-12-31

    Boscan field is located in the western coast of Maracaibo lake and is operated by Maraven S.A., affiliate of Petroleos de Venezuela S.A. It has 315 active wells, 252 of which are produced with sucker rod pumping. Other artificial lift methods currently applied in this field are hydraulic (piston) pumping (39 wells) and ESP (24 wells). This paper presents the results of the production optimization of two sucker rod pumping wells of Boscan field producing viscous oil. This optimization has been possible due to the development of a new production scheme and the application of system analysis in completion design.more » The new production scheme involves the utilization of a subsurface stuffing box assembly and a slotted housing, both designed and patented by Intevep S.A., affiliate of Petroleos de Venezuela S.A. The completion design method and software used in the optimization study were also developed by Intevep S.A. The new production scheme and design method proved to be effective in preventing the causes of the above mentioned problems, allowing the increase of oil production under better operating conditions.« less

  8. Wireline system for multiple direct push tool usage

    DOEpatents

    Bratton, Wesley L.; Farrington, Stephen P.; Shinn, II, James D.; Nolet, Darren C.

    2003-11-11

    A tool latching and retrieval system allows the deployment and retrieval of a variety of direct push subsurface characterization tools through an embedded rod string during a single penetration without requiring withdrawal of the string from the ground. This enables the in situ interchange of different tools, as well as the rapid retrieval of soil core samples from multiple depths during a single direct push penetration. The system includes specialized rods that make up the rod string, a tool housing which is integral to the rod string, a lock assembly, and several tools which mate to the lock assembly.

  9. 7 CFR 29.2270 - Foreign matter.

    Code of Federal Regulations, 2010 CFR

    2010-01-01

    ... 7 Agriculture 2 2010-01-01 2010-01-01 false Foreign matter. 29.2270 Section 29.2270 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Foreign matter. Any extraneous substance or material such as stalks, suckers, straw, strings, rubber bands...

  10. 7 CFR 29.2270 - Foreign matter.

    Code of Federal Regulations, 2013 CFR

    2013-01-01

    ... 7 Agriculture 2 2013-01-01 2013-01-01 false Foreign matter. 29.2270 Section 29.2270 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Foreign matter. Any extraneous substance or material such as stalks, suckers, straw, strings, rubber bands...

  11. 7 CFR 29.3023 - Foreign matter.

    Code of Federal Regulations, 2011 CFR

    2011-01-01

    ... 7 Agriculture 2 2011-01-01 2011-01-01 false Foreign matter. 29.3023 Section 29.3023 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Foreign matter. Any extraneous substance or material such as stalks, suckers, straw, strings, rubber bands...

  12. 7 CFR 29.2270 - Foreign matter.

    Code of Federal Regulations, 2012 CFR

    2012-01-01

    ... 7 Agriculture 2 2012-01-01 2012-01-01 false Foreign matter. 29.2270 Section 29.2270 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Foreign matter. Any extraneous substance or material such as stalks, suckers, straw, strings, rubber bands...

  13. 7 CFR 29.3023 - Foreign matter.

    Code of Federal Regulations, 2014 CFR

    2014-01-01

    ... 7 Agriculture 2 2014-01-01 2014-01-01 false Foreign matter. 29.3023 Section 29.3023 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Foreign matter. Any extraneous substance or material such as stalks, suckers, straw, strings, rubber bands...

  14. 7 CFR 29.3023 - Foreign matter.

    Code of Federal Regulations, 2013 CFR

    2013-01-01

    ... 7 Agriculture 2 2013-01-01 2013-01-01 false Foreign matter. 29.3023 Section 29.3023 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Foreign matter. Any extraneous substance or material such as stalks, suckers, straw, strings, rubber bands...

  15. 7 CFR 29.3023 - Foreign matter.

    Code of Federal Regulations, 2012 CFR

    2012-01-01

    ... 7 Agriculture 2 2012-01-01 2012-01-01 false Foreign matter. 29.3023 Section 29.3023 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Foreign matter. Any extraneous substance or material such as stalks, suckers, straw, strings, rubber bands...

  16. 7 CFR 29.2270 - Foreign matter.

    Code of Federal Regulations, 2011 CFR

    2011-01-01

    ... 7 Agriculture 2 2011-01-01 2011-01-01 false Foreign matter. 29.2270 Section 29.2270 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Foreign matter. Any extraneous substance or material such as stalks, suckers, straw, strings, rubber bands...

  17. 7 CFR 29.2270 - Foreign matter.

    Code of Federal Regulations, 2014 CFR

    2014-01-01

    ... 7 Agriculture 2 2014-01-01 2014-01-01 false Foreign matter. 29.2270 Section 29.2270 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Foreign matter. Any extraneous substance or material such as stalks, suckers, straw, strings, rubber bands...

  18. 7 CFR 29.3023 - Foreign matter.

    Code of Federal Regulations, 2010 CFR

    2010-01-01

    ... 7 Agriculture 2 2010-01-01 2010-01-01 false Foreign matter. 29.3023 Section 29.3023 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Foreign matter. Any extraneous substance or material such as stalks, suckers, straw, strings, rubber bands...

  19. A simple wave driver

    NASA Astrophysics Data System (ADS)

    Kağan Temiz, Burak; Yavuz, Ahmet

    2015-08-01

    This study was done to develop a simple and inexpensive wave driver that can be used in experiments on string waves. The wave driver was made using a battery-operated toy car, and the apparatus can be used to produce string waves at a fixed frequency. The working principle of the apparatus is as follows: shortly after the car is turned on, the wheel starts to turn at a constant angular speed. A rod that is fixed on the wheel turns at the same constant angular speed, too. A tight string that the wave will be created on is placed at a distance where the rod can touch the string. During each rotation of the wheel, the rod vibrates the string up and down. The vibration frequency of this rod equals the wheel’s rotation frequency, and this frequency value can be measured easily with a small magnet and a bicycle speedometer. In this way, the frequency of the waves formed in the rope can also be measured.

  20. Experience reveals ways to minimize failures in rod-pumped wells

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Patterson, J.C.; Bucaram, S.M.; Curfew, J.V.

    From the experience gained over the past 25 years, ARCO Oil and Gas Co. has developed recommendations to reduce equipment failure in sucker-rod pumping installations. These recommendations include equipment selection and design, operating procedures, and chemical treatment. Equipment failure and its attendant costs are extremely important in today's petroleum industry. Because rod pumping is the predominant means of artificial lift, minimizing equipment failure in rod pumped wells can have a significant impact on profitability. This compilation of recommendations comes from field locations throughout the US and other countries. The goal is to address and solve problems on a well-by-well basis.

  1. Research concerning the evaluation of the connection forces in the joints of the sucker rod pumping units mechanism

    NASA Astrophysics Data System (ADS)

    Bădoiu, D.; Petrescu, M. G.; Antonescu, N. N.; Toma, G.

    2018-01-01

    At present, the sucker rod pumping installations are the most used in the case of the wells in production, when an eruptive exploitation is not possible. The practice has demonstrated that an important role in increasing safety in the operation of the pumping units has the design of the various component bearings because of the extremely high values of the connection forces to which they are loaded. That is why it is necessary to establish as accurately as possible the values of these connecting forces. In the paper is analyzed the dynamics of a conventional pumping unit mechanism. The dynamic study which allows establishing the values of the connecting forces in the joints is performed within the Assur structural groups. The dynamic analysis was implemented into a computer program using Maple programming environment and finally it has been presented some simulation results in the case of a C-320D-256-100 pumping unit.

  2. Model for Sucker-Rod Pumping Unit Operating Modes Analysis Based on SimMechanics Library

    NASA Astrophysics Data System (ADS)

    Zyuzev, A. M.; Bubnov, M. V.

    2018-01-01

    The article provides basic information about the process of a sucker-rod pumping unit (SRPU) model developing by means of SimMechanics library in the MATLAB Simulink environment. The model is designed for the development of a pump productivity optimal management algorithms, sensorless diagnostics of the plunger pump and pumpjack, acquisition of the dynamometer card and determination of a dynamic fluid level in the well, normalization of the faulty unit operation before troubleshooting is performed by staff as well as equilibrium ratio determining by energy indicators and outputting of manual balancing recommendations to achieve optimal power consumption efficiency. Particular attention is given to the application of various blocks from SimMechanics library to take into account the pumpjack construction principal characteristic and to obtain an adequate model. The article explains in depth the developed tools features for collecting and analysis of simulated mechanism data. The conclusions were drawn about practical implementation possibility of the SRPU modelling results and areas for further development of investigation.

  3. Device and method to relieve cordelle action in a chain driven pump

    DOEpatents

    Dysarz, Edward D.

    1994-01-01

    A cordelle action relief apparatus or device for use in sucker rod pumps in a petroleum or water well. The device is incorporated in a chain driven pump to prevent the chain from forming a bow or archlike configuration as the chain rolls off of the sprocket and down into the well. When the chain is allowed to form this bow or arch it could damage the well and well casing. The device includes a first rod on the side of the chain and a second rod on the second side of the chain that will allow the rollers of the chain to roll on the rod and further prevent the chain from bowing or arching and will further allow the rollers on the chain to roll on the rods which will further prevent damage to the well casing, the well, and the chain.

  4. [A new technic for esophago-enteral anastomosis with a mechanical stapler without purse-string sutures].

    PubMed

    Liboni, A; Mari, C; Zamboni, P; Uzzau, A; Noce, L; Bucoliero, F; Mele, M; Masala, C

    1989-01-01

    Staplers have improved the results of esophageal surgery, in our experience and in others experience, as esophago-enteric anastomoses have become safer and faster than when manual suturing is used. Probably one of the last problems in the stapler technique, especially in the thoracic area, is the performance of on adequate esophageal purse-string suture: an improper performance of this suture can cause a dangerous leak of the anastomosis. So, many surgeons, to reduce the risk of esophageal dehiscence connected with the esophageal purse-string, use either purse-string devices or alternative methods such as a second handsewn purse-string, U stitches of the esophagus, etc. We think that the risk of improper anastomoses after esophageal resection can be reduced if the need for the esophageal purse-string can be eliminated. This work shows our personal technique for performing esophagoenterostomy, especially in the thoracic area, using the new CEEA stapler (Autosuture) without esophageal purse-string sutures. According to the modified procedure the stapler anvil and the mini rod are introduced in the esophagectomy and a 2-0 thread is knotted around the CEEA mini rod. Then the esophageal mutilated part is closed by a linear stapler keeping a syringe needle, which contains the thread, through the linear suture. Then, using the thread as a pulling system, the surgeon makes the needle and the tip of the mini rod slide out of the esophageal suture. Now the surgeon can reassemble the CEEA and perform the anastomosis. There are many clinical reports that cite no leaks following circular stapled anastomoses across linear stapled closures.

  5. Hierarchical Nanostructures Self-Assembled from a Mixture System Containing Rod-Coil Block Copolymers and Rigid Homopolymers

    PubMed Central

    Li, Yongliang; Jiang, Tao; Lin, Shaoliang; Lin, Jiaping; Cai, Chunhua; Zhu, Xingyu

    2015-01-01

    Self-assembly behavior of a mixture system containing rod-coil block copolymers and rigid homopolymers was investigated by using Brownian dynamics simulations. The morphologies of formed hierarchical self-assemblies were found to be dependent on the Lennard-Jones (LJ) interaction εRR between rod blocks, lengths of rod and coil blocks in copolymer, and mixture ratio of block copolymers to homopolymers. As the εRR value decreases, the self-assembled structures of mixtures are transformed from an abacus-like structure to a helical structure, to a plain fiber, and finally are broken into unimers. The order parameter of rod blocks was calculated to confirm the structure transition. Through varying the length of rod and coil blocks, the regions of thermodynamic stability of abacus, helix, plain fiber, and unimers were mapped. Moreover, it was discovered that two levels of rod block ordering exist in the helices. The block copolymers are helically wrapped on the homopolymer bundles to form helical string, while the rod blocks are twistingly packed inside the string. In addition, the simulation results are in good agreement with experimental observations. The present work reveals the mechanism behind the formation of helical (experimentally super-helical) structures and may provide useful information for design and preparation of the complex structures. PMID:25965726

  6. A modified efficient purse-string stapling technique (mEST) that uses a new metal rod for intracorporeal esophagojejunostomy in laparoscopic total gastrectomy

    PubMed Central

    Moon, Jeong-Ho; Yamamoto, Kazuyoshi; Yanagimoto, Yoshitomo; Sugimura, Keijirou; Miyata, Hiroshi; Yano, Masahiko; Sakon, Masato

    2017-01-01

    Intracorporeal esophagojejunostomy after laparoscopic total gastrectomy is technically difficult because this procedure should be performed in a narrow surgical field in the upper abdomen even when completely laparoscopic approaches are used. The placement of the anvil of a circular stapling device into the esophagus and connection the instrument to the anvil are extremely difficult steps in this surgery. Therefore, we developed a simple technique for intracorporeal esophagojejunostomy using hemi-double stapling technique; we named this technique the efficient purse-string stapling technique (EST). More recently, we have developed a modified EST (mEST) that utilizes a new stainless steel anvil rod instead of a plastic rod. Relative to the plastic rod, the steel rod is reusable and shorter; thus, it was easier to perform anvil placement into the esophagus with the steel rod. Anvil preparation for mEST: a stainless steel rod is attached to the shaft of the anvil, and the needle and thread are sutured to the tip of the rod. After complete insertion of the anvil into the esophageal cavity, the needle and thread are used to penetrate the anterior esophageal wall, and the esophagus is then clamped using a linear stapler just distal to the site penetrated by the thread. The linear stapler is fired, and anvil placement in the esophagus is simultaneously accomplished. After the rod is removed from the anvil, the instrument is intracorporeally connected to the anvil and then fired to complete the gastrojejunostomy. This technique is simple and facilitates intracorporeal reconstruction procedures in laparoscopic total gastrectomy. PMID:28815221

  7. Lunar drill footplate and casing

    NASA Technical Reports Server (NTRS)

    Maassen, Erik C.; Hendrix, Thomas H.; Morrison, Eddie W.; Phillips, Rodrick B.; Le, Vu Quang; Works, Bruce A.

    1989-01-01

    To prevent hole collapse during lunar drilling operations, a casing has been devised of a graphite reinforced polyimide composite which will be able to withstand the lunar environment. Additionally, this casing will be inserted into the ground in segments two meters long which will penetrate the regolith simultaneously with the auger. The vertical action of the mobile platform will provide a downward force to the casing string through a special adaptor, giving the casing the needed impetus to sink the anticipated depth of ten meters. Casing segments will be connected with a simple snap arrangement. Excess casing will be cut off by a cylindrical cutting tool which will also transport the excess casing away from the hole. A footplate will be incorporated to grasp the auger rod string during rod segment additions or removals. The footplate grasping mechanism will consist of a set of vice-like arms, one end of each bearing threaded to a common power screw. The power screw will be threaded such that one end's thread pitch opposes that of the other end. The weight of the auger and rod string will be transmitted through the arms to the power screw and absorbed by a set of three ball bearing assemblies. The power screw will be driven by a one-half horsepower brushless motor actuated by radio control. The footplate will rest on four short legs and be anchored with pins that are an integral part of each leg.

  8. Simple Excitation of Standing Waves in Rubber Bands and Membranes

    NASA Astrophysics Data System (ADS)

    Cortel, Adolf

    2004-04-01

    Many methods to excite standing waves in strings, plates, membranes, rods, tubes, and soap bubbles have been described. Usually a loudspeaker or a vibrating reed is driven by the amplified output of an audio oscillator. A novel and simple method consists of using a tuning fork or a singing rod to excite transversal standing waves in stretched rubber membranes sprinkled with fine sand.

  9. One-dimensional reduction of viscous jets. II. Applications

    NASA Astrophysics Data System (ADS)

    Pitrou, Cyril

    2018-04-01

    In a companion paper [Phys. Rev. E 97, 043115 (2018), 10.1103/PhysRevE.97.043115], a formalism allowing to describe viscous fibers as one-dimensional objects was developed. We apply it to the special case of a viscous fluid torus. This allows to highlight the differences with the basic viscous string model and with its viscous rod model extension. In particular, an elliptic deformation of the torus section appears because of surface tension effects, and this cannot be described by viscous string nor viscous rod models. Furthermore, we study the Rayleigh-Plateau instability for periodic deformations around the perfect torus, and we show that the instability is not sufficient to lead to the torus breakup in several droplets before it collapses to a single spherical drop. Conversely, a rotating torus is dynamically attracted toward a stationary solution, around which the instability can develop freely and split the torus in multiple droplets.

  10. Inter-annual variability in apparent relative production, survival, and growth of juvenile Lost River and shortnose suckers in Upper Klamath Lake, Oregon, 2001–15

    USGS Publications Warehouse

    Burdick, Summer M.; Martin, Barbara A.

    2017-06-15

    Executive SummaryPopulations of the once abundant Lost River (Deltistes luxatus) and shortnose suckers (Chasmistes brevirostris) of the Upper Klamath Basin, decreased so substantially throughout the 20th century that they were listed under the Endangered Species Act in 1988. Major landscape alterations, deterioration of water quality, and competition with and predation by exotic species are listed as primary causes of the decreases in populations. Upper Klamath Lake populations are decreasing because fish lost due to adult mortality, which is relatively low for adult Lost River suckers and variable for adult shortnose suckers, are not replaced by new young adult suckers recruiting into known adult spawning aggregations. Catch-at-age and size data indicate that most adult suckers presently in Upper Klamath Lake spawning populations were hatched around 1991. While, a lack of egg production and emigration of young fish (especially larvae) may contribute, catch-at-length and age data indicate high mortality during the first summer or winter of life may be the primary limitation to the recruitment of young adults. The causes of juvenile sucker mortality are unknown.We compiled and analyzed catch, length, age, and species data on juvenile suckers from Upper Klamath Lake from eight prior studies conducted from 2001 to 2015 to examine annual variation in apparent production, survival, and growth of young suckers. We used a combination of qualitative assessments, general linear models, and linear regression to make inferences about annual differences in juvenile sucker dynamics. The intent of this exercise is to provide information that can be compared to annual variability in environmental conditions with the hopes of understanding what drives juvenile sucker population dynamics.Age-0 Lost River suckers generally grew faster than age-0 shortnose suckers, but the difference in growth rates between the two species varied among years. This unsynchronized annual variation in daily growth may be an indication that environmental conditions are affecting growth rates of these species in different ways.The combined evidence outlined in this report and in Simon and others (2012) indicates that years of relatively high age-0 sucker production occurred in the late 1990s through at least 2000, in 2006, and in 2011. Our analysis of annual age-0 sucker catch per unit effort (CPUE), which accounted for zero inflated data and annual variation in sampling gears and locations, indicated that 2006 had the greatest apparent relative production of age-0 suckers ≥ 45 mm standard length (SL) during the time period examined. Midsummer trap net effort by the U.S. Geological Survey (USGS) was too sparse to examine age-0 sucker CPUE from 2011 to 2013. Relatively frequent catches of age-1 suckers in 2001, 2007, and 2012 corroborated relatively high CPUE for age-0 suckers during 1999–2000, 2006, and 2011, as reported by USGS or Simon and others (2012).There were several indications in the data that juvenile sucker survival is low from at least midsummer of the first year of life through mid-September of the second year of life. Our estimated index of relative apparent age-0 sucker late-summer survival, which accounted for zero inflated data and variations in sampling gears and locations, was higher in 2009 than in 2004. Our index of apparent age-0 sucker mortality for all other years from 2001 to 2015 was similar among years. Seventy-five percent of age-1 suckers were captured prior to July 17 each year. In 2007, the one year with substantial age-1 sucker summertime catches, the proportion of nets to capture age-1 suckers decreased from July to mid-September. Maximum annual age-2+ sucker CPUE was 0.02 fish per net, 10,000 times less than the maximum annual age-0 sucker CPUE.Analysis of species data indicated that juvenile Lost River suckers may have greater apparent mortality than shortnose suckers. Lost River suckers made up a smaller proportion of age-0 suckers captured in July each year than would be expected, based on the abundance of adult Lost River suckers relative to shortnose suckers, and higher Lost River than shortnose sucker fecundity. The proportion of age-0 suckers captured that were Lost River suckers decreased from July to September in several years. Only 14 percent of age-1 or older juvenile suckers identified to species over the 15-year time period were Lost River suckers.

  11. Spring and Summer Spatial Distribution of Endangered Juvenile Lost River and Shortnose Suckers in Relation to Environmental Variables in Upper Klamath Lake, Oregon: 2007 Annual Report

    USGS Publications Warehouse

    Burdick, Summer M.; VanderKooi, Scott P.; Anderson, Greer O.

    2009-01-01

    Lost River sucker Deltistes luxatus and shortnose sucker Chasmistes brevirostris were listed as endangered in 1988 for a variety of reasons including apparent recruitment failure. Upper Klamath Lake, Oregon, and its tributaries are considered the most critical remaining habitat for these two species. Age-0 suckers are often abundant in Upper Klamath Lake throughout the summer months, but catches decline dramatically between late August and early September each year, and age-1 and older subadult suckers are rare. These rapid declines in catch rates and a lack of substantial recruitment into adult sucker populations in recent years suggests sucker populations experience high mortality between their first summer and first spawn. A lack of optimal rearing habitat may exacerbate juvenile sucker mortality or restrict juvenile growth or development. In 2007, we continued research on juvenile sucker habitat use begun by the U.S. Geological Survey (USGS) in 2001. Age-0 catch rates in 2006 were more than an order of magnitude greater than in previous years, which prompted us to refocus our research from age-0 suckers to age-1 sucker distributions and habitat use. We took a two-phased approach to our research in 2007 that included preliminary spring sampling and intense summer sampling components. Spring sampling was a pilot study designed to gather baseline data on the distribution of age-1 suckers as they emerge from winter in shoreline environments throughout Upper Klamath Lake (Chapter 1). Whereas, summer sampling was designed to quantitatively estimate the influence of environmental variables on age-0 and age-1 sucker distribution throughout Upper Klamath Lake, while accounting for imperfect detection (Chapter 2). In addition to these two components, we began a project to evaluate passive integrated transponder (PIT) tag loss and the effects of PIT tags on mortality of age-1 Lost River suckers (Chapter 3). The spring pilot study built the foundation for future research on post-wintering juvenile sucker distribution and habitat use studies. Only 34 percent of nets set during spring sampling (April 2 to May 29) caught juvenile suckers and catch rates were low (0.038 to 0.405 suckers/hour) and widely distributed throughout shoreline areas. Of 13 suckers sacrificed for identification, only one was determined to be a Lost River sucker. All others were either shortnose suckers or Klamath largescale Catostomus snyderi suckers, but were not identified to species. Suckers caught during the spring averaged 93 +- 2 millimeter (mm) standard length (SL; mean +- SE) and were all estimated to be a year old. Spring catches did not vary in respect to nearness to tributary streams or rivers, substrate type, area of the lake, or distance from shore. On the other hand, a higher percentage of nets caught at least one sucker when they were set within 50 meters (m) of a wetland edge (60 percent) compared to nets set 200 m from a wetland (30 percent) or in other shoreline areas (29 percent). Our results also suggest that in the spring age-1 suckers use habitats less than 2 m deep at a greater frequency than deeper environments, a trend that was reversed in the summer. Temporal trends in summer catch rates of age-0 suckers generally were similar to those in previous years, with a peak during the week of August 5. In contrast, age-1 sucker catches were relatively high until the week of July 16, but rapidly declined each week for the rest of the sampling season. Age-0 suckers were caught at higher rates than age-1 suckers though the summer, but both age groups were captured at a similar percentage of sites (age-0, 26.5 percent and age-1, 27.4 percent). Age-0 catches were composed of slightly more Lost River suckers (53.2 percent) than shortnose suckers (42.1 percent). In contrast, most age-1 suckers were shortnose suckers (72.7 percent). Our summer sampling indicates age-0 suckers within Upper Klamath Lake primarily are habitat generalists, whe

  12. Colonial waterbird predation on Lost River and Shortnose suckers in the Upper Klamath Basin

    USGS Publications Warehouse

    Evans, Allen F.; Hewitt, David A.; Payton, Quinn; Cramer, Bradley M.; Collis, Ken; Roby, Daniel D.

    2016-01-01

    We evaluated predation on Lost River Suckers Deltistes luxatus and Shortnose Suckers Chasmistes brevirostris by American white pelicans Pelecanus erythrorhynchos and double-crested cormorants Phalacrocorax auritus nesting at mixed-species colonies in the Upper Klamath Basin of Oregon and California during 2009–2014. Predation was evaluated by recovering (detecting) PIT tags from tagged fish on bird colonies and calculating minimum predation rates, as the percentage of available suckers consumed, adjusted for PIT tag detection probabilities but not deposition probabilities (i.e., probability an egested tag was deposited on- or off-colony). Results indicate that impacts of avian predation varied by sucker species, age-class (adult, juvenile), bird colony location, and year, demonstrating dynamic predator–prey interactions. Tagged suckers ranging in size from 72 to 730 mm were susceptible to cormorant or pelican predation; all but the largest Lost River Suckers were susceptible to bird predation. Minimum predation rate estimates ranged annually from <0.1% to 4.6% of the available PIT-tagged Lost River Suckers and from <0.1% to 4.2% of the available Shortnose Suckers, and predation rates were consistently higher on suckers in Clear Lake Reservoir, California, than on suckers in Upper Klamath Lake, Oregon. There was evidence that bird predation on juvenile suckers (species unknown) in Upper Klamath Lake was higher than on adult suckers in Upper Klamath Lake, where minimum predation rates ranged annually from 5.7% to 8.4% of available juveniles. Results suggest that avian predation is a factor limiting the recovery of populations of Lost River and Shortnose suckers, particularly juvenile suckers in Upper Klamath Lake and adult suckers in Clear Lake Reservoir. Additional research is needed to measure predator-specific PIT tag deposition probabilities (which, based on other published studies, could increase predation rates presented herein by a factor of roughly 2.0) and to better understand biotic and abiotic factors that regulate sucker susceptibility to bird predation.

  13. Performance and scaling of a novel locomotor structure: adhesive capacity of climbing gobiid fishes.

    PubMed

    Maie, Takashi; Schoenfuss, Heiko L; Blob, Richard W

    2012-11-15

    Many species of gobiid fishes adhere to surfaces using a sucker formed from fusion of the pelvic fins. Juveniles of many amphidromous species use this pelvic sucker to scale waterfalls during migrations to upstream habitats after an oceanic larval phase. However, adults may still use suckers to re-scale waterfalls if displaced. If attachment force is proportional to sucker area and if growth of the sucker is isometric, then increases in the forces that climbing fish must resist might outpace adhesive capacity, causing climbing performance to decline through ontogeny. To test for such trends, we measured pressure differentials and adhesive suction forces generated by the pelvic sucker across wide size ranges in six goby species, including climbing and non-climbing taxa. Suction was achieved via two distinct growth strategies: (1) small suckers with isometric (or negatively allometric) scaling among climbing gobies and (2) large suckers with positively allometric growth in non-climbing gobies. Species using the first strategy show a high baseline of adhesive capacity that may aid climbing performance throughout ontogeny, with pressure differentials and suction forces much greater than expected if adhesion were a passive function of sucker area. In contrast, large suckers possessed by non-climbing species may help compensate for reduced pressure differentials, thereby producing suction sufficient to support body weight. Climbing Sicyopterus species also use oral suckers during climbing waterfalls, and these exhibited scaling patterns similar to those for pelvic suckers. However, oral suction force was considerably lower than that for pelvic suckers, reducing the ability for these fish to attach to substrates by the oral sucker alone.

  14. Distribution and condition of larval and juvenile Lost River and shortnose suckers in the Williamson River Delta restoration project and Upper Klamath Lake, Oregon

    USGS Publications Warehouse

    Burdick, Summer M.; Brown, Daniel T.

    2010-01-01

    Federally endangered Lost River sucker (Deltistes luxatus) and shortnose sucker (Chasmistes brevirostris) were once abundant throughout their range but populations have declined. They were extirpated from several lakes in the 1920s and may no longer reproduce in others. Poor recruitment to the adult spawning populations is one of several reasons cited for the decline and lack of recovery of these species and may be the consequence of high mortality during juvenile life stages. High larval and juvenile sucker mortality may be exacerbated by an insufficient quantity of suitable or high quality rearing habitat. In addition, larval suckers may be swept downstream from suitable rearing areas in Upper Klamath Lake into Keno Reservoir, which is seasonally anoxic. The Nature Conservancy flooded about 3,600 acres (1,456 hectares) to the north of the Williamson River mouth (Tulana Unit) in October 2007 and about 1,400 acres (567 hectares) to the south and east of the Williamson River mouth (Goose Bay Unit) a year later to retain larval suckers in Upper Klamath Lake, create nursery habitat, and improve water quality. The U.S. Geological Survey joined a long-term research and monitoring program in collaboration with The Nature Conservancy, the Bureau of Reclamation, and Oregon State University in 2008 to assess the effects of the Williamson River Delta restoration on the early life-history stages of Lost River and shortnose suckers. The primary objectives of the research were to describe habitat colonization and use by larval and juvenile suckers and non-sucker fishes and to evaluate the effects of the restored habitat on the health and condition of juvenile suckers. This report summarizes data collected in 2009 by the U.S. Geological Survey as a part of this monitoring effort. The Williamson River Delta appeared to provide suitable rearing habitat for endangered larval Lost River and shortnose suckers in 2008 and 2009. Larval suckers captured in this delta typically were larger than those captured in the adjacent lake habitat in 2008, but the opposite was true for larval shortnose suckers in 2009. Mean sample density was greater for both species in the Williamson River Delta than adjacent lake habitats in both years. Larval suckers captured in the restoration area, however, had less food in their guts compared to those captured in Upper Klamath or Agency Lakes. Differential distribution among sucker species within the Williamson River Delta and between the delta and adjacent lakes indicated that shortnose suckers likely benefited more from the restored Williamson River Delta than Lost River or Klamath largescale suckers (Catostomus snyderi). Catch rates in shallow-water habitats with vegetation within the delta were higher for shortnose and Klamath largescale suckers than for larval Lost River suckers in 2008 and 2009.However, catch rates at the mouth of the Williamson River in 2008 and in Upper Klamath Lake in 2009 were higher for larval Lost River suckers than for larvae identified as either shortnose or Klamath largescale suckers. Shortnose suckers also comprised the greatest portion of age-0 suckers captured in the Williamson River Delta in 2008 and 2009. The relative abundance of age-1 shortnose suckers was high in our catches compared to age-1 Lost River suckers in 2009 in the delta and adjacent lakes, which may or may not indicate shortnose suckers experienced better survival than Lost River suckers in 2008. Age-0 and age-1 suckers were similarly distributed throughout the Williamson River Delta in 2008 and 2009. Age-0 suckers used shallow vegetated and unvegetated habitats primarily in mid- to late July in both years. A comparison of catch rates between our study and a concurrent study in Upper Klamath Lake indicated that Goose Bay was the most used habitat in 2009 and the Tulana Unit was the one of the least used habitats in 2008 and 2009 by age-0 suckers. Catch rates for age-1 suckers, however, indicated that bo

  15. Population characteristics and the influence of discharge on Bluehead Sucker and Flannelmouth Sucker

    USGS Publications Warehouse

    Klein, Zachary B.; Breen, Matthew J.; Quist, Michael C.

    2017-01-01

    Rivers are among some of the most complex and important ecosystems in the world. Unfortunately, many fishes endemic to rivers have suffered declines in abundance and distribution suggesting that alterations to lotic environments have negatively influenced native fish populations. Of the 35 fishes native to the Colorado River basin (CRB), seven are considered either endangered, threatened, or species of special concern. As such, the conservation of fishes native to the CRB is a primary interest for natural resource management agencies. One of the major factors limiting the conservation and management of fishes endemic to the CRB is the lack of basic information on their ecology and population characteristics. We sought to describe the population dynamics and demographics of three populations of Bluehead Suckers (Catostomus discobolus) and Flannelmouth Suckers (C. latipinnis) in Utah. Additionally, we evaluated the potential influence of altered flow regimes on the recruitment and growth of Bluehead Suckers and Flannelmouth Suckers. Mortality of Bluehead Suckers and Flannelmouth Suckers from the Green, Strawberry, and White rivers was comparable to other populations. Growth of Bluehead Suckers and Flannelmouth Suckers was higher in the Green, Strawberry, and White rivers when compared to other populations in the CRB. Similarly, recruitment indices suggested that Bluehead Suckers and Flannelmouth Suckers in the Green, Strawberry, and White rivers had more stable recruitment than other populations in the CRB. Models relating growth and recruitment to hydrological indices provided little explanatory power. Notwithstanding, our results indicate that Bluehead Suckers and Flannelmouth Suckers in the Green, Strawberry, and White rivers represent fairly stable populations and provide baseline information that will be valuable for the effective management and conservation of the species.

  16. Temporal and spatial distribution of endangered juvenile Lost River and shortnose suckers in relation to environmental variables in Upper Klamath Lake, Oregon: 2009 annual data summary

    USGS Publications Warehouse

    Bottcher, Jared L.; Burdick, Summer M.

    2010-01-01

    Lost River sucker (Deltistes luxatus) and shortnose sucker (Chasmistes brevirostris) were listed as endangered in 1988 for a variety of reasons including apparent recruitment failure. Upper Klamath Lake, Oregon, and its tributaries are considered the most critical remaining habitat for these two species. Age-0 suckers are often abundant in Upper Klamath Lake throughout the summer months, but catches decline dramatically between late August and early September each year. Similar declines of age-1 suckers between spring and late summer also occur annually. These rapid declines in catch rates and a lack of substantial recruitment into adult sucker populations in recent years suggests sucker populations experience high mortality between their first summer and first spawn. Summer age-0 sucker habitat use and distribution have been studied extensively, but many uncertainties remain about age-1 and older juvenile habitat use, distribution, and movement patterns within Upper Klamath Lake. This study was designed to examine seasonal changes in distribution of age-1 suckers in Upper Klamath Lake as they relate to depth and water quality. The results of our third annual spring and summer sampling effort are presented in this report. Catch data collected in 2009 indicate seasonal changes in age-1 and older juvenile sucker habitat use coincident with changes in water quality. Although age-1 sucker catch rates were again concentrated along the western shore in June and early July, as they were in 2007 and 2008, very few age-1 suckers were captured in Eagle Ridge Trench in 2009 - a deepwater area along the western shore extending from Howard Bay to Eagle Ridge Point. Instead, suckers in 2009 were concentrated in the relatively shallow bays along the western shore. Nevertheless, as dissolved-oxygen concentrations decreased in mid-July below sublethal thresholds around the Eagle Ridge Trench, age-1 suckers apparently moved away from the western shore, and subsequently were captured in main lake areas and the eastern shore. Age-1 suckers were noticeably absent from the tributaries of Upper Klamath Lake during periods of chronically low dissolved-oxygen concentrations in the lake, refuting a previously untested hypothesis that tributaries were important age-1 sucker refuge habitats. In addition, declines in overall catch rates for age-1 suckers in August and September, despite intensive sampling, indicates that the apparent declines in abundance may be due to increased mortality and not due to sampling the wrong environments or poor detection probability. The remote detection of an age-1 juvenile sucker tagged in Short Creek and subsequently recaptured in the Link River array, more than 30 kilometers away, indicates the capacity of juvenile suckers to migrate relatively long distances. This knowledge, coupled with other remotely detected suckers in the Williamson River, indicates that juvenile sucker movement in Upper Klamath Lake may be common. In order to better quantify movement and potentially survival, future research should focus on tagging more juvenile suckers and taking advantage of the significant passive integrated transponder tag infrastructure throughout Upper Klamath Lake and its tributaries. In this data summary, we also describe the distribution of age-0 suckers in Upper Klamath Lake and its tributaries. These data corroborate findings from 2007 and 2008, which describe age-0 sucker habitat as shallow relative to depths available in Upper Klamath Lake. Similar to age-1 suckers, age-0 sucker abundances also appeared to decline in late summer, despite continued sampling throughout Upper Klamath Lake and its tributaries. In addition to low dissolved-oxygen concentrations, increased opercle deformity and anchor worm (Lernaea spp.) infection rates, as well as increased abundances of fathead minnows and other piscivorous non-native fish, may provide potential insight into the causes of juvenile sucker rarity. Opercle deformity r

  17. Tributary use by imperiled Flannelmouth and Bluehead Suckers in the upper Colorado River Basin

    USGS Publications Warehouse

    Fraser, Gregory S.; Winkelman, Dana L.; Bestgen, Kevin R.; Thompson, Kevin G.

    2017-01-01

    Habitat alterations and establishment of nonnative fishes have reduced the distributions of Flannelmouth Sucker Catostomus latipinnis and Bluehead Sucker C. discobolus to less than 50% of their historical ranges in the Colorado River basin. Tributaries are sometimes less altered than main-stem habitat in the basin and may be important to support various life history processes, but their role in the maintenance of Flannelmouth Sucker and Bluehead Sucker populations is poorly understood. Using mark–recapture techniques, we show tributaries are important habitat for native suckers in the upper Colorado River basin and report three main findings. First, both Flannelmouth and Bluehead suckers likely respond to a thermal cue that initiates spawning movement patterns. Suckers moved into Coal Creek from the White River beginning in mid-May of 2012 and 2013 to spawn. The majority of sucker spawning movements occurred when water temperatures in White River exceeded 11–14°C and those in Coal Creek were 2.5–4°C warmer, while flows varied between years. Second, based on PIT tag detection arrays, 13–45% of suckers showed spawning site fidelity. Sampling only with fyke nets would have resulted in the conclusion that site fidelity by native suckers was only 1–17%, because nets were less efficient at detecting marked fish. Third, most suckers of both species emigrated from Coal Creek within 48 h after being captured while suckers that were detected only via arrays remained resident for 10–12 d. The posthandling flight response we observed was not anticipated and to our knowledge has not been previously reported for these species. Remote PIT tag antenna arrays allowed for a stronger inference regarding movement and tributary use by these species than what could be achieved using just fyke nets. Tributaries are an important part of Flannelmouth Sucker and Bluehead Sucker life history and thus important to conservation strategies for these species.

  18. Juvenile Lost River and shortnose sucker year class strength, survival, and growth in Upper Klamath Lake, Oregon, and Clear Lake Reservoir, California—2016 Monitoring Report

    USGS Publications Warehouse

    Burdick, Summer M.; Ostberg, Carl O.; Hoy, Marshal S.

    2018-04-20

    Executive SummaryThe largest populations of federally endangered Lost River (Deltistes luxatus) and shortnose suckers (Chasmistes brevirostris) exist in Upper Klamath Lake, Oregon, and Clear Lake Reservoir, California. Upper Klamath Lake populations are decreasing because adult mortality, which is relatively low, is not being balanced by recruitment of young adult suckers into known spawning aggregations. Most Upper Klamath Lake juvenile sucker mortality appears to occur within the first year of life. Annual production of juvenile suckers in Clear Lake Reservoir appears to be highly variable and may not occur at all in very dry years. However, juvenile sucker survival is much higher in Clear Lake, with non-trivial numbers of suckers surviving to join spawning aggregations. Long-term monitoring of juvenile sucker populations is needed to (1) determine if there are annual and species-specific differences in production, survival, and growth, (2) to identify the season (summer or winter) in which most mortality occurs, and (3) to help identify potential causes of high juvenile sucker mortality, particularly in Upper Klamath Lake.We initiated an annual juvenile sucker monitoring program in 2015 to track cohorts in 3 months (June, August, and September) annually in Upper Klamath Lake and Clear Lake Reservoir. We tracked annual variability in age-0 sucker apparent production, juvenile sucker apparent survival, and apparent growth. Using genetic markers, we were able to classify suckers as one of three taxa: shortnose or Klamath largescale suckers, Lost River, or suckers with genetic markers of both species (Intermediate Prob[LRS]). Using catch data, we generated taxa-specific indices of year class strength, August–September apparent survival, and overwinter apparent survival. We also examined prevalence and severity of afflictions such as parasites, wounds, and deformities.Indices of year class strength in Upper Klamath Lake were similar for shortnose suckers in 2015 and 2016, but about twice as high for Lost River suckers and suckers having intermediate Prob[LRS] in 2016 than in 2015. Indices of apparent August–September survival were lower in 2016 (0.41) than in 2015 (1.07) for shortnose suckers and suckers identified as having intermediate Prob [LRS] (0.14 in 2016 and 1.69 in 2015). Indices of apparent August—September survival were similar in 2016 (0.16) and 2015 (0.07) for Lost River suckers. Indices of apparent survival were lower for age-0 Lost River suckers than age-0 shortnose suckers in both years. Although samples sizes are small, a declining trend in the ratio of Lost River to shortnose suckers from 28/23 (1.22) as age-0 fish in September of 2015 to 1/9 (0.11) as age-1 fish in June of 2016 is consistent with higher over winter apparent mortality for Lost River suckers than shortnose suckers in Upper Klamath Lake.Shortnose sucker year class strength was greater in years with high Willow Creek inflows and Clear Lake surface elevation during the spawning season, indicating that access to spawning habitat was an important contributing factor. In previous sampling, age-0 sucker catch per unit effort (CPUE) was relatively high in 2011 and 2012, moderately high in 2013, and zero in 2014 and 2015. The 2011 and 2012 year classes continued to be detected, but the 2013 year class went undetected for the first time in 2016. The 2014 year class continued to be undetected in 2016. Three suckers with one annulus each on fin rays were captured in Clear Lake in 2016. Although these fish are potential representatives of the 2015 year class, they were small for their age, indicating they may have hatched in 2016. Age-0 shortnose and Lost River suckers were captured in Clear Lake in 2016, indicating new cohorts of both taxa were produced. Moderate to abundant year classes were produced in 2011, 2012, and 2016 when lake surface elevation greater than 1,378.9 m (4,524 ft) during the February–June spawning season. Also in 2011 and 2016, rapid increases in lake-surface elevation indicated potentially high Willow Creek inflows. A somewhat less abundant year class produced in 2012 than in 2011 and 2016 was associated with lower spawning season inflows. The apparently smaller 2013 year class was formed when Willow Creek inflows were apparently low and lake surface never exceeded 1,379.2 m (4,524.9 ft). In 2014 and 2015, when year-classes were small or not detected, the Clear Lake surface elevations were at or below 1,378.2 m (4,522 ft), and there was very little spring time Willow Creek inflow.Age-0 shortnose sucker CPUE in Clear Lake was correlated with seasonal decreases in water volumes in 2016 and could not be used to create indices of August–September survival. Age-0 shortnose sucker catch rates in Clear Lake Reservoir were about seven times less in August than in September. Meanwhile, the water volume in Clear Lake Reservoir declined by about 36 percent between these two sampling periods. Higher September catch rates may have resulted from additional age-0 suckers entering the lake from the river, a concentrating effect of declining water volumes, or both.Differences in August standard length, apparent growth rates, and the prevalence of abnormalities were consistent with healthier age-0 suckers in Clear Lake Reservoir than in Upper Klamath Lake. Age-0 suckers were larger in August in Clear Lake Reservoir than in Upper Klamath Lake, which may be due to an earlier hatch date, faster growth, or both in Clear Lake Reservoir. Sample sizes were only large enough to compare growth rates of age-0 shortnose suckers from Upper Klamath Lake in 2015 to Clear Lake Reservoir in 2016. Age-0 shortnose suckers grew more between August and September in Clear Lake Reservoir in 2016 than in Upper Klamath Lake in 2015. Petechial hemorrhages of the skin on age-0 suckers were more prevalent in Upper Klamath Lake than in Clear Lake Reservoir in 2016. Deformed opercula, black-spot forming parasites, and infections presumed to be Columnaris sp. were observed on less than 12 percent of suckers from Upper Klamath Lake but were not observed on suckers from Clear Lake Reservoir in 2016.

  19. Age and condition of juvenile catostomids in Clear Lake Reservoir, California

    USGS Publications Warehouse

    Burdick, Summer M.; Rasmussen, Josh

    2013-01-01

    Although infrequent recruitment of new individuals into the adult spawning populations of Lost River suckers (Deltistes luxatus) and shortnose suckers (Chasmistes brevirostris) limits recovery of these species in Upper Klamath Lake, it is not clear that populations are recruitment limited in Clear Lake Reservoir (hereafter Clear Lake). Specifically, some evidence indicates that shortnose suckers may regularly recruit to the adult spawning population in Clear Lake. Therefore, a study of early life history patterns and recruitment dynamics in Clear Lake may lead to a better understanding of what is limiting recovery of suckers in both lakes. Adult suckers in Clear Lake migrate up Willow Creek and its tributaries to spawn in some years, but low flow in Willow Creek may inhibit spawning migrations in other years. It is unclear whether spawning is successful, larvae survive, or how frequently juveniles persist to adulthood. Environmental variables associated with successful spawning or young-of-year survival have not been identified, and early life history for these populations is poorly understood. The U.S. Geological Survey, in cooperation with the U.S. Fish and Wildlife Service, initiated a study in 2011 to better understand juvenile sucker life history in Clear Lake, and to identify constraints in the early life history that may limit recruitment to the adult spawning populations. The relative weights of shortnose suckers from Clear Lake and Upper Klamath Lake were compared to examine differences in condition. However, it is unclear whether the disparity in relative weights between the populations reflects differences in condition, phenotype, or both. Approximately 80 percent of juvenile suckers in Clear Lake are shortnose suckers with some morphologic features similar to Klamath largescale suckers (Catostomus snyderi), whereas juvenile suckers in Upper Klamath Lake can be clearly classified as either shortnose or Lost River suckers. The presence of juvenile suckers age-3 and older indicate that production, larval survival, and juvenile survival are at least periodically sufficient to lead to recruitment into the adult population of shortnose suckers in Clear Lake.

  20. Distribution and condition of larval and juvenile Lost River and shortnose suckers in the Williamson River Delta restoration project and Upper Klamath Lake, Oregon

    USGS Publications Warehouse

    Burdick, Summer M.

    2012-01-01

    Federally endangered Lost River sucker (Deltistes luxatus) and shortnose sucker (Chasmistes brevirostris) were once abundant throughout their range but populations have declined. They were extirpated from several lakes in the 1920s and may no longer reproduce in other lakes. Poor recruitment to the adult spawning populations is one of several reasons cited for the decline and lack of recovery of these species and may be the consequence of high mortality during juvenile life stages. High larval and juvenile sucker mortality may be exacerbated by an insufficient quantity of suitable or high-quality rearing habitat. In addition, larval suckers may be swept downstream from suitable rearing areas in Upper Klamath Lake into Keno Reservoir, where they are assumed lost to Upper Klamath Lake populations. The Nature Conservancy flooded about 3,600 acres (1,456 hectares) to the north of the Williamson River mouth (Tulana) in October 2007, and about 1,400 acres (567 hectares) to the south and east of the Williamson River mouth (Goose Bay Farms) in October 2008, in order to retain larval suckers in Upper Klamath Lake, create nursery habitat, and improve water quality. The U.S. Geological Survey joined a long-term research and monitoring program in collaboration with The Nature Conservancy, the Bureau of Reclamation, and Oregon State University in 2008 to assess the effects of the Williamson River Delta restoration on the early life-history stages of Lost River and shortnose suckers. The primary objectives of the research were to describe habitat colonization and use by larval and juvenile suckers and non-sucker fishes and to evaluate the effects of the restored habitat on the health and condition of juvenile suckers. This report summarizes data collected in 2010 by the U.S. Geological Survey as a part of this monitoring effort and follows two annual reports on data collected in 2008 and 2009. Restoration modifications made to the Williamson River Delta appeared to provide additional suitable rearing habitat for endangered Lost River and shortnose suckers from 2008 to 2010 based on sucker catches. Mean larval sample density was greater for both species in the Williamson River Delta than adjacent lake habitats in all 3 years. In addition to larval suckers, at least three age classes of juvenile suckers were captured in the delta. The shallow Goose Bay Farms and Tulana Emergent were among the most used habitats by age-0 suckers in 2009. Both of these environments became inaccessible due to low water in 2010, however, and were not sampled after July 19, 2010. In contrast, age-1 sucker catches shifted from the shallow water (about 0.5-1.5 m deep) on the eastern side of the Williamson River Delta in May, to deeper water environments (greater than 2 m) by the end of June or early July in all 3 years. Differential distribution among sucker species within the Williamson River Delta and between the delta and adjacent lakes indicated that shortnose suckers likely benefited more from the restored Williamson River Delta than Lost River or Klamath largescale suckers (Catostomus snyderi). Catch rates in shallow-water habitats within the delta were higher for shortnose and Klamath largescale sucker larvae than for larval Lost River suckers in 2008, 2009, and 2010. Shortnose suckers also comprised the greatest portion of age-0 suckers captured in the Williamson River Delta in all 3 years of the study. The relative abundance of age-1 shortnose suckers was high in our catches compared to age-1 Lost River suckers in 2009 and 2010. The restored delta also created habitat for several piscivorous fishes, but only two appeared to pose a meaningful threat of predation to suckers - fathead minnows (Pimephales promelas) and yellow perch (Perca flavescens). Fathead minnows that prey on larval but not juvenile suckers dominated catches in all sampling areas. Yellow perch also were abundant throughout the study area, but based on their gape size and co-occurrence with suckers, most were only capable of preying on larvae. Low May lake-surface elevation, below average snow pack, and anticipated irrigation demands indicated late summer water levels in Upper Klamath Lake would be unusually low in 2010. In response to concerns by the Fish and Wildlife Service and The Nature Conservancy that low-water conditions might strand fish on the delta, low water seine surveys were implemented. Eleven fishes, including both endangered suckers, were captured in seine surveys, including both species of suckers, which continued to use shallow water less than 0.4 m deep through September 21. Lake elevation declined to 1,261.54 m (4,138.9 feet) in mid-September 2010, but did not appear to strand fish or cause large-scale fish mortality.

  1. Demographics and 2008 Run Timing of Adult Lost River (Deltistes luxatus) and Shortnose (Chasmistes brevirostris) Suckers in Upper Klamath Lake

    USGS Publications Warehouse

    Janney, Eric C.; Hayes, Brian S.; Hewitt, David A.; Barry, Patrick M.; Scott, Alta; Koller, Justin; Johnson, Mark; Blackwood, Greta

    2009-01-01

    We used capture-recapture data to assess population dynamics of endangered Lost River suckers (Deltistes luxatus) and shortnose suckers (Chasmistes brevirostris) in Upper Klamath Lake, Oregon. The Cormack-Jolly-Seber method was used to estimate apparent survival probabilities, and a temporal symmetry model was used to estimate annual seniority probabilities. Information theoretic modeling was used to assess variation in parameter estimates due to time, gender, and species. In addition, length data were used to detect multiple year-class failures and events of high recruitment into adult spawning populations. Survival of adult Lost River and shortnose suckers varied substantially across years. Relatively high annual mortality was observed for the lakeshore-spawning Lost River sucker subpopulation in 2002 and for the river spawning subpopulation in 2001. Shortnose suckers experienced high mortality in 2001 and 2004. This indicates that high mortality events are not only species specific, but also are specific to subpopulations for Lost River suckers. Seniority probability estimates and length composition data indicate that recruitment of new individuals into adult sucker populations has been sparse. The overall fitness of Upper Klamath Lake sucker populations are of concern given the low observed survival in some years and the paucity of recent recruitment. During most years, estimates of survival probabilities were lower than seniority probabilities, indicating net losses in adult sucker population abundances. The evidence for decline was more marked for shortnose suckers than for Lost River suckers. Our data indicated that sucker survival for both species, but especially shortnose suckers, was sometimes low in years without any observed fish kills. This indicates that high mortality can occur over a protracted period, resulting in poor annual survival, but will not necessarily be observed in association with a fish kill. A better understanding of the factors influencing adult survival and recruitment into spawning populations is needed. Monitoring these vital parameters will provide a quantitative means to evaluate population status and assess the effectiveness of conservation and recovery efforts.

  2. Predation on larval suckers in the Williamson River Delta revealed by molecular genetic assays—A pilot study

    USGS Publications Warehouse

    Hereford, Danielle M.; Ostberg, Carl O.; Burdick, Summer M.

    2016-06-13

    Predation of endangered Lost River suckers (Deltistes luxatus) and shortnose suckers (Chasmistes brevirostris) during larval egress to Upper Klamath Lake from the Williamson River is poorly understood but may be an important factor limiting recruitment into adult spawning populations. Native and non-native piscivores are abundant in nursery wetland habitat, but larval predation has not been directly studied for all species. Larvae lack hard body structures and digest rapidly in predator digestive systems. Therefore, traditional visual methods for diet analysis may fail to identify the extent of predation on larvae. The goals of this study were to (1) use quantitative polymerase chain reaction (qPCR) and single nucleotide polymorphism (SNP) assays developed for Lost River and shortnose suckers to assay predator stomach contents for sucker DNA, and (2) to assess our ability to use this technique to study predation. Predators were captured opportunistically during larval sucker egress. Concurrent feeding trials indicate that most predators—yellow perch (Perca flaverscens), fathead minnow (Pimephales promelas), blue chub (Gila coerulea), Klamath tui chub (Siphatales bicolor bicolor), Klamath Lake sculpin (Cottus princeps), slender sculpin (Cottus tenuis)—preyed on sucker larvae in the laboratory. However, sucker DNA was not detected in fathead minnow stomachs. Of the stomachs screened from fish captured in the Williamson River Delta, 15.6 percent of yellow perch contained sucker DNA. This study has demonstrated that the application of qPCR and SNP assays is effective for studying predation on larval suckers. We suggest that techniques associated with dissection or detection of sucker DNA from fathead minnow stomachs need improvement.

  3. Aspen Root Sucker Formation and Apical Dominance

    Treesearch

    Robert E. Farmer

    1962-01-01

    Root suckering is the primary mode of regeneration in the aspens, Populus tremuloides Michx. and P. grandidentata Michx. When stems of these species are cut, numerous suckers originating in the root pericycle are formed on their extensive lateral root systems. During their first season of growth, suckers ordinarily reach a height...

  4. Temporal and Spatial Distribution of Endangered Juvenile Lost River and Shortnose Suckers in Relation to Environmental Variables in Upper Klamath Lake, Oregon: 2008 Annual Data Summary

    USGS Publications Warehouse

    Burdick, Summer M.; VanderKooi, Scott P.

    2010-01-01

    Lost River sucker (Deltistes luxatus) and shortnose sucker (Chasmistes brevirostris) were listed as endangered in 1988 for a variety of reasons including apparent recruitment failure. Upper Klamath Lake, Oregon, and its tributaries are considered the most critical remaining habitat for these two species. Age-0 suckers are often abundant in Upper Klamath Lake throughout the summer months, but catches decline dramatically between late August and early September each year and age-1 and older sub-adult suckers are rare. These rapid declines in catch rates and a lack of substantial recruitment into adult sucker populations in recent years suggests sucker populations experience high mortality between their first summer and first spawn. A lack of access to, or abundance of, optimal rearing habitat may exacerbate juvenile sucker mortality or restrict juvenile growth or development. Summer age-0 sucker habitat use and distribution has been studied extensively, but many uncertainties remain about age-1 and older juvenile habitat use, distribution, and movement patterns within Upper Klamath Lake. We designed a study to examine seasonal changes in distribution of age-1 suckers in Upper Klamath Lake as they relate to depth and water quality. In this document, which meets our annual data summary and reporting obligations, we discuss the results of our second annual spring and summer sampling effort. Catch data collected in 2007 and 2008 indicate seasonal changes in age-1 and older juvenile sucker habitat use coincident with changes in water quality, which were previously undocumented. In both years during April and May, age-1 and older juvenile suckers were found in shallow water environments. Then, as water temperatures began to warm throughout Upper Klamath Lake in June, age-1 and older juvenile suckers primarily were captured along the western shore in some of the deepest available environments. Following a dramatic decrease in dissolved oxygen concentrations in Eagle Ridge Trench, juvenile suckers were no longer found along the western shore but were captured throughout the rest of Upper Klamath Lake. When dissolved oxygen concentrations were 4 milligrams per liter or greater along the western shore, juvenile sucker captures were again concentrated in that area. Although this pattern indicates that low dissolved oxygen concentration or another related water-quality limitation may force juvenile suckers to leave the western shore, it is unclear as to why age-1 and older juveniles might be attracted to the area in the first place. Understanding this apparent behavior could be important to managing habitat for these species. In this data summary, we also describe the distribution of catches of age-0 suckers and other fishes in Upper Klamath Lake. These data corroborate previous studies that describe age-0 sucker habitat as shallow relative to depths available in Upper Klamath Lake. In this study, we did not seek, nor find additional clarification on age-0 sucker habitat use and distribution in Upper Klamath Lake. Our brief description of the distribution and abundance of all other fish species caught provides a context in which to assess the rarity of juvenile suckers within the fish community of Upper Klamath Lake.

  5. 76 FR 76337 - Endangered and Threatened Wildlife and Plants; Designation of Critical Habitat for Lost River...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2011-12-07

    ... ecology, as well as the technological advancements made available since preparing the 1994 proposed rule..., advancement in our understanding of Lost River sucker's and shortnose sucker's ecology, and the technological... required for Lost River sucker and shortnose sucker from studies of this species' habitat, ecology, and...

  6. Predation on lake whitefish eggs by longnose suckers

    USGS Publications Warehouse

    Nester, Robert T.; Poe, Thomas P.

    1984-01-01

    In November 1981, we observed intense predation on lake whitefish (Coregonus clupeaformis) eggs by longnose suckers (Catostomus catostomus) on lake whitefish spawning grounds in northwestern Lake Huron. Since longnose suckers commonly frequent the same habitat used by spawning lake whitefish, there exists the potential for high losses of eggs due to sucker predation.

  7. BIOLOGY OF THE LEECH ACTINOBDELLA INEQUIANNULATA MOORE, 1901 (ANNELIDA: HIRUDINEA: RHYNCHOBDELLIDA: GLOSSIPHONIIDAE), PARASITIC ON THE WHITE SUCKER, CATOSTOMUS COMMERSONI LACEPEDE, 1803 AND THE LONGNOSE SUCKER, CATOSTOMUS CATOSTOMUS FORSTER, 1773, IN ALGONQUIN PROVINCIAL PARK, ONTARIO, CANADA

    EPA Science Inventory

    Actinobdella inequiannulata was found on the white sucker, Catostomus commersoni, and less frequently on the longnose sucker, Catostomus catostomus, in Algonquin Provincial Park, Ontario, Canada. Catostomus commersoni parasitized with Act. inequiannulata was collected from July ...

  8. Age-0 Lost River sucker and shortnose sucker nearshore habitat use in Upper Klamath Lake, Oregon: A patch occupancy approach

    USGS Publications Warehouse

    Burdick, S.M.; Hendrixson, H.A.; VanderKooi, S.P.

    2008-01-01

    We examined habitat use by age-0 Lost River suckers Deltistes luxatus and shortnose suckers Chasmistes brevirostris over six substrate classes and in vegetated and nonvegetated areas of Upper Klamath Lake, Oregon. We used a patch occupancy approach to model the effect of physical habitat and water quality conditions on habitat use. Our models accounted for potential inconsistencies in detection probability among sites and sampling occasions as a result of differences in fishing gear types and techniques, habitat characteristics, and age-0 fish size and abundance. Detection probability was greatest during mid- to late summer, when water temperatures were highest and age-0 suckers were the largest. The proportion of sites used by age-0 suckers was inversely related to depth (range = 0.4-3.0 m), particularly during late summer. Age-0 suckers were more likely to use habitats containing small substrate (64 mm) and habitats with vegetation than those without vegetation. Relatively narrow ranges in dissolved oxygen, temperature, and pH prevented us from detecting effects of these water quality features on age-0 sucker nearshore habitat use.

  9. Distribution, Health, and Development of Larval and Juvenile Lost River and Shortnose Suckers in the Williamson River Delta Restoration Project and Upper Klamath Lake, Oregon: 2008 Annual Data Summary

    USGS Publications Warehouse

    Burdick, Summer M.; Ottinger, Christopher; Brown, Daniel T.; VanderKooi, Scott P.; Robertson, Laura; Iwanowicz, Deborah

    2009-01-01

    Federally endangered Lost River sucker Deltistes luxatus and shortnose sucker Chasmistes brevirostris were once abundant throughout their range but populations have declined; they have been extirpated from several lakes, and may no longer reproduce in others. Poor recruitment into the adult spawning populations is one of several reasons cited for the decline and lack of recovery of these species, and may be the consequence of high mortality during juvenile life stages. High larval and juvenile sucker mortality may be exacerbated by an insufficient quantity of suitable rearing habitat. Within Upper Klamath Lake, a lack of marshes also may allow larval suckers to be swept from suitable rearing areas downstream into the seasonally anoxic waters of the Keno Reservoir. The Nature Conservancy (TNC) flooded about 3,600 acres to the north of the Williamson River mouth (Tulana Unit) in October 2007, and about 1,400 acres to the south and east of the Williamson River mouth (Goose Bay Unit) a year later, to retain larval suckers in Upper Klamath Lake, create nursery habitat for suckers, and improve water quality. In collaboration with TNC, the Bureau of Reclamation, and Oregon State University, we began a long-term collaborative research and monitoring program in 2008 to assess the effects of the Williamson River Delta restoration on the early life-history stages of Lost River and shortnose suckers. Our approach includes two equally important aspects. One component is to describe habitat use and colonization processes by larval and juvenile suckers and non-sucker fish species. The second is to evaluate the effects of the restored habitat on the health and condition of juvenile suckers. This report contains a summary of the first year of data collected as a part of this monitoring effort.

  10. When experiment and energy conservation collide: video analysis of an unrolling mat

    NASA Astrophysics Data System (ADS)

    Mungan, Carl E.; Lipscombe, Trevor C.

    2018-03-01

    A mat consisting of round bamboo rods connected by strings perpendicular to their axes unrolls without slipping on a horizontal table. Video analysis is used to measure the position of the centre of the remaining roll as a function of time. It is found to accelerate with time due to the ‘rocket effect’ of the roll ejecting rods backward relative to itself. Mechanical energy is not conserved because of the inelastic collisions of the rods with the table. The fitted coefficient of restitution (COR) is 0.59 ± 0.04 which is consistent with known values for wood on wood. In support of this explanation, progressively smaller values of the COR are found when the mat is unrolled on a flat woven rug and on a shock-absorbing pad. The level of analysis is appropriate to an undergraduate course in physical mechanics.

  11. Demographic analysis of Lost River sucker and shortnose sucker populations in Upper Klamath Lake, Oregon

    USGS Publications Warehouse

    Janney, E.C.; Shively, R.S.; Hayes, B.S.; Barry, P.M.; Perkins, D.

    2008-01-01

    We used 13 years (1995-2007) of capture-mark-recapture data to assess population dynamics of endangered Lost River suckers Deltistes luxatus and shortnose suckers Chasmistes brevirostris in Upper Klamath Lake, Oregon. The Cormack-Jolly-Seber method was used to estimate survival, and information theoretic modeling was used to assess variation due to time, gender, species, and spawning subpopulations. Length data were used to detect multiple year-class failures and events of high recruitment into adult spawning populations. Average annual survival probability was 0.88 for Lost River suckers and 0.76 for shortnose suckers. Mean life span estimates based on these survival rates indicated that Lost River suckers survived long enough on average to attempt reproduction eight times, whereas shortnose suckers only survived to spawn three to four times. Shortnose sucker survival was not only poor in years of fish kills (1995-1997) but also was low in years without fish kills (i.e., 2002 and 2004). This suggests that high mortality occurs in some years but is not necessarily associated with fish kills. Annual survival probabilities were not only different between the two species but also differed between two spawning subpopulations of Lost River suckers. Length composition data indicated that recruitment into spawning populations only occurred intermittently. Populations of both species transitioned from primarily old individuals with little size diversity and consistently poor recruitment in the late 1980s and early 1990s to mostly small, recruit-sized fish by the late 1990s. A better understanding of the factors influencing adult survival and recruitment into spawning populations is needed. Monitoring these vital parameters will provide a quantitative means to evaluate population status and assess the effectiveness of conservation and recovery efforts.

  12. Development of a quantitative assay to measure expression of transforming growth factor ß (TGF-ß) in Lost River sucker (Deltistes luxatus) and shortnose sucker (Chasmistes brevirostris) and evaluation of potential pitfalls in use with field-collected samples

    USGS Publications Warehouse

    Robertson, Laura S.; Ottinger, Christopher A.; Burdick, Summer M.; VanderKooi, Scott P.

    2012-01-01

    The Nature Conservancy is in the process of restoring the Williamson River Delta in an attempt to recreate important juvenile habitat for the endangered shortnose sucker Chasmistes brevirostris and the endangered Lost River sucker Deltistes luxatus. Measurement of TGF-β mRNA expression level was one of the indicators chosen to evaluate juvenile sucker health during the restoration process. TGF-β mRNA expression level has been correlated with disease status in several laboratory studies and TGF-β mRNA expression level has been used as a species-specific indicator of immune status in field-based fish health assessments. We describe here the identification of TGF-β and a possible splice variant from shortnose sucker and from Lost River sucker. The performance of a quantitative RT-PCR assay to measure TGF-β mRNA expression level was evaluated in field-collected spleen and kidney tissue samples. The quality of extracted RNA was higher in tissues harvested in September compared to July and higher in tissues harvested at lower temperature compared to higher temperature. In addition, the expression level of both TGF-β and 18S as assessed by qRT-PCR was higher in samples with higher quality RNA. TGF-β mRNA expression was lower in kidney than in spleen in both Lost River sucker and shortnose sucker.

  13. Health and condition of endangered young-of-the-year Lost River and Shortnose suckers relative to water quality in Upper Klamath Lake, Oregon, 2014–2015

    USGS Publications Warehouse

    Burdick, Summer M.; Conway, Carla M.; Elliott, Diane G.; Hoy, Marshal S.; Dolan-Caret, Amari; Ostberg, Carl O.

    2017-10-19

    Most mortality of endangered Lost River (Deltistes luxatus) and shortnose (Chasmistes brevirostris) suckers in Upper Klamath Lake, Oregon, occurs within the first year of life. Juvenile suckers in Clear Lake Reservoir, California, survive longer and may even recruit to the spawning populations. In a previous (2013–2014) study, the health and condition of juvenile suckers and the dynamics of water quality between Upper Klamath Lake and Clear Lake Reservoir were compared. That study found that apparent signs of stress or exposure to irritants, such as peribiliary cuffing in liver tissue and mild inflammation and necrosis in gill tissues, were present in suckers from both lakes and were unlikely to be clues to the cause of differential mortality between lakes. Seasonal trends in energy storage as glycogen and triglycerides were also similar between lakes, indicating prey limitation was not a likely factor in differential mortality. To better understand the relationship between juvenile sucker health and water quality, we examined suckers collected in 2014–2015 from Upper Klamath Lake, where water quality can be dynamic and, at times, extreme.While there were notable differences in water quality and fish health between years, we were not able to identify any specific water-quality-related causes for differential fish condition. Water quality was generally better in 2014 than in 2015. When considered together afflictions and abnormalities generally indicated healthier suckers in 2014 than 2015. Low dissolved-oxygen events (< 4 milligrams per liter) were less frequent and occurred earlier; high pH events (≥ 9.5) were less frequent and shorter in duration; large diel fluctuations in pH (≥ 1.4) were less frequent; water temperatures were warmer, particularly in July and September; and concentrations of microcystin in both large and small fractions of samples were lower in 2014 than in 2015. Total and therefore also un-ionized ammonia were low in 2014–2015 relative to concentrations known to affect suckers. Petechial hemorrhages of the skin, attached Lernaea spp. and eosinophilic hyaline droplets in the kidney tubules were less prevalent in 2014 than in 2015; however, hyperplastic and hypertrophic gill tissue and trichodinids on the gills were observed more frequently in 2014. There were more suckers with normal liver color and texture in 2014 than in 2015. The prevalence of suckers with liver inflammation was greater in 2014 and only observed in suckers collected after August 5, whereas liver inflammation occurred intermittently in 2015. Liver glycogen among suckers decreased in late-August 2014 and increased from early August to mid-September 2015. Lost River suckers had greater whole-body triglyceride content but a larger proportion with an absence of visceral fat observed in 2014 than in 2015. In contrast, shortnose suckers were similar between years in regard to both whole-body triglyceride and visceral fat. Black-spot-forming parasites (trematode metacercariae) were observed in a higher prevalence on shortnose suckers but not Lost River suckers in 2014 than in 2015. Opercular deformities were less prevalent in both species in 2014 than in 2015.Neither gross nor histological examination revealed a high prevalence of abnormalities in suckers that clearly indicate a primary mechanism for juvenile mortality in Upper Klamath Lake. Histological abnormalities were almost always focal and minimal or mild except where associated with parasites. Mild to severe focal abnormalities associated with Lernaea sp. attachment sites and encysted digenean (trematode) metacercariae are unlikely to be associated with mortality. Severe and diffuse inflammation and hyperplasia of the gills associated with Ichthyobodo sp. on one Lost River sucker, may indicate a potential cause of mortality. High mortality may have primarily occurred outside our study period (for example, in spring or over winter), or was caused by a factor that could not be detected with our methods (for example, predation). Alternatively, abnormalities in a small percentage of passively captured suckers in Upper Klamath Lake may indicate health-related issues that were more prevalent in populations than in our samples. Temporary decreases in liver glycogen stores may also indicate periods of stress, which may eventually lead to mortality of young suckers.

  14. Seasonal Distribution and Abundance of Larval and Juvenile Lost River and Shortnose Suckers in Hanks Marsh, Upper Klamath National Wildlife Refuge, Upper Klamath Lake, Oregon: 2007 Annual Report

    USGS Publications Warehouse

    Anderson, Greer O.; Wilkens, Alexander X.; Burdick, Summer M.; VanderKooi, Scott P.

    2009-01-01

    In the summer of 2007, we undertook an assessment of larval and juvenile sucker use of Hanks Marsh in Upper Klamath Lake, Oregon. This 1,200-acre marsh on the southeastern shoreline of the lake represents part of the last remaining natural emergent wetland habitat in the lake. Because of the suspected importance of this type of habitat to larval and juvenile endangered Lost River and shortnose suckers, it was thought that sucker abundance in the marsh might be comparatively greater than in other non-vegetated areas of the lake. It also was hoped that Hanks Marsh would serve as a reference site for wetland restoration projects occurring in other areas of the lake. Our study had four objectives: to (1) examine seasonal distribution and relative abundance of larval suckers in and adjacent to Hanks Marsh in relation to habitat features such as depth, vegetation, water quality, and relative abundance of non-sucker species; (2) determine the presence or absence and describe the distribution of juvenile suckers [35 to 80 mm standard length (SL)] along the periphery of Hanks Marsh; (3) assess spatial and temporal overlap between larval suckers and their potential predators; and (4) assess suitability of water quality throughout the summer for young-of-the-year suckers. Due to the low number of suckers found in the marsh and our inability to thoroughly sample all marsh habitats due to declining lake levels during the summer, we were unable to completely address these objectives in this pilot study. The results, however, do give some indication of the relative use of Hanks Marsh by sucker and non-sucker species. Through sampling of larval and juvenile suckers in various habitat types within the marsh, we determined that sucker use of Hanks Marsh may be very low in comparison with other areas of the lake. We caught only 42 larval and 19 juvenile suckers during 12 weeks of sampling throughout the marsh. Sucker catches were rare in Hanks Marsh, and were lower than catch rates in other marshes of Upper Klamath Lake and in other nearshore and offshore areas of the lake. Based on the few suckers we did capture in Hanks Marsh, larvae tended to be found more often in vegetated habitats. A modified sampling design and approach may be necessary to address the objectives in this study, given that declining lake-surface elevation prevented us from adequately sampling all portions of the marsh throughout the sampling season. Common non-sucker species in Hanks Marsh included juvenile and adult brown bullhead, larval blue chub, tui chub, fathead minnow, and yellow perch. This species composition was similar to that of other marshes in Upper Klamath Lake but most species were found in lower numbers in Hanks Marsh than other marshes. It may be that use of Hanks Marsh is limited by poor water quality, which we found to exist at many sites after June. It also may be that access to or habitat in the marsh is limited at certain times of the year by low water. Although the results from this initial study of Hanks Marsh indicate that the area may have little direct benefit for sucker species, indirect benefits for these species possibly may come from its positive influence on some aspects of water quality in the lake, such as regulation of pH. It also may be the case that use of Hanks Marsh may vary by year and conditions; however, under the current scope of the study, we were unable to investigate inter-annual variability.

  15. An acoustic study of the Brazilian cuica

    NASA Astrophysics Data System (ADS)

    Wheeler, Paul A.

    2002-11-01

    The cuica is a friction drum of African origin played in the batucada (an ensemble of instruments used for the samba) during the Brazilian carnival. It is played by rubbing a bamboo rod which is connected to the center of a drum head, giving a rhythmic grunting sound. Pitch is changed by applying pressure to the membrane. This paper discusses several acoustic aspects of a folk cuica (made of a gourd) including the waveforms, spectra, and time envelopes produced. Rubbing the bamboo rod gives a primitive saw-toothed excitation, similar to a bowed violin string. This is connected to the center of a membrane which modifies and radiates the sound. The body of the cuica contributes little to the sound.

  16. Clone expansion and competition between quaking and bigtooth aspen suckers after clearcutting.

    Treesearch

    Donald A. Perala

    1981-01-01

    Quaking aspen clones expanded over more area, and regenerated greater sucker stem densities and biomass than did bigtooth aspen clones. However, sucker height growth was similar between the two species.

  17. Assessing movement and sources of mortality of juvenile catostomids using passive integrated transponder tags, Upper Klamath Lake, Oregon - Summary of 2012 effort

    USGS Publications Warehouse

    Burdick, Summer M.

    2013-01-01

    Survival of juvenile endangered Lost River and shortnose suckers is thought to limit recruitment into the adult populations and ultimately limit the recovery of these species in Upper Klamath Lake, Oregon. Although many hypotheses exist about the sources of mortality, the contribution of each speculated source of mortality has not been examined. To examine causes of mortality, validate estimated age to maturity, and examine movement patterns for juvenile suckers in Upper Klamath Lake, passive integrated transponder (PIT) tags and remote tag detection systems were used. Age-1 suckers were opportunistically tagged in 2009 and 2010 during another study on juvenile sucker distribution. After the distribution study concluded in 2010, USGS redirected sampling efforts to target age-1 suckers for tagging. Tags were redetected using an existing infrastructure of remote PIT tag readers and tag scanning surveys at American white pelican (Pelecanus erythrorhynchos), double-crested cormorant (Phalacrocorax auritus), and Forster’s tern (Sterna forsteri) breeding and loafing areas. Individual fish histories are used to describe the distance, direction, and timing of juvenile sucker movement. Sucker PIT tag detections in the Sprague and Williamson Rivers in mid-summer and in autumn indicate tagged juvenile suckers use these tributaries outside of the known spring spawning season. PIT tags detected in bird habitats indicate predation by birds was a cause of mortality.

  18. Aspen Sucker Production and Growth from Outplanted Root Cuttings

    Treesearch

    Donald A. Perala

    1978-01-01

    Aspen suckers from 1-m-long root cuttings survived and grew better than those from 12.5-cm-long cuttings. Sucker survival and growth were also inversely related to parent root diameter. Discusses the practical implications for aspen management.

  19. Spawning migration movements of Lost River and shortnose suckers in the Williamson and Sprague Rivers, Oregon, following the removal of Chiloquin Dam-2009 Annual Report

    USGS Publications Warehouse

    Ellsworth, Craig M.; VanderKooi, Scott P.

    2011-01-01

    The Chiloquin Dam was located at river kilometer (rkm) 1.3 on the Sprague River near the town of Chiloquin, Oregon. The dam was identified as a barrier that potentially inhibited or prevented the upstream spawning migrations and other movements of endangered Lost River suckers (Deltistes luxatus), shortnose suckers (Chasmistes brevirostris), and other fish in the Sprague River. Our research objectives in 2009 were to evaluate adult catostomid spawning migration patterns using radio telemetry to identify and describe shifts in spawning area distribution and migration behavior following the removal of Chiloquin Dam in 2008. We attached external radio transmitters to 58 Lost River suckers and 59 shortnose suckers captured at the Williamson River fish weir. A total of 17 radio-tagged Lost River suckers and one radio-tagged shortnose sucker were detected approaching the site of the former Chiloquin Dam but only two radio-tagged fish (one male Lost River sucker and one female Lost River sucker) were detected crossing upstream of the dam site. A lower proportion of radio-tagged shortnose suckers were detected migrating into the Sprague River when compared with previous years. Detections on remote passive integrated transponder (PIT) tag arrays located in the Sprague River show that although the proportion of fish coming into the Sprague River is small when compared to the number of fish crossing the Williamson River fish weir, the number of fish migrating upstream of the Chiloquin Dam site increased exponentially in the first year since its removal. These data will be used in conjunction with larval production and adult spawning distribution data to evaluate the effectiveness of dam removal in order to provide increased access to underutilized spawning habitat located further upstream in the Sprague River and to reduce the crowding of spawning fish below the dam site.

  20. Structure and mechanical properties of Octopus vulgaris suckers.

    PubMed

    Tramacere, Francesca; Kovalev, Alexander; Kleinteich, Thomas; Gorb, Stanislav N; Mazzolai, Barbara

    2014-02-06

    In this study, we investigate the morphology and mechanical features of Octopus vulgaris suckers, which may serve as a model for the creation of a new generation of attachment devices. Octopus suckers attach to a wide range of substrates in wet conditions, including rough surfaces. This amazing feature is made possible by the sucker's tissues, which are pliable to the substrate profile. Previous studies have described a peculiar internal structure that plays a fundamental role in the attachment and detachment processes of the sucker. In this work, we present a mechanical characterization of the tissues involved in the attachment process, which was performed using microindentation tests. We evaluated the elasticity modulus and viscoelastic parameters of the natural tissues (E ∼ 10 kPa) and measured the mechanical properties of some artificial materials that have previously been used in soft robotics. Such a comparison of biological prototypes and artificial material that mimics octopus-sucker tissue is crucial for the design of innovative artificial suction cups for use in wet environments. We conclude that the properties of the common elastomers that are generally used in soft robotics are quite dissimilar to the properties of biological suckers.

  1. Structure and mechanical properties of Octopus vulgaris suckers

    PubMed Central

    Tramacere, Francesca; Kovalev, Alexander; Kleinteich, Thomas; Gorb, Stanislav N.; Mazzolai, Barbara

    2014-01-01

    In this study, we investigate the morphology and mechanical features of Octopus vulgaris suckers, which may serve as a model for the creation of a new generation of attachment devices. Octopus suckers attach to a wide range of substrates in wet conditions, including rough surfaces. This amazing feature is made possible by the sucker's tissues, which are pliable to the substrate profile. Previous studies have described a peculiar internal structure that plays a fundamental role in the attachment and detachment processes of the sucker. In this work, we present a mechanical characterization of the tissues involved in the attachment process, which was performed using microindentation tests. We evaluated the elasticity modulus and viscoelastic parameters of the natural tissues (E ∼ 10 kPa) and measured the mechanical properties of some artificial materials that have previously been used in soft robotics. Such a comparison of biological prototypes and artificial material that mimics octopus-sucker tissue is crucial for the design of innovative artificial suction cups for use in wet environments. We conclude that the properties of the common elastomers that are generally used in soft robotics are quite dissimilar to the properties of biological suckers. PMID:24284894

  2. 21 CFR 870.4420 - Cardiopulmonary bypass cardiotomy return sucker.

    Code of Federal Regulations, 2011 CFR

    2011-04-01

    ... 21 Food and Drugs 8 2011-04-01 2011-04-01 false Cardiopulmonary bypass cardiotomy return sucker... Cardiopulmonary bypass cardiotomy return sucker. (a) Identification. A cardiopulmonary bypass cardiotomy return... from the chest or heart during cardiopulmonary bypass surgery. (b) Classification. Class II...

  3. 21 CFR 870.4420 - Cardiopulmonary bypass cardiotomy return sucker.

    Code of Federal Regulations, 2010 CFR

    2010-04-01

    ... 21 Food and Drugs 8 2010-04-01 2010-04-01 false Cardiopulmonary bypass cardiotomy return sucker... Cardiopulmonary bypass cardiotomy return sucker. (a) Identification. A cardiopulmonary bypass cardiotomy return... from the chest or heart during cardiopulmonary bypass surgery. (b) Classification. Class II...

  4. Musculoskeletal determinants of pelvic sucker function in Hawaiian stream gobiid fishes: interspecific comparisons and allometric scaling.

    PubMed

    Maie, Takashi; Schoenfuss, Heiko L; Blob, Richard W

    2013-07-01

    Gobiid fishes possess a distinctive ventral sucker, formed from fusion of the pelvic fins. This sucker is used to adhere to a wide range of substrates including, in some species, the vertical cliffs of waterfalls that are climbed during upstream migrations. Previous studies of waterfall-climbing goby species have found that pressure differentials and adhesive forces generated by the sucker increase with positive allometry as fish grow in size, despite isometry or negative allometry of sucker area. To produce such scaling patterns for pressure differential and adhesive force, waterfall-climbing gobies might exhibit allometry for other muscular or skeletal components of the pelvic sucker that contribute to its adhesive function. In this study, we used anatomical dissections and modeling to evaluate the potential for allometric growth in the cross-sectional area, effective mechanical advantage (EMA), and force generating capacity of major protractor and retractor muscles of the pelvic sucker (m. protractor ischii and m. retractor ischii) that help to expand the sealed volume of the sucker to produce pressure differentials and adhesive force. We compared patterns for three Hawaiian gobiid species: a nonclimber (Stenogobius hawaiiensis), an ontogenetically limited climber (Awaous guamensis), and a proficient climber (Sicyopterus stimpsoni). Scaling patterns were relatively similar for all three species, typically exhibiting isometric or negatively allometric scaling for the muscles and lever systems examined. Although these scaling patterns do not help to explain the positive allometry of pressure differentials and adhesive force as climbing gobies grow, the best climber among the species we compared, S. stimpsoni, does exhibit the highest calculated estimates of EMA, muscular input force, and output force for pelvic sucker retraction at any body size, potentially facilitating its adhesive ability. Copyright © 2013 Wiley Periodicals, Inc.

  5. Inspiration, simulation and design for smart robot manipulators from the sucker actuation mechanism of cephalopods.

    PubMed

    Grasso, Frank W; Setlur, Pradeep

    2007-12-01

    Octopus arms house 200-300 independently controlled suckers that can alternately afford an octopus fine manipulation of small objects and produce high adhesion forces on virtually any non-porous surface. Octopuses use their suckers to grasp, rotate and reposition soft objects (e.g., octopus eggs) without damaging them and to provide strong, reversible adhesion forces to anchor the octopus to hard substrates (e.g., rock) during wave surge. The biological 'design' of the sucker system is understood to be divided anatomically into three functional groups: the infundibulum that produces a surface seal that conforms to arbitrary surface geometry; the acetabulum that generates negative pressures for adhesion; and the extrinsic muscles that allow adhered surfaces to be rotated relative to the arm. The effector underlying these abilities is the muscular hydrostat. Guided by sensory input, the thousands of muscle fibers within the muscular hydrostats of the sucker act in coordination to provide stiffness or force when and where needed. The mechanical malleability of octopus suckers, the interdigitated arrangement of their muscle fibers and the flexible interconnections of its parts make direct studies of their control challenging. We developed a dynamic simulator (ABSAMS) that models the general functioning of muscular hydrostat systems built from assemblies of biologically constrained muscular hydrostat models. We report here on simulation studies of octopus-inspired and artificial suckers implemented in this system. These simulations reproduce aspects of octopus sucker performance and squid tentacle extension. Simulations run with these models using parameters from man-made actuators and materials can serve as tools for designing soft robotic implementations of man-made artificial suckers and soft manipulators.

  6. Health and condition of endangered juvenile Lost River and shortnose suckers relative to water quality and fish assemblages in Upper Klamath Lake, Oregon, and Clear Lake Reservoir, California

    USGS Publications Warehouse

    Burdick, Summer M.; Elliott, Diane G.; Ostberg, Carl O.; Conway, Carla M.; Dolan-Caret, Amari; Hoy, Marshal S.; Feltz, Kevin P.; Echols, Kathy R.

    2015-11-25

    Differences in sucker health and condition between lakes were considered the most promising clues to the causes of differential juvenile sucker morality between lakes. A low prevalence of petechial hemorrhaging of the skin (16 percent) and deformed opercula (8 percent) in Upper Klamath Lake suckers may indicate exposure to a toxin other than microcystin. Suckers grew slower in their first year of life, but had similar or greater triglyceride and glycogen levels in Upper Klamath Lake compared to Clear Lake Reservoir. These findings do not suggest a lack of prey quantity but may indicate lower prey quality in Upper Klamath Lake.

  7. Properties of Interfacial Tribo-Films

    DTIC Science & Technology

    1993-06-01

    cf these rods is such as to have the center of gravity of or the attraction of water into the re-entrant peripheral gap the whole sample as close as...difference between the fluid dynamics, acoustic effects in stringed musical static and the kinetic friction coefficients increases with instruments...interfacial fluid molecules to static minimize oscillations, the center of gravity of the sample friction have been explored and, in this regard, adsorbed

  8. BIOLOGY AND OCCURRENCE OF THE LEECH, ACTINOBDELLA INEQUIANNULATA (ANNELIDA: HIRUDINEA: GLOSSIPHONIIDAE) PARASITIC ON TWO SPECIES OF SUCKERS

    EPA Science Inventory

    Actinobdella inequiannulata was found on the white sucker, Catostomus commersoni, and less frequently on the longnose sucker, Catostomus catostomus, in Algonquin Provincial Park, Ontario, Canada. This study established the presence of only one species of leech, Actinobdela inequ...

  9. Regeneration

    Treesearch

    George A. Schier; Wayne D. Shepperd; John R. Jones

    1985-01-01

    There are basically two approaches to regenerating aspen stands-sexual reproduction using seed, or vegetative regeneration by root suckering. In the West, root suckering is the most practical method. The advantage of having an existing, well established root system capable of producing numerous root suckers easily outweighs natural or artificial reforestation in the...

  10. The effects of habitat restoration on endangered fishes in the Upper Klamath Basin

    NASA Astrophysics Data System (ADS)

    Vanderkooi, S.; Burdick, S.; Ellsworth, C.

    2009-12-01

    The Klamath Basin has been prominent in the debate over the use of water in the arid west for nearly a decade. Principle competing demands include threatened and endangered species, agriculture, recreation, and Tribal rights. In an effort to recover two endangered fish species, the Lost River sucker and the shortnose sucker, several large-scale restoration projects have recently been undertaken in the basin. These include restoration of 7000 acres of drained wetlands in the Williamson River Delta in 2007 and 2008 and the removal of Chiloquin Dam on the Sprague River in 2008. The objectives of these projects include increasing amounts of usable aquatic habitats, restoring watershed function, improving water quality, and improving access to and connectivity among habitats. The U.S. Geological Survey is involved in a series of collaborative, interdisciplinary research efforts to evaluate the effectiveness of these restoration projects. Effectiveness is being evaluated by physical, biological and ecological metrics. Our focus is on the biology and ecology of endangered suckers. In the Williamson River Delta, we’re collecting data on the distribution, habitat use, relative abundance, and health and condition of early life-history stages of endangered suckers. Results to date indicate larval as well as age-0 and age-1 juvenile suckers are using newly created habitats in the delta. Preliminary results from comparisons of age-0 suckers captured within and outside of the delta suggest those using the restored habitats are in better condition. In the Sprague River, we’re studying the behavior, run timing, and distribution of adult suckers during spawning migrations as well as the relative abundance, species composition, and timing of emigrating larval suckers. Preliminary results indicate adult suckers migrated into the reach previously impounded by Chiloquin Dam in greater numbers than in the past. While these results indicate a positive response to dam removal, population-level effects of such restoration projects on suckers will not be fully evident for years to come because of the long life span and slow maturation rates of these fish. Continued monitoring of physical and biological changes due to restoration is essential to determining the effectiveness of these actions and informing future efforts.

  11. Life history and ecological characteristics of the Santa Ana sucker, Catostomus santaanae

    USGS Publications Warehouse

    Saiki, Michael K.; Martin, Barbara A.; Knowles, Glen W.; Tennant, Patrick W.

    2007-01-01

    This study was conducted to document the life history and ecological characteristics of the Santa Ana sucker, Catostomus santaanae, within its native range in southern California. Electrofishing surveys were conducted at 3-month intervals from December 1998 to December 1999 at one site on the San Gabriel River and two sites on the Santa Ana River. Suckers were captured in the San Gabriel River (average, 6.6 fish/10-minutes electrofishing) and at an upstream Santa Ana River site (average, 2.3 fish/10-minutes electrofishing) but not at a downstream Santa Ana River site. Length frequency distributions indicated that at least three year classes (modal groups) of suckers were present in the San Gabriel River, whereas one or two year classes were present in the Santa Ana River. Collection of 21-30 mm standard length (SL) juveniles in June in the Santa Ana River and in September in the San Gabriel River indicated that reproduction occurred over several months. In December, Age-0 suckers averaged 36-48 mm SL in the San Gabriel River and 63-65 mm SL in the Santa Ana River, whereas Age-1 suckers averaged 86 mm SL in the San Gabriel River and 115 mm SL in the Santa Ana River. On average, suckers were in better body condition in the San Gabriel River than in the Santa Ana River. Highest abundance of suckers was associated with relativelypristine environmental conditions (especially low specific conductance) where other native fishes were also common or abundant.

  12. Octopus-like suction cups: from natural to artificial solutions.

    PubMed

    Tramacere, F; Follador, M; Pugno, N M; Mazzolai, B

    2015-05-13

    Octopus suckers are able to attach to all nonporous surfaces and generate a very strong attachment force. The well-known attachment features of this animal result from the softness of the sucker tissues and the surface morphology of the portion of the sucker that is in contact with objects or substrates. Unlike artificial suction cups, octopus suckers are characterized by a series of radial grooves that increase the area subjected to pressure reduction during attachment. In this study, we constructed artificial suction cups with different surface geometries and tested their attachment performances using a pull-off setup. First, smooth suction cups were obtained for casting; then, sucker surfaces were engraved with a laser cutter. As expected, for all the tested cases, the engraving treatment enhanced the attachment performance of the elastomeric suction cups compared with that of the smooth versions. Moreover, the results indicated that the surface geometry with the best attachment performance was the geometry most similar to octopus sucker morphology. The results obtained in this work can be utilized to design artificial suction cups with higher wet attachment performance.

  13. Techniques for Monitoring Razorback Sucker in the Lower Colorado River, Hoover to Parker Dams, 2006-2007, Final Report

    USGS Publications Warehouse

    Mueller, Gordon A.; Wydoski, Richard; Best, Eric; Hiebert, Steve; Lantow, Jeff; Santee, Mark; Goettlicher, Bill; Millosovich, Joe

    2008-01-01

    Trammel netting is generally the accepted method of monitoring razorback sucker in reservoirs, but this method is ineffective for monitoring this fish in rivers. Trammel nets set in the current become fouled with debris, and nets set in backwaters capture high numbers of nontarget species. Nontargeted fish composed 97 percent of fish captured in previous studies (1999-2005). In 2005, discovery of a large spawning aggregation of razorback sucker in midchannel near Needles, Calif., prompted the development of more effective methods to monitor this and possibly other riverine fish populations. This study examined the effectiveness of four methods of monitoring razorback sucker in a riverine environment. Hoop netting, electrofishing, boat surveys, and aerial photography were evaluated in terms of data accuracy, costs, stress on targeted fish, and effect on nontargeted fish as compared with trammel netting. Trammel netting in the riverine portion of the Colorado River downstream of Davis Dam, Arizona-Nevada yielded an average of 43 razorback suckers a year (1999 to 2005). Capture rates averaged 0.5 razorback suckers per staff day effort, at a cost exceeding $1,100 per fish. Population estimates calculated for 2003-2005 were 3,570 (95 percent confidence limits [CL] = 1,306i??i??i??-8,925), 1,768 (CL = 878-3,867) and 1,652 (CL = 706-5,164); wide confidence ranges reflect the small sample size. By-catch associated with trammel netting included common carp, game fish and, occasionally, shorebirds, waterfowl, and muskrats. Hoop nets were prone to downstream drift owing to design and anchoring problems aggravated by hydropower ramping. Tests were dropped after the 2006 field season and replaced with electrofishing. Electrofishing at night during low flow and when spawning razorback suckers moved to the shoreline proved extremely effective. In 2006 and 2007, 263 and 299 (respectively) razorback suckers were taken. Capture rates averaged 8.3 razorback suckers per staff day at a cost of $62 per fish. The adult population was estimated at 1,196 (925-1,546) fish. Compared with trammel netting, confidence limits narrowed substantially, from +or- 500 percent to +or- 30 percent, reflecting more precise estimates. By-catch was limited to two common carp. No recreational game fish, waterfowl, or mammals were captured or handled during use of electrofishing. Aerial photography (2006 and 2007) suggested an annual average of 580 fish detected on imagery. Identification of species was not possible; carp commonly have been mistaken for razorback sucker. Field verification determined that the proportion of razorback suckers to other fish was 3:1. On that basis, we estimated 435 razorback suckers were photographed, which equals 8.4 razorback suckers per staff day at a cost of $78 per fish. The data did not lend itself to population estimates. Fish were more easily identified from boats, where their lateral rather than their dorsal aspect is visible. On average, 888 razorback suckers were positively identified each year. Observation rates averaged 29.6 razorback suckers per staff day at a cost less than $18 per fish observed. Sucker densities averaged 20.5 and 9.6 fish/hectare which equated to an average spawning population at Needles, Calif., of 2,520 in 2006 and 1152 in 2007. The lower 2007 estimate reflected a refinement in sampling approach which removed a sampling bias. Electrofishing and boat surveys were more cost effective than other methods tested, and they provided more accurate information without the by-catch associated with trammel netting. However, they provided different types of data. Handling fish may be necessary for research purposes but unnecessary for general trend analysis. Electrofishing was extremely effective but can harm fish if not used with caution. Unnecessary electrofishing increases the likelihood of spinal damage and possible damage to eggs and potential young, and it may alter spawning behavior or duration. B

  14. Bullfrog tadpole (Rana catesbeiana) and red swamp crayfish (Procambarus clarkii) predation on early life stages of endangered razorback sucker (Xyrauchen texanus)

    USGS Publications Warehouse

    Mueller, G.A.; Carpenter, J.; Thornbrugh, D.

    2006-01-01

    Bullfrog tadpoles (Rana catesbeiana) and red swamp crayfish (Procambarus clarkii) are widespread introduced taxa that are problematic throughout the western United States. Their impact on native amphibians and crustaceans is well documented, but less is known regarding their influence on native fishes. Predator-prey tank tests showed both species consumed eggs and larvae of the endangered razorback sucker (Xyrauchen texanus) in a laboratory setting. Tadpoles consumed 2.2 razorback sucker eggs/d and 1.4 razorback sucker larvae/d, while crayfish ate 6.0 eggs/d and 3.5 larvae/d. Relatively high densities of bullfrog tadpoles and crayfish in razorback sucker spawning areas suggest that these nonnative taxa might pose a threat to the recruitment success of this and other imperiled native fish.

  15. Hydro-ball in-core instrumentation system and method of operation

    DOEpatents

    Tower, Stephen N.; Veronesi, Luciano; Braun, Howard E.

    1990-01-01

    A hydro-ball in-core instrumentation system employs detector strings each comprising a wire having radiation sensitive balls affixed diametrically at spaced positions therealong and opposite tip ends of which are transportable by fluid drag through interior passageways. In the passageways primary coolant is caused to flow selectively in first and second opposite directions for transporting the detector strings from stored positions in an exterior chamber to inserted positions within the instrumentation thimbles of the fuel rod assemblies of a pressure vessel, and for return. The coolant pressure within the detector passageways is the same as that within the vessel; face contact, disconnectable joints between sections of the interior passageways within the vessel facilitate assembly and disassembly of the vessel for refueling and routine maintenance operations. The detector strings may pass through a very short bend radius thereby minimizing space requirements for the connections of the instrumentation system to the vessel and concomitantly the vessel containment structure. Improved radiation mapping and a significant reduction in potential exposure of personnel to radiation are provided. Both top head and bottom head penetration embodiments are disclosed.

  16. Using on-site bioassays to determine selenium risk to propagated endangered fishes

    USGS Publications Warehouse

    Allert, Ann L.; Fairchild, James F.; May, Thomas W.; Sappington, Linda C.; Darnall, N.; Wilson, M.

    2006-01-01

    The Utah Reclamation, Mitigation and Conservation Commission is determining the feasibility of establishing a hatchery and grow-out facility for endangered June suckers Chasmistes liorus at Goshen Warm Springs, Utah. A survey of water quality indicated that selenium and other contaminants may be of concern at Goshen Warm Springs. We conducted an ecotoxicological study with three objectives: (1) to determine the growth rates of juvenile June suckers in two ponds (Lily Pond and North Pond) at Goshen Warm Springs and in Utah Lake, (2) to determine the uptake and depuration rates of selenium in juvenile June suckers, and (3) to evaluate limnological factors that may influence growth and selenium accumulation in June suckers. Fish growth was significantly greater at Utah Lake and North Pond than at Lily Pond or under current hatchery conditions. At the end of the uptake phase of the study (day 87), selenium concentrations in June suckers from Lily Pond, North Pond, and Utah Lake were 1.62, 1.90, and 1.32 μg/g of dry weight, respectively. Selenium uptake in June suckers was statistically significant at Lily Pond (0.005 μg·g−1 ·d−1), North Pond (0.010 μg·g−1 ·d−1), and Utah Lake (0.003 μg·g−1 ·d−1). At day 87, the fish were transferred to well water for selenium depuration. Significant selenium depuration occurred after the transfer of June suckers to clean water. Results indicated that selenium bioaccumulated to statistically significant levels at both Lily and North ponds. However, these concentrations are not likely to be of concern because they are not known to cause chronic toxicity. Depuration experiments indicated that June suckers stocked into Utah Lake would eliminate accumulated selenium residues within 3 months. Results indicated that Goshen Warm Springs could be used for fish propagation. In addition, further evaluation of Utah Lake as an interim hatchery site for June suckers should be considered.

  17. Effect of pruning the parent root on growth of aspen suckers

    Treesearch

    Ashbel F. Hough

    1965-01-01

    Various portions of the root systems of bigtooth aspen (Populus grandidentata) suckers were severed, and the subsequent height and radial growth of stems were measured. Aspen vegetative regeneration is heavily dependent on the parent roots for at least 25 years following initial suckering. The distal portion of the parent root contributes more to...

  18. Population Dynamics of Adult Lost River (Deltistes luxatus) and Shortnose (Chasmistes brevirostris) Suckers in Clear Lake Reservoir, California, 2006-08

    USGS Publications Warehouse

    Barry, Patrick M.; Janney, Eric C.; Hewitt, David A.; Hayes, Brian S.; Scott, Alta C.

    2009-01-01

    We report results from ongoing research into the population dynamics of endangered Lost River and shortnose suckers in Clear Lake Reservoir, California. Results are included for sampling that occurred from fall 2006 to spring 2008. We summarize catches and passive integrated transponder tagging efforts from trammel net sampling in fall 2006 and fall 2007, and report on detections of tagged suckers on remote antennas in the primary spawning tributary, Willow Creek, in spring 2007 and spring 2008. Results from trammel net sampling were similar to previous years, although catches of suckers in fall 2006 were lower than in 2007 and past years. Lost River and shortnose suckers combined made up about 80 percent of the sucker catch in each year, and more than 2,000 new fish were tagged across the 2 years. Only a small number of the suckers captured in fall sampling were recaptures of previously tagged fish, reinforcing the importance of remote detections of fish for capture-recapture analysis. Detections of tagged suckers in Willow Creek were low in spring 2007, presumably because of low flows. Nonetheless, the proportions of tagged fish that were detected were reasonably high and capture-recapture analyses should be possible after another year of data collection. Run timing for Lost River and shortnose suckers was well described by first detections of individuals by antennas in Willow Creek, although we may not have installed the antennas early enough in 2008 to monitor the earliest portion of the Lost River sucker migration. The duration and magnitude of the spawning runs for both species were influenced by flows and water temperature. Flows in Willow Creek were much higher in 2008 than in 2007, and far more detections were recorded in 2008 and the migrations were more protracted. In both years and for both species, migrations began in early March at water temperatures between 5 and 6 deg C and peaks were related to periods of increasing water temperature. The sex ratio of Lost River suckers detected in Willow Creek was skewed toward males, despite consistently more females having been tagged in fall sampling. This pattern indicates that some tagged female Lost River suckers may be spawning elsewhere in the system, and we intend to investigate this possibility to verify or alter the representativeness of our spring monitoring. Length frequency analysis of fall trammel net catches showed that the populations of both species in Clear Lake Reservoir have undergone major demographic transitions during the last 15 years. In the mid-1990s, the populations were dominated by larger fish and showed little evidence of recent recruitment. These larger fish apparently disappeared in the late 1990s and early 2000s, and the populations are now dominated by fish that recruited into the adult populations in the late 1990s. The length frequencies from the last 4 years provide evidence of consistent recruitment into the Lost River sucker population, but provide no such evidence for the shortnose sucker population. Overall, annual growth rates for both species in Clear Lake were 2-4 times greater than growth rates for conspecifics in Upper Klamath Lake. However, little or no growth occurred for either species in Clear Lake between 2006 and 2007. Based on available evidence, we are unable to fully explain differences in growth rates between systems or among years within Clear Lake.

  19. The morphology and adhesion mechanism of Octopus vulgaris suckers.

    PubMed

    Tramacere, Francesca; Beccai, Lucia; Kuba, Michael; Gozzi, Alessandro; Bifone, Angelo; Mazzolai, Barbara

    2013-01-01

    The octopus sucker represents a fascinating natural system performing adhesion on different terrains and substrates. Octopuses use suckers to anchor the body to the substrate or to grasp, investigate and manipulate objects, just to mention a few of their functions. Our study focuses on the morphology and adhesion mechanism of suckers in Octopus vulgaris. We use three different techniques (MRI, ultrasonography, and histology) and a 3D reconstruction approach to contribute knowledge on both morphology and functionality of the sucker structure in O. vulgaris. The results of our investigation are two-fold. First, we observe some morphological differences with respect to the octopus species previously studied (i.e., Octopus joubini, Octopus maya, Octopus bimaculoides/bimaculatus and Eledone cirrosa). In particular, in O. vulgaris the acetabular chamber, that is a hollow spherical cavity in other octopuses, shows an ellipsoidal cavity which roof has an important protuberance with surface roughness. Second, based on our findings, we propose a hypothesis on the sucker adhesion mechanism in O. vulgaris. We hypothesize that the process of continuous adhesion is achieved by sealing the orifice between acetabulum and infundibulum portions via the acetabular protuberance. We suggest this to take place while the infundibular part achieves a completely flat shape; and, by sustaining adhesion through preservation of sucker configuration. In vivo ultrasonographic recordings support our proposed adhesion model by showing the sucker in action. Such an underlying physical mechanism offers innovative potential cues for developing bioinspired artificial adhesion systems. Furthermore, we think that it could possibly represent a useful approach in order to investigate any potential difference in the ecology and in the performance of adhesion by different species.

  20. The Morphology and Adhesion Mechanism of Octopus vulgaris Suckers

    PubMed Central

    Tramacere, Francesca; Beccai, Lucia; Kuba, Michael; Gozzi, Alessandro; Bifone, Angelo; Mazzolai, Barbara

    2013-01-01

    The octopus sucker represents a fascinating natural system performing adhesion on different terrains and substrates. Octopuses use suckers to anchor the body to the substrate or to grasp, investigate and manipulate objects, just to mention a few of their functions. Our study focuses on the morphology and adhesion mechanism of suckers in Octopus vulgaris. We use three different techniques (MRI, ultrasonography, and histology) and a 3D reconstruction approach to contribute knowledge on both morphology and functionality of the sucker structure in O. vulgaris. The results of our investigation are two-fold. First, we observe some morphological differences with respect to the octopus species previously studied (i.e., Octopus joubini, Octopus maya, Octopus bimaculoides/bimaculatus and Eledone cirrosa). In particular, in O. vulgaris the acetabular chamber, that is a hollow spherical cavity in other octopuses, shows an ellipsoidal cavity which roof has an important protuberance with surface roughness. Second, based on our findings, we propose a hypothesis on the sucker adhesion mechanism in O. vulgaris. We hypothesize that the process of continuous adhesion is achieved by sealing the orifice between acetabulum and infundibulum portions via the acetabular protuberance. We suggest this to take place while the infundibular part achieves a completely flat shape; and, by sustaining adhesion through preservation of sucker configuration. In vivo ultrasonographic recordings support our proposed adhesion model by showing the sucker in action. Such an underlying physical mechanism offers innovative potential cues for developing bioinspired artificial adhesion systems. Furthermore, we think that it could possibly represent a useful approach in order to investigate any potential difference in the ecology and in the performance of adhesion by different species. PMID:23750233

  1. Tegumental ultrastructure of Echinoparyphium recurvatum according to developmental stages

    PubMed Central

    Woo, Ho-Choon; Hong, Sung-Jong

    2002-01-01

    The present study was performed to observe tegumental ultrastructure of Echinoparyphium recurvatum according to developmental stages. Worms (1, 3, 5 and 15-day old) were recovered from chicks experimentally infected with metacercariae from Radix auricularia coreana. One-day old worms were elongated and ventrally concave, and covered with peg-like tegumental spines except the adjacent areas of the head crown and excretory pore. Type I sensory papillae were distributed on the lip of the oral sucker, and grouped ciliated papillae were around the oral sucker. Peg-like tegumental spines were densely distributed on the anterior surface of the ventral sucker level. The ventral sucker had an aspinous tegument and no sensory papillae. Tegumental spines on the posterior surface of the ventral sucker level were sparsely distributed and disappeared posteriorly. In 3 and 5-day old worms, the tegument around the oral sucker was aspinose and wrinkled concentrically. The ventral sucker had a wrinkled tegument and many bulbous papillae. Type I sensory papillae were distributed between the bulbous papillae. Tegumental spines were spade-shaped with a terminal tip. A total of 45 collar spines including 4 end group ones on both ventral corners was alternately arranged in 2 rows. The 15-day old worms were very stout and their tegumental spines were tongue-shaped without a terminal tip. From the above results, it is confirmed that the surface ultrastructure of E. recurvatum was generally similar to that of other echinostomatid flukes. However, some features, i.e., morphological change of tegumental spines and appearence of sensory papillae on the ventral sucker according to development, and number, shape and arrangement of collar spines, were characteristic, which may be of taxonomic and bioecological significance. PMID:12073731

  2. Evaluation of techniques to protect aspen suckers from ungulate browsing in the Black Hills

    Treesearch

    Andrew M. Kota; Dale L. Bartos

    2010-01-01

    Excessive browsing by cattle (Bos taurus L.) and wild ungulates, particularly elk (Cervus elaphus L.), sometimes inhibits growth and maturity of quaking aspen (Populus tremuloides Michx.) suckers in Western North America. In areas where aspen stands are in poor condition or declining, protecting suckers from ungulates may be necessary. This study compared the utility...

  3. Suckers and other bursal structures of Pomphorhynchus bulbocolli and Acanthocephalus dirus (Acanthocephala).

    PubMed

    Doyle, L R; Gleason, L N

    1991-06-01

    Accessory copulatory structures in the bursa of Pomphorhynchus bulbocolli include 2 suckers on the inner surface and bursal rays. A stylet was present in the penis of this species. Two suckers were present in the bursa of Acanthocephalus dirus, but bursal rays and a stylet in the penis were not observed. This is the first report of suckers present in the bursas of acanthocephalans. Copulatory cement caps and holes surrounded by a ring approximating the inner bursal diameter and not covered by a cap were present on the sides of some P. bulbocolli, indicating the possibility of hypodermic insemination in this species.

  4. Population impacts in white sucker (Catostomus commersonii) exposed to oil sands-derived contaminants in the Athabasca River.

    PubMed

    Arens, Collin J; Arens, Jennifer C; Hogan, Natacha S; Kavanagh, Richard J; Berrue, Fabrice; Van Der Kraak, Glen J; van den Heuvel, Michael R

    2017-08-01

    Biological and chemical endpoints were measured in white sucker collected downstream of Athabasca oil sands developments (AB, Canada) and compared with those at Calling Lake (AB, Canada), a reference location upstream of the Athabasca oil sands deposit. Naphthenic acid concentrations were also measured at 14 sites in the Athabasca River watershed. Concentrations of naphthenic acids were elevated in tributaries adjacent to oil sands mining developments. Tributary naphthenic acid profiles were more similar to aged oil sands process water than samples from the Athabasca River, suggesting an influence of tailings in the tributaries. White sucker showed higher energy storage in the Athabasca River as indicated by significantly higher condition and liver size. White sucker were not investing that energy into reproductive effort as measured by gonad size and fecundity, which were significantly reduced relative to the reference location. White sucker showed increased exposure to polycyclic aromatic hydrocarbons as indicated by hepatic cytochrome P4501A (CYP1A) activity and fluorescent bile metabolites, as well as higher concentrations of naphthenic acids in bile. Cadmium, copper, nickel, and selenium were also elevated in white sucker liver tissue compared with the reference location. Based on the exposure profile and response pattern observed, effects on energy storage and utilization in white sucker from the Athabasca River most likely resulted from exposure to polycyclic aromatic hydrocarbons derived from petrogenic and pyrolytic sources. Environ Toxicol Chem 2017;36:2058-2067. © 2017 SETAC. © 2017 SETAC.

  5. Use of pultruded reinforced plastics in energy generation and energy related applications

    NASA Astrophysics Data System (ADS)

    Anderson, R.

    Applications of pultrusion-formed fiber-reinforced plastics (FRP) in the wind, oil, and coal derived energy industries are reviewed. FRP is noted to be a viable alternative to wood, aluminum, and steel for reasons of availability, price, and weight. Attention is given to the development of FRP wind turbine blades for the DOE 8 kW low cost, high reliability wind turbine program. The blades feature a NACA 23112 profile with a 15 in. chord on the system which was tested at Rocky Flats, CO. Fabricating the blades involved a plus and minus 45 deg roving orientation, a heavy fiber-glass nose piece to assure blade strength, and a separately manufactured foam core. Additional uses for FRP products have been found in the structural members of coal stack scrubbers using a vinyl ester resin in a fire retardant formulation, and as low cost, light weight sucker rods for deep well oil drilling.

  6. Environmental Assessment: Space Innovation and Development Center Schriever AFB, Colorado

    DTIC Science & Technology

    2006-03-01

    Ptychocheilus lucius E Gunnison sage-grouse Centrocercus minimus c Humpback chub Gila cypha E Razorback sucker© Xyrauchentexanus E Uinta Basin hookless...debilis c Razorback sucker© Xyrauchen texanus E Uinta Basin hookless cactus Sclerocactus glaucus T Yellow-billed cuckoo Coccyzus americanus c GILPIN...Razorback sucker© Xyrauchen texanus E Uinta Basin hookless cactus Sclerocactus glaucus T Yellow-billed cuckoo Coccyzus americanus c MINERAL Bald

  7. Regeneration of aspen by suckering on burned sites in western Wyoming

    Treesearch

    Dale L. Bartos; Walter F. Mueggler; Robert B. Campbell

    1991-01-01

    Numbers of suckers produced following burning of aspen stands varied greatly between areas but followed similar trends over a 6-year period: a relative abundance the first year, followed by an abrupt decline by the third year and a gradual decline thereafter. First-year sucker numbers that ranged from 34,000 to 147,000/ha suffered mortality between 54 and 93 percent by...

  8. A comparative analysis of genetic variation in rootstocks and scions of old olive trees - a window into the history of olive cultivation practices and past genetic variation.

    PubMed

    Barazani, Oz; Westberg, Erik; Hanin, Nir; Dag, Arnon; Kerem, Zohar; Tugendhaft, Yizhar; Hmidat, Mohammed; Hijawi, Thameen; Kadereit, Joachim W

    2014-05-28

    Past clonal propagation of olive trees is intimately linked to grafting. However, evidence on grafting in ancient trees is scarce, and not much is known about the source of plant material used for rootstocks. Here, the Simple Sequence Repeat (SSR) marker technique was used to study genetic diversity of rootstocks and scions in ancient olive trees from the Levant and its implications for past cultivation of olives. Leaf samples were collected from tree canopies (scions) and shoots growing from the trunk base (suckers). A total of 310 trees were sampled in 32 groves and analyzed with 14 SSR markers. In 82.7% of the trees in which both scion and suckers could be genotyped, these were genetically different, and thus suckers were interpreted to represent the rootstock of grafted trees. Genetic diversity values were much higher among suckers than among scions, and 194 and 87 multi-locus genotypes (MLGs) were found in the two sample groups, respectively. Only five private alleles were found among scions, but 125 among suckers. A frequency analysis revealed a bimodal distribution of genetic distance among MLGs, indicating the presence of somatic mutations within clones. When assuming that MLGs differing by one mutation are identical, scion and sucker MLGs were grouped in 20 and 147 multi-locus lineages (MLLs). The majority of scions (90.0%) belonged to a single common MLL, whereas 50.5% of the suckers were single-sample MLLs. However, one MLL was specific to suckers and found in 63 (22.6%) of the samples. Our results provide strong evidence that the majority of olive trees in the study are grafted, that the large majority of scions belong to a single ancient cultivar containing somatic mutations, and that the widespread occurrence of one sucker genotype may imply rootstock selection. For the majority of grafted trees it seems likely that saplings were used as rootstocks; their genetic diversity probably is best explained as the result of a long history of sexual reproduction involving cultivated, feral and wild genotypes.

  9. A comparative analysis of genetic variation in rootstocks and scions of old olive trees – a window into the history of olive cultivation practices and past genetic variation

    PubMed Central

    2014-01-01

    Background Past clonal propagation of olive trees is intimately linked to grafting. However, evidence on grafting in ancient trees is scarce, and not much is known about the source of plant material used for rootstocks. Here, the Simple Sequence Repeat (SSR) marker technique was used to study genetic diversity of rootstocks and scions in ancient olive trees from the Levant and its implications for past cultivation of olives. Leaf samples were collected from tree canopies (scions) and shoots growing from the trunk base (suckers). A total of 310 trees were sampled in 32 groves and analyzed with 14 SSR markers. Results In 82.7% of the trees in which both scion and suckers could be genotyped, these were genetically different, and thus suckers were interpreted to represent the rootstock of grafted trees. Genetic diversity values were much higher among suckers than among scions, and 194 and 87 multi-locus genotypes (MLGs) were found in the two sample groups, respectively. Only five private alleles were found among scions, but 125 among suckers. A frequency analysis revealed a bimodal distribution of genetic distance among MLGs, indicating the presence of somatic mutations within clones. When assuming that MLGs differing by one mutation are identical, scion and sucker MLGs were grouped in 20 and 147 multi-locus lineages (MLLs). The majority of scions (90.0%) belonged to a single common MLL, whereas 50.5% of the suckers were single-sample MLLs. However, one MLL was specific to suckers and found in 63 (22.6%) of the samples. Conclusions Our results provide strong evidence that the majority of olive trees in the study are grafted, that the large majority of scions belong to a single ancient cultivar containing somatic mutations, and that the widespread occurrence of one sucker genotype may imply rootstock selection. For the majority of grafted trees it seems likely that saplings were used as rootstocks; their genetic diversity probably is best explained as the result of a long history of sexual reproduction involving cultivated, feral and wild genotypes. PMID:24886387

  10. Near-shore and off-shore habitat use by endangered juvenile Lost River and Shortnose Suckers in Upper Klamath Lake, Oregon: 2006 data summary

    USGS Publications Warehouse

    Burdick, Summer M.; Wilkens, Alexander X.; VanderKooi, Scott P.

    2008-01-01

    We continued sampling juvenile suckers in 2006 as part of an effort to develop bioenergetics models for juvenile Lost River and shortnose suckers. This study required us to collect fish to determine growth rates and energy content of juvenile suckers. We followed the sampling protocols and methods described by Hendrixson et al. (2007b) to maintain continuity and facilitate comparisons with data collected in recent years, but sampled at a reduced level of effort compared to previous years (approximately one-third) due to limited funding. Here we present a summary of catch data collected in 2006. Bioenergetics models will be reported separately

  11. Preliminary testing of the role of exercise and predator recognition for bonytail and razorback sucker

    USGS Publications Warehouse

    Mueller, Gordon A.; Carpenter, Jeanette; Krapfel, Robert; Figiel, Chester

    2007-01-01

    Razorback suckers exercised (treatment) in water current (<0.3 m/s) for 10 weeks exhibited greater swimming stamina than unexercised, control fish. When exercised and unexercised razorback suckers were placed together with large predators in 2006, treatment fish had significantly fewer (n = 9, z = 1.69, p = 0.046) mortalities than control fish, suggesting increased stamina improved predator escape skills. Predator/prey tests comparing razorback suckers that had been previously exposed to a predation event with control fish, found treatment fish also had significantly fewer losses than predatornaïve fish (p = 0.017). Similar tests exposing predator-savvy and predator-naïve bonytail with largemouth bass showed a similar trend; predator-savvy bonytail suffered 38 percent fewer losses than control fish. However, there was not a statistically significant difference between the test groups (p = 0.143) due to small sample size. All exercise and predator exposure trials increased the survival rate of razorback sucker and bonytail compared to untreated counterparts.

  12. A new species of Petasiger (Digenea: Echinostomiformes: Echinostomatidae) in the brown pelican, Pelecanus occidentalis, (Aves: Pelecaniformes: Pelecanidae), from the Area de Conservación Guanacaste, Costa Rica.

    PubMed

    Zamparo, David; Overstreet, Robin M; Brooks, Daniel R

    2005-12-01

    A new species of Petasiger inhabits Pelecanus occidentalis, from the Area de Conservación Guanacaste, Costa Rica. The new species most closely resembles Petasiger novemdecim Lutz 1929 and Petasiger caribbensis Nassi, 1980 by having 19 circumoral spines and vitelline follicles confluent between the ventral sucker and gonads. The new species differs from both of these species by having a cirrus sac that is anteroposteriorly elongate and that reaches posteriorly to the midventral sucker, an ovary and Mehlis' gland that overlap the anterior testis dorsally, a uterus that lies dorsal to the ventral sucker, and a dextromedial genital pore. In P. novemdecim and P. caribbensis the cirrus sac is round and does not extend posteriorly to the anterior margin of the ventral sucker, both the ovary and Mehlis' gland are anterior to the anterior testis, the uterus runs lateral to and not dorsal to the ventral sucker, and the genital pore opens medially and sinistromedially, respectively.

  13. Distribution and condition of young-of-year Lost River and shortnose suckers in the Williamson River Delta restoration project and Upper Klamath Lake, Oregon, 2008-10--Final Report

    USGS Publications Warehouse

    Burdick, Summer M.; Hewitt, David A.

    2012-01-01

    We were unable to compare health and condition of juvenile suckers among habitats, due to their movement among habitats. However, documentation of length-weight relationships, afflictions and deformities, and histology indicated juvenile suckers captured in all habitats maintained a similar level of health among the 3 years of our study.

  14. Hazard assessment of selenium to endangered razorback suckers (Xyrauchen texanus)

    USGS Publications Warehouse

    Hamilton, S.J.; Holley, K.M.; Buhl, K.J.

    2002-01-01

    A hazard assessment was conducted based on information derived from two reproduction studies conducted with endangered razorback suckers (Xyrauchen texanus) at three sites near Grand Junction, CO, USA. Selenium contamination of the upper and lower Colorado River basin has been documented in water, sediment, and biota in studies by US Department of the Interior agencies and academia. Concern has been raised that this selenium contamination may be adversely affecting endangered fish in the upper Colorado River basin. The reproduction studies with razorback suckers revealed that adults readily accumulated selenium in various tissues including eggs, and that 4.6 μg/g of selenium in food organisms caused increased mortality of larvae. The selenium hazard assessment protocol resulted in a moderate hazard at the Horsethief site and high hazards at the Adobe Creek and North Pond sites. The selenium hazard assessment was considered conservative because an on-site toxicity test with razorback sucker larvae using 4.6 μg/g selenium in zooplankton caused nearly complete mortality, in spite of the moderate hazard at Horsethief. Using the margin of uncertainty ratio also suggested a high hazard for effects on razorback suckers from selenium exposure. Both assessment approaches suggested that selenium in the upper Colorado River basin adversely affects the reproductive success of razorback suckers.

  15. Hazard assessment of selenium to endangered razorback suckers (Xyrauchen texanus).

    PubMed

    Hamilton, Steven J; Holley, Kathleen M; Buhl, Kevin J

    2002-05-27

    A hazard assessment was conducted based on information derived from two reproduction studies conducted with endangered razorback suckers (Xyrauchen texanus) at three sites near Grand Junction, CO, USA. Selenium contamination of the upper and lower Colorado River basin has been documented in water, sediment, and biota in studies by US Department of the Interior agencies and academia. Concern has been raised that this selenium contamination may be adversely affecting endangered fish in the upper Colorado River basin. The reproduction studies with razorback suckers revealed that adults readily accumulated selenium in various tissues including eggs, and that 4.6 microg/g of selenium in food organisms caused increased mortality of larvae. The selenium hazard assessment protocol resulted in a moderate hazard at the Horsethief site and high hazards at the Adobe Creek and North Pond sites. The selenium hazard assessment was considered conservative because an on-site toxicity test with razorback sucker larvae using 4.6 microg/g selenium in zooplankton caused nearly complete mortality, in spite of the moderate hazard at Horsethief. Using the margin of uncertainty ratio also suggested a high hazard for effects on razorback suckers from selenium exposure. Both assessment approaches suggested that selenium in the upper Colorado River basin adversely affects the reproductive success of razorback suckers.

  16. Distribution of Rio Grande cutthroat trout and its co-occurrence with the Rio Grande sucker and Rio Grande chub on the Carson and Santa Fe National Forests

    Treesearch

    Bob Calamusso; John N. Rinne

    1996-01-01

    Studies were initiated in June, 1994 by the USDA Forest Service, Rocky Mountain Forest and Range Experiment Station to update knowledge on the distribution of the Rio Grande cutthroat trout a Forest Service Sensitive Species, and its co-occurrence with two native cypriniforms, Rio Grande sucker and Rio Grande Chub. The Rio Grande sucker IS listed as endangered by the...

  17. An introduced and a native vertebrate hybridize to form a genetic bridge to a second native species

    USGS Publications Warehouse

    McDonald, D.B.; Parchman, T.L.; Bower, M.R.; Hubert, W.A.; Rahel, F.J.

    2008-01-01

    The genetic impacts of hybridization between native and introduced species are of considerable conservation concern, while the possibility of reticulate evolution affects our basic understanding of how species arise and shapes how we use genetic data to understand evolutionary diversification. By using mitochondrial NADH dehydrogenase subunit 2 (ND2) sequences and 467 amplified fragment-length polymorphism nuclear DNA markers, we show that the introduced white sucker (Catostomus commersoni) has hybridized with two species native to the Colorado River Basin - the flannelmouth sucker (Catostomus latipinnis) and the bluehead sucker (Catostomus discobolus). Hybrids between the flannelmouth sucker and white sucker have facilitated introgression between the two native species, previously isolated by reproductive barriers, such that individuals exist with contributions from all three genomes. Most hybrids had the mitochondrial haplotype of the introduced white sucker, emphasizing its pivotal role in this three-way hybridization. Our findings highlight how introduced species can threaten the genetic integrity of not only one species but also multiple previously reproductively isolated species. Furthermore, this complex three-way reticulate (as opposed to strictly bifurcating) evolution suggests that seeking examples in other vertebrate systems might be productive. Although the present study involved an introduced species, similar patterns of hybridization could result from natural processes, including stream capture or geological formations (e.g., the Bering land bridge). ?? 2008 by The National Academy of Sciences of the USA.

  18. Commentary: selenium study on endangered razorback sucker is flawed

    USGS Publications Warehouse

    Hamilton, Steven J.

    2005-01-01

    The razorback sucker (Xyrauchen texanus) is listed as federally endangered throughout its range. A massive recovery effort by the Recovery Implementation Program for Endangered Fish Species in the Upper Colorado River Basin has focused its efforts in the upper Colorado River. The upper Colorado River basin also has two locations that have been identified by the National Irrigation Water Quality Program as having substantial selenium contamination. Selenium is toxic to fishes, affecting reproductive success. Thus, there is concern about potential effects of selenium on the endangered razorback sucker. Two sets of studies have investigated the effects of selenium on razorback suckers, but study results are conflicting. This commentary evaluates studies that claim selenium is not a problem for razorback sucker. We find that study bias was so pervasive that purported conclusions were unwarranted. Contaminated control water, older life stages of fish tested, lack of methodology for analysis of selenium in water, diet, or fish, use of rotifer food, low feeding rates, low growth rates of fish, and improper storage of site waters resulted in an apparent erroneous linkage of high selenium in whole-body residues with no adverse effects.

  19. Surface ultrastructure of juvenile and adult Acanthoparyphium tyosenense (Digenea: Echinostomatidae).

    PubMed

    Han, Eun-Taek; Choi, Moon-Seok; Choi, Sung-Yil; Chai, Jong-Yil

    2011-12-01

    The tegumental ultrastructure of juvenile and adult Acanthoparyphium tyosenense (Digenea: Echinostomatidae) was observed by scanning electron microscopy. One- to 3-day-old juveniles and 10-day-old adults were harvested from chicks experimentally fed metacercariae from a bivalve, Mactra veneriformis. The juvenile worms were minute, curved ventrally, and had 23 collar spines characteristically arranged in a single row. The lips of the oral sucker had 7 single aciliated sensory papillae and 4 grouped uniciliated sensory papillae. The ventral sucker had 25 aciliated round swellings on its lip. The anterolateral surface between the 2 suckers was densely packed with tongue-shaped tegumental spines, and the ventral surface just posterior to the ventral sucker was covered with peg-like spines. Retractile, peg-like spines were seen on the anterolateral surface, whereas scale-like spines with round tips and broad bases were sparsely distributed posterior to the ventral sucker. The cirrus was characteristically protruding and armed with minute spines. The surface ultrastructure of A. tyosenense was unique, especially in the number and arrangement of collar spines, shape, and distribution of tegumental spines and in distribution of sensory papillae.

  20. Unveiling the morphology of the acetabulum in octopus suckers and its role in attachment

    PubMed Central

    Tramacere, Francesca; Pugno, Nicola M.; Kuba, Michael J.; Mazzolai, Barbara

    2015-01-01

    In recent years, the attachment mechanism of the octopus sucker has attracted the interest of scientists from different research areas, including biology, engineering, medicine and robotics. From a technological perspective, the main goal is to identify the underlying mechanisms involved in sucker attachment for use in the development of new generations of artificial devices and materials. Recently, the understanding of the morphology of the sucker has been significantly improved; however, the mechanisms that allow attachment remain largely unknown. In this work, we present new anatomical findings: specifically, a protuberance in the acetabular roof in five different octopus species; previously, this protuberance was identified by the authors in Octopus vulgaris. Moreover, we discuss the role of the protuberance and other anatomical structures in attachment with minimal energy consumption. PMID:25657834

  1. 7 CFR 29.3558 - Type 35.

    Code of Federal Regulations, 2011 CFR

    2011-01-01

    ... INSPECTION Standards Official Standard Grades for Dark Air-Cured Tobacco (u.s. Types 35, 36, 37 and Foreign..., Kentucky-Tennessee-Indiana One Sucker, or Dark Air-cured One Sucker, including the upper Cumberland...

  2. 7 CFR 29.3558 - Type 35.

    Code of Federal Regulations, 2010 CFR

    2010-01-01

    ... INSPECTION Standards Official Standard Grades for Dark Air-Cured Tobacco (u.s. Types 35, 36, 37 and Foreign..., Kentucky-Tennessee-Indiana One Sucker, or Dark Air-cured One Sucker, including the upper Cumberland...

  3. 7 CFR 29.3558 - Type 35.

    Code of Federal Regulations, 2014 CFR

    2014-01-01

    ... INSPECTION Standards Official Standard Grades for Dark Air-Cured Tobacco (u.s. Types 35, 36, 37 and Foreign..., Kentucky-Tennessee-Indiana One Sucker, or Dark Air-cured One Sucker, including the upper Cumberland...

  4. 7 CFR 29.3558 - Type 35.

    Code of Federal Regulations, 2012 CFR

    2012-01-01

    ... INSPECTION Standards Official Standard Grades for Dark Air-Cured Tobacco (u.s. Types 35, 36, 37 and Foreign..., Kentucky-Tennessee-Indiana One Sucker, or Dark Air-cured One Sucker, including the upper Cumberland...

  5. 7 CFR 29.3558 - Type 35.

    Code of Federal Regulations, 2013 CFR

    2013-01-01

    ... INSPECTION Standards Official Standard Grades for Dark Air-Cured Tobacco (u.s. Types 35, 36, 37 and Foreign..., Kentucky-Tennessee-Indiana One Sucker, or Dark Air-cured One Sucker, including the upper Cumberland...

  6. Characterization of a novel hepadnavirus in the white sucker (Catostomus commersonii) from the Great Lakes Region of the USA

    USGS Publications Warehouse

    Hahn, Cassidy M.; Iwanowicz, Luke R.; Cornman, Robert S.; Conway, Carla M.; Winton, James R.; Blazer, Vicki S.

    2015-01-01

    The white sucker Catostomus commersonii is a freshwater teleost often utilized as a resident sentinel. Here, we sequenced the full genome of a hepatitis B-like virus that infects white suckers from the Great Lakes Region of the USA. Dideoxysequencing confirmed the white sucker hepatitis B virus (WSHBV) has a circular genome (3542 bp) with the prototypical codon organization of hepadnaviruses. Electron microscopy demonstrated that complete virions of approximately 40 nm were present in the plasma of infected fish. Compared to avi- and orthohepadnaviruses, sequence conservation of the core, polymerase and surface proteins was low and ranged from 16-27% at the amino acid level. An X protein homologue common to the orthohepadnaviruses was not present. The WSHBV genome included an atypical, presumptively non-coding region absent in previously described hepadnaviruses. Phylogenetic analyses confirmed WSHBV as distinct from previously documented hepadnaviruses. The level of divergence in protein sequences between WSHBV other hepadnaviruses, and the identification of an HBV-like sequence in an African cichlid provide evidence that a novel genus of the family Hepadnaviridae may need to be established that includes these hepatitis B-like viruses in fishes. Viral transcription was observed in 9.5% (16 of 169) of white suckers evaluated. The prevalence of hepatic tumors in these fish was 4.9%, of which only 2.4% were positive for both virus and hepatic tumors. These results are not sufficient to draw inferences regarding the association of WSHBV and carcinogenesis in white sucker.

  7. Small nonnative fishes as predators of larval razorback suckers

    USGS Publications Warehouse

    Carpenter, J.; Mueller, G.A.

    2008-01-01

    The razorback sucker (Xyrauchen texanus), an endangered big-river fish of the Colorado River basin, has demonstrated no sustainable recruitment in 4 decades, despite presence of spawning adults and larvae. Lack of adequate recruitment has been attributed to several factors, including predation by nonnative fishes. Substantial funding and effort has been expended on mechanically removing nonnative game fishes, typically targeting large predators. As a result, abundance of larger predators has declined, but the abundance of small nonnative fishes has increased in some areas. We conducted laboratory experiments to determine if small nonnative fishes would consume larval razorback suckers. We tested adults of three small species (threadfin shad, Dorosoma petenense; red shiner, Cyprinella lutrensis; fathead minnow, Pimephales promelas) and juveniles of six larger species (common carp, Cyprinus carpio; yellow bullhead, Ameiurus natalis; channel catfish, Ictalurus punctatus; rainbow trout, Oncorhynchus mykiss; green sunfish, Lepomis cyanellus; bluegill, L. macrochirus). These nonnative fishes span a broad ecological range and are abundant within the historical range of the razorback sucker. All nine species fed on larval razorback suckers (total length, 9-16 mm). Our results suggest that predation by small nonnative fishes could be responsible for limiting recovery of this endangered species.

  8. Tagging age-1 Lost River and shortnose suckers with passive integrated transponders, Upper Klamath Lake, Oregon–Summary of 2009–11 effort

    USGS Publications Warehouse

    Burdick, Summer M.

    2012-01-01

    A passive integrated transponder (PIT) tagging study was initiated in 2009 for age-1 endangered Lost River and shortnose suckers in Upper Klamath Lake, Oregon, for the purpose of examining causes of mortality, validating estimated age to maturity, and examining movement patterns. This study, which was done opportunistically in 2009 and 2010, received funding in 2011 for a directed tagging effort. Tags were redetected using an existing infrastructure of remote PIT tag readers and tag scanning surveys at American white pelican and double-crested cormorant breeding and loafing areas. Individual fish histories are used to describe the distance, direction, and timing of age-1 sucker movement. Sucker PIT tag detections in the Sprague and Williamson rivers in mid-summer and in autumn indicate age-1 suckers use these tributaries outside of the known spring spawning season. PIT tags detected in bird habitats indicate predation by birds may have been a cause of mortality in 2009. Field conditions prevented scanning bird breeding and loafing areas in Upper Klamath Wildlife National Refuge for tags in 2011, however, limiting our ability to make inferences about bird predation in those years.

  9. Survival, movement, and health of hatchery-raised juvenile Lost River suckers within a mesocosm in Upper Klamath Lake, Oregon

    USGS Publications Warehouse

    Hereford, Danielle M.; Burdick, Summer M.; Elliott, Diane G.; Dolan-Caret, Amari; Conway, Carla M.; Harris, Alta C.

    2016-01-28

    The recovery of endangered Lost River suckers (Deltistes luxatus) in Upper Klamath Lake is limited by poor juvenile survival and failure to recruit into the adult population. Poor water quality, degradation of rearing habitat, and toxic levels of microcystin are hypothesized to contribute to low juvenile survival. Studies of wild juvenile suckers are limited in that capture rates are low and compromised individuals are rarely captured in passive nets. The goal of this study was to assess the use of a mesocosm for learning about juvenile survival, movement, and health. Hatchery-raised juvenile Lost River suckers were PIT (passive integrated transponder) tagged and monitored by three vertically stratified antennas. Fish locations within the mesocosm were recorded at least every 30 minutes and were assessed in relation to vertically stratified water-quality conditions. Vertical movement patterns were analyzed to identify the timing of mortality for each fish. Most mortality occurred from July 28 to August 16, 2014. Juvenile suckers spent daylight hours near the benthos and moved throughout the entire water column during dark hours. Diel movements were not in response to dissolved-oxygen concentrations, temperature, or pH. Furthermore, low dissolved-oxygen concentrations, high temperatures, high pH, high un-ionized ammonia, or high microcystin levels did not directly cause mortality, although indirect effects may have occurred. However, water-quality conditions known to be lethal to juvenile Lost River suckers did not occur during the study period. Histological assessment revealed severe gill hyperplasia and Ichthyobodo sp. infestations in most moribund fish. For these fish, Ichthyobodo sp. was likely the cause of mortality, although it is unclear if this parasite originated in the rearing facility because fish were not screened for this parasite prior to introduction. This study has demonstrated that we can effectively use a mesocosm equipped with antennas to learn about the timing of mortality, movement, and health of PIT-tagged hatchery-raised juvenile Lost River suckers.

  10. The Causes and Prevention Measures of Stuck Pump Phenomenon of Rod-pumped Well in CBM Field

    NASA Astrophysics Data System (ADS)

    Yonggui, Mei

    2018-02-01

    In the process of CBM field exploitation, in order to realize the drainage equipment to work continuous stably, the article pays attention to study and solve the stuck pump problem, and aim of reducing reservoir damage and lowing production costs. Through coal particles stuck pump experiment and sediment composition analysis, we find out five primary cause of stuck pump phenomenon: sand from coal seam, sediment from ground, iron corrosion, iron scrap caused by eccentric wear, coal cake. According to stuck pump mechanism, the article puts forward 8 measures to prevent stuck pump phenomenon, and the measures are focused on technology optimization, operation management and drainage process control. After 7 years production practice, the yearly stuck pump rate has dropped from 8.9% to 1.2%, and the pump inspection period has prolonged 2 times. The experiment result shows that pure coal particles cannot cause stuck pump, but sand, scrap iron, and iron corrosion are the primary cause of stuck pump. The article study and design the new pipe string structure that the bottom of the pipe string is open. This kind of pipe string applied the sedimentation terminal velocity theory to solve the stuck pump phenomenon, and it can be widely used in CBM drainage development.

  11. Effect of wave action on near-well zone cleaning

    NASA Astrophysics Data System (ADS)

    Pen'kovskii, V. I.; Korsakova, N. K.

    2017-10-01

    Drilling filtrate invasion into the producing formation and native water accumulating of the near-well zone in well operation reduce the well productivity. As a result of that, depending on characteristic capillary pressure scale and differential pressure drawdown, oil production rate may become lower than expected one. In this paper, it is considered the hysteresis effects of capillary pressure after reversion of displacement. As applied to laboratory experiment conditions, the solution of problem of oil flow in formation model with a pressure drop on the model sides harmonically varied with time is presented. It was estimated a range of fluid vibration effective action on the near-well zone cleaning from capillary locking water. The plant simulating extraction of oil from formation using widely practised sucker-rod pump has been created. Formation model is presented as a slot filled with broken glass between two plates. In the process, natural oil and sodium chloride solution were used as working fluids. The experiments qualitatively confirm a positive effect of jack pumps on the near-well zone cleaning.

  12. Selenium concentrations in the razorback sucker (Xyrauchen texanus): Substitution of non-lethal muscle plugs for muscle tissue in contaminant assessment

    USGS Publications Warehouse

    Waddell, B.; May, T.

    1995-01-01

    A single muscle plug was collected from each of 25 live razorback suckers inhabiting the Colorado River basin and analyzed for selenium by instrumental neutron activation. Eight fish from Ashley Creek and three from Razorback Bar exhibited selenium concentrations exceeding 8 μg/g, a level associated with reproductive failure in fish. Concentrations of selenium in eggs and milt were significantly correlated with selenium concentrations in muscle plugs and together indicate a possible explanation for the decline of this species in the Colorado River basin. Muscle plugs (<50mg) and muscle tissue (20 g) were collected from dorsal, anterior, and posterior areas of common carp, flannelmouth sucker, and an archived razorback sucker and analyzed for selenium. Concentrations of selenium in muscle plugs were significantly correlated with selenium concentrations in muscle tissue from the same location and fish (r=0.97). Coefficients of variation for selenium concentrations in each fish were <6.5% for muscle tissue, but ranged from 1.5 to 32.4% for muscle plugs. Increased variation in muscle plugs was attributed to lower selenium concentrations found in the anterior muscle plugs of flannelmouth suckers. Mean selenium concentrations in muscle plugs and tissue from dorsal and posterior areas and muscle tissue from the anterior area were not significantly different. The non-lethal collection of a muscle plug from dorsal and posterior areas of the razorback sucker and other fish species may provide an accurate assessment of selenium concentrations that exist in adjacent muscle tissue.

  13. A review of possible causes of nutrient enrichment and decline of endangered sucker populations in Upper Klamath Lake, Oregon

    USGS Publications Warehouse

    Bortleson, Gilbert C.; Fretwell, Marvin O.

    1993-01-01

    Ten possible causes for this excessive enrichment in nutrients are described. Three of these hypotheses are suggested for immediate testing because of large-scale changes in nutrient loading that may have occurred as a result of man’s activities. These three hypotheses relate nutrient enrichment to (1) conversion of marshland to agricultural land, (2) agricultural drainage from the basin, and (3) reservoir regulation. Eleven possible hypothetical causes for the decline in sucker populations also are described. The decline in sucker population may be related to excessive nutrient enrichment (eutrophication) of the lake.

  14. Tegumental ultrastructure of the juvenile and adult Himasthla alincia (Digenea: Echinostomatidae)

    PubMed Central

    Han, Eun-Taek; Han, Kye-Young

    2003-01-01

    The tegumental ultrastructure of juvenile and adult Himasthla alincia (Digenea: Echinostomatidae) was observed by scanning electron microscopy. One-, 5- (juveniles) and 20-day-old worms (adults) were harvested from chicks experimentally fed metacercariae from a bivalve, Mactra veneriformis. The juvenile worms were elongated and curved ventrally. The head crown bore 31 collar spines, arranged in a single row. The lip of the oral sucker had 12 paired, and 3 single type I sensory papillae, and the ventral sucker had about 25 type II sensory papillae. The anterolateral surface between the two suckers was densely packed with tegumental spines with 4-7 pointed tips. The adult worms were more elongated and filamentous, and had severe transverse folds over the whole body surface. On the head crown and two suckers, type I and II sensory papillae were more densely distributed than in the juvenile worms. Retractile brush-like spines, with 8-10 digits, were seen on the anterolateral surface, whereas claw-shaped spines, with 2-5 digits, were sparsely distributed posteriorly to the ventral sucker. The cirrus characteristically protruded out, and was armed with small spines distally. The surface ultrastructure of H. alincia was shown to be unique among echinostomes, especially in the digitation of its tegumental spines, the distribution of sensory papillae and by severe folds of the tegument. PMID:12666726

  15. A comparison of age, size, and fecundity of harvested and reference White Sucker populations

    USGS Publications Warehouse

    Begley, Meg; Coghlan, Stephen M.; Zydlewski, Joseph D.

    2017-01-01

    White Suckers Catostomus commersonii are an important source of fresh bait for the Maine lobster fishery. The Maine Department of Inland Fisheries and Wildlife began issuing commercial harvest permits in 1991, without reporting requirements or limits on the number of permits. There is recent concern that overfishing may be occurring. To infer impact, we investigated demographic differences between White Sucker populations in lakes open to harvest and those in lakes closed to harvest. Each of three harvested lakes was paired to a nearby closed lake as a reference based on general size, morphometry, and information on harvest pressure. In total, 976 spawning White Suckers were collected from the six lakes in 2014 (120–282 individuals/lake). Fish size, estimated age, fecundity, and mortality rates were compared between lakes. We hypothesized that we would find smaller, younger, and less-fecund individuals in harvested lakes compared to reference lakes. Size and age distributions for both sexes differed between nearly all lake pairs (except between males from one pair). White Suckers from reference lakes were larger and older and had greater gonadosomatic indices and fecundity than fish from harvested lakes. Estimated annual mortality rates were at least twofold higher in harvested lakes than in reference lakes. We detected some differences in von Bertalanffy growth parameters between lake pairs, as might occur under selective harvest pressure. The growth coefficient was smaller for reference lakes than for harvested lakes, while asymptotic length was greater for reference lakes than for harvested lakes. The data suggest that current levels of exploitation are resulting in greater age truncation in existing White Sucker populations.

  16. Hazard assessment of selenium and other trace elements in wild larval razorback sucker from the Green River, Utah

    USGS Publications Warehouse

    Hamilton, S.J.; Muth, R.T.; Waddell, B.; May, T.W.

    2000-01-01

    Contaminant investigations of the Green River in northeastern Utah have documented selenium contamination at sites receiving irrigation drainage. The Green River provides critical habitat for four endangered fishes including the largest extant riverine population of endangered razorback sucker. Although 2175 larval razorback suckers were collected from the river between 1992 and 1996, very few juveniles have been captured within recent decades. Selenium concentrations were measured in larval razorback suckers collected from five sites in the Green River (Cliff Creek, Stewart Lake Drain, Sportsman's Drain, Greasewood Corral, and Old Charlie Wash) to assess the potential for adverse effects on recruitment of larvae to the juvenile stage and the adult population. Larvae from all sites contained mean selenium concentrations ranging from 4.3 to 5.8 ??g/g. These values were at or above the proposed toxic threshold of 4 ??g/g for adverse biological effects in fish, which was derived from several laboratory and field studies with a wide range of fish species. At two sites, Cliff Creek and Stewart Lake Drain, selenium concentrations in larvae increased over time as fish grew, whereas selenium concentrations decreased as fish grew at Sportsman's Drain. Evaluation of a 279-larvae composite analyzed for 61 elements demonstrated that selenium and, to a lesser extent, vanadium were elevated to concentrations reported to be toxic to a wide range of fish species. Elevated selenium concentrations in larval razorback suckers from the five sites suggest that selenium contamination may be widespread in the Green River, and that survival and recruitment of larvae to the juvenile stage may be limited due to adverse biological effects. Selenium contamination may be adversely affecting the reproductive success and recruitment of endangered razorback sucker.

  17. Physiological differences between root suckers and saplings enlarge the regeneration niche in Eucryphia cordifolia Cav.

    PubMed

    Escandón, Antonio B; Rojas, Roke; Morales, Loreto V; Corcuera, Luis J; Coopman, Rafael E; Paula, Susana

    2018-01-01

    Many clonal plants produce vegetative recruits that remain connected to the parent plant. Such connections permit resource sharing among ramets, explaining the high survival rates of vegetative recruits during establishment under suboptimal conditions for sexual regeneration. We propose that differences in the regeneration niches of sexual and vegetative recruits reflect different physiological adjustments caused by parental supply of resources to the ramets. We conducted ecophysiological measurements in saplings and root suckers of Eucryphia cordifolia Cav., a tree species of the temperate rainforest of southern South America. We compared the following traits of saplings and suckers: gas exchange at the leaf level, crown architecture, daily crown carbon balance, biomass allocation to above-ground tissues (leaf-to-stem mass ratio, leaf mass area and leaf area ratio), xylem anatomy traits (lumen vessel fraction, vessel density and size) and stem ring width. We also correlated the growth rates of saplings and suckers with relevant environmental data (light and climate). Saplings showed morphological, architectural and physiological traits that enhance daily crown carbon balance and increase water-use efficiency, in order to supply their growth demands while minimizing water loss per unit of carbon gained. The radial growth of saplings diminished under dry conditions, which suggests a strong stomatal sensitivity to water availability. Suckers have low stomatal conductance, likely because the carbon supplied by the parent plant diminishes the necessity of high rates of photosynthesis. The low responsiveness of sucker growth to temporal changes in water availability also supports the existence of parental supply. The physiological differences between sexual and vegetative recruits satisfactorily explain the ecological niche of E. cordifolia, with saplings restricted to more closed and humid sites. © The Author 2017. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  18. CHANGES IN GENETIC DIVERSITY OF A WHITE SUCKER POPULATION FOLLOWING EXPERIMENTAL WHOLE-LAKE ACIDIFICATION

    EPA Science Inventory

    Despite great strides to reduce sulfur dioxide emissions over the last decade, acid precipitation remains a persistent threat to North American fish communities. A demographic analysis of white suckers in an experimentally acidified oligotrophic lake in northwest Ontario demonst...

  19. Demographics and run timing of adult Lost River (Deltistes luxatus) and shortnose (Chasmistes brevirostris) suckers in Upper Klamath Lake, Oregon, 2011

    USGS Publications Warehouse

    Hewitt, David A.; Janney, Eric C.; Hayes, Brian S.; Harris, Alta C.

    2012-01-01

    Despite relatively high survival in most years, both species have experienced substantial declines in the abundance of spawning fish because losses from mortality have not been balanced by recruitment of new individuals. Although capture-recapture data indicate substantial recruitment of new individuals into the adult spawning populations for SNS and river spawning LRS in some years, size data do not corroborate these estimates. In fact, fork length data indicate that all populations are largely comprised of fish that were present in the late 1990s and early 2000s. As a result, the status of the endangered sucker populations in Upper Klamath Lake remains worrisome, and the situation is most dire for shortnose suckers. Future investigations should explore the connections between sucker recruitment and survival and various environmental factors, such as water quality and disease. Our monitoring program provides a robust platform for estimating vital population parameters, evaluating the status of the populations, and assessing the effectiveness of conservation and recovery efforts.

  20. Preliminary juvenile Lost River and shortnose sucker investigations in Clear Lake, California--2011 pilot study summary

    USGS Publications Warehouse

    Burdick, Summer M.; Rasmussen, Josh

    2012-01-01

    Poor recruitment appears to limit the recovery of Lost River and shortnose sucker populations in Clear Lake Reservoir, California, but the cause is unknown. Adult suckers migrate up Willow Creek and its tributaries to spawn in some years, but low flow in Willow Creek may inhibit spawning migrations in other years. It is unclear whether spawning is successful, larvae survive, or juveniles persist to adulthood. Environmental variables associated with successful spawning or young-of-year survival have not been identified and early life history for these populations is poorly understood. The U.S. Geological Survey in cooperation with the U.S. Fish and Wildlife Service and Ruby Pipeline L.L.C. Corporation (El Paso, Tex.) initiated a study in 2011 to better understand juvenile sucker life history in Clear Lake Reservoir, and to identify constraints in the early life history that may limit recruitment to the adult spawning populations. This is a report on the 2011 pilot study for this project.

  1. Characterization of a Novel Hepadnavirus in the White Sucker (Catostomus commersonii) from the Great Lakes Region of the United States.

    PubMed

    Hahn, Cassidy M; Iwanowicz, Luke R; Cornman, Robert S; Conway, Carla M; Winton, James R; Blazer, Vicki S

    2015-12-01

    The white sucker Catostomus commersonii is a freshwater teleost often utilized as a resident sentinel. Here, we sequenced the full genome of a hepatitis B-like virus that infects white suckers from the Great Lakes Region of the United States. Dideoxy sequencing confirmed that the white sucker hepatitis B virus (WSHBV) has a circular genome (3,542 bp) with the prototypical codon organization of hepadnaviruses. Electron microscopy demonstrated that complete virions of approximately 40 nm were present in the plasma of infected fish. Compared to avi- and orthohepadnaviruses, sequence conservation of the core, polymerase, and surface proteins was low and ranged from 16 to 27% at the amino acid level. An X protein homologue common to the orthohepadnaviruses was not present. The WSHBV genome included an atypical, presumptively noncoding region absent in previously described hepadnaviruses. Phylogenetic analyses confirmed WSHBV as distinct from previously documented hepadnaviruses. The level of divergence in protein sequences between WSHBV and other hepadnaviruses and the identification of an HBV-like sequence in an African cichlid provide evidence that a novel genus of the family Hepadnaviridae may need to be established that includes these hepatitis B-like viruses in fishes. Viral transcription was observed in 9.5% (16 of 169) of white suckers evaluated. The prevalence of hepatic tumors in these fish was 4.9%, and only 2.4% of fish were positive for both virus and hepatic tumors. These results are not sufficient to draw inferences regarding the association of WSHBV and carcinogenesis in white sucker. We report the first full-length genome of a hepadnavirus from fishes. Phylogenetic analysis of this genome indicates divergence from genomes of previously described hepadnaviruses from mammalian and avian hosts and supports the creation of a novel genus. The discovery of this novel virus may better our understanding of the evolutionary history of hepatitis B-like viruses of other hosts. In fishes, knowledge of this virus may provide insight regarding possible risk factors associated with hepatic neoplasia in the white sucker. This may also offer another model system for mechanistic research. Copyright © 2015 Hahn et al.

  2. Characterization of a Novel Hepadnavirus in the White Sucker (Catostomus commersonii) from the Great Lakes Region of the United States

    PubMed Central

    Hahn, Cassidy M.; Cornman, Robert S.; Conway, Carla M.; Winton, James R.; Blazer, Vicki S.

    2015-01-01

    ABSTRACT The white sucker Catostomus commersonii is a freshwater teleost often utilized as a resident sentinel. Here, we sequenced the full genome of a hepatitis B-like virus that infects white suckers from the Great Lakes Region of the United States. Dideoxy sequencing confirmed that the white sucker hepatitis B virus (WSHBV) has a circular genome (3,542 bp) with the prototypical codon organization of hepadnaviruses. Electron microscopy demonstrated that complete virions of approximately 40 nm were present in the plasma of infected fish. Compared to avi- and orthohepadnaviruses, sequence conservation of the core, polymerase, and surface proteins was low and ranged from 16 to 27% at the amino acid level. An X protein homologue common to the orthohepadnaviruses was not present. The WSHBV genome included an atypical, presumptively noncoding region absent in previously described hepadnaviruses. Phylogenetic analyses confirmed WSHBV as distinct from previously documented hepadnaviruses. The level of divergence in protein sequences between WSHBV and other hepadnaviruses and the identification of an HBV-like sequence in an African cichlid provide evidence that a novel genus of the family Hepadnaviridae may need to be established that includes these hepatitis B-like viruses in fishes. Viral transcription was observed in 9.5% (16 of 169) of white suckers evaluated. The prevalence of hepatic tumors in these fish was 4.9%, and only 2.4% of fish were positive for both virus and hepatic tumors. These results are not sufficient to draw inferences regarding the association of WSHBV and carcinogenesis in white sucker. IMPORTANCE We report the first full-length genome of a hepadnavirus from fishes. Phylogenetic analysis of this genome indicates divergence from genomes of previously described hepadnaviruses from mammalian and avian hosts and supports the creation of a novel genus. The discovery of this novel virus may better our understanding of the evolutionary history of hepatitis B-like viruses of other hosts. In fishes, knowledge of this virus may provide insight regarding possible risk factors associated with hepatic neoplasia in the white sucker. This may also offer another model system for mechanistic research. PMID:26378165

  3. Effects of pool formation and flash flooding on relative abundance of young-of-year flannelmouth suckers in the Paria River, Arizona

    USGS Publications Warehouse

    Thieme, Michele L.; McIvor, Carole; Brouder, Mark J.; Hoffnagle, Timothy L.

    2001-01-01

    Flannelmouth sucker, Catostomus latipinnis, a fish endemic to the Colorado River basin in the western United States, appears to experience poor recruitment to adult size in the Colorado River, downstream of Glen Canyon Dam. Lack or impermanence of rearing areas for young-of-year (YOY) fish is hypothesized to be the problem. Knowing the importance of tributary mouths as rearing areas in other river systems, we studied use of the mouth of the Paria River, a tributary of the Colorado River, by YOY flannelmouth suckers, and the availability of rearing area in the mouth at different flow levels in the Colorado River in 1996 and 1997. We also examined the relationship between flash floods in the Paria River and catch-per-unit-effort (CPUE) of YOY in the Paria River between 1991 and 1996.Maximum mean daily discharge in the Paria River was inversely correlated with CPUE of YOY flannelmouth suckers (Spearman Rho=−0.9856, p=0.0003) during their critical rearing period (15 March–30 June). Thus, it appears that YOY flannelmouth suckers rear longer in the Paria River in years when flash flooding is minimal.Recruitment of YOY flannelmouth suckers at the Paria River may also be improved by enhancing pool formation during spring and summer rearing seasons. YOY flannelmouth sucker was captured in a pool created by high Colorado River flows (≥336 m3/s) that inundated the mouth of the Paria River during spring and summer, 1996. In 1997, high flows (about 550–750 m3/s) in the Colorado River during winter and spring initially inundated the Paria River and formed a pool in the mouth. However, these high flows eventually caused 0.5–1.0 m of suspended sediment from the incoming Paria River to deposit in the mouth. Thus, despite higher flows than 1996, the slackwater area formed only occasionally in 1997. Differences in pool formation between 1996 and 1997 demonstrate that pool formation cannot be inferred solely from Colorado River flows. 

  4. [Spatial organization of musculature in the Himasthla elongata cercaria (Trematoda: Echinostomatidae)].

    PubMed

    Krupenko, D Iu

    2011-01-01

    Somatic muscles (body-wall and "parenchyma" musculature), muscles of suckers, alimentary tract and excretory bladder of Himasthla elongata cercaria were investigated using fluorescent phalloidin labelling and confocal microscopy. The arrangement of body-wall muscles differs between the certain parts of cercarial body and appears to be the most complicated in the collar district. Among the body-wall musculature, we described U-shaped muscles, which have never been found previously in trematodes. Muscles of oral and ventral suckers are grouped into 6-7 independent layers. In some of those layers, they are arranged bilaterally, which contradicts the tradition to consider the sucker as radially symmetric.

  5. National Program for Inspection of Non-Federal Dams. Sucker Brook Dam (VT 00212), Richelieu River Basin, Town of Salisbury, Addison County, Vermont. Phase I Inspection Report.

    DTIC Science & Technology

    1980-02-01

    BENNINGTON STATE OF VERMONT APPROX. SCALE IN MILES SUCKER BROOK DAM LOCATION MAP GORDON E AINSWORTH & ASSOCIATES INC Fviil 20 SU6ARLOAF STSUHDEFEDMASS...New Hampshire and Vermont - Water Year 1977", USGS Water-Data Report NH-Vf-77-l, U.S. Geologi - 0 cal Survey, Boston, Ma., 1978. 20. "Climatological...34’.’ -_’_,’’- ".’’° , ’J LrLi . m""_"’"L % " , , " °. • .. RICHELIEU RIVER BASIN TOWN OF SALISBURY ADDISON COUNTY, VERMONT 0o (0 Ln SUCKER

  6. Development of 20 TaqMan assays differentiating the endangered shortnose and Lost River suckers

    USGS Publications Warehouse

    Hoy, Marshal S.; Ostberg, Carl O.

    2015-01-01

    Accurate species identification is vital to conservation and management of species at risk. Species identification is challenging when taxa express similar phenotypic characters and form hybrids, for example the endangered shortnose sucker (Chasmistes brevirostris) and Lost River sucker (Deltistes luxatus). Here, we developed 20 Taqman assays that differentiate these species (19 nuclear DNA and one mitochondrial DNA). Assays were evaluated in 160 young-of-the-year identified to species using meristic counts. Alleles were not fixed between species, but species were highly differentiated (F ST = 0.753, P < 0.001). The assays developed herein will be a valuable tool for resource managers.

  7. Abundance and ultrastructural diversity of neuronal gap junctions in the OFF and ON sublaminae of the inner plexiform layer of rat and mouse retina.

    PubMed

    Kamasawa, N; Furman, C S; Davidson, K G V; Sampson, J A; Magnie, A R; Gebhardt, B R; Kamasawa, M; Yasumura, T; Zumbrunnen, J R; Pickard, G E; Nagy, J I; Rash, J E

    2006-11-03

    Neuronal gap junctions are abundant in both outer and inner plexiform layers of the mammalian retina. In the inner plexiform layer (IPL), ultrastructurally-identified gap junctions were reported primarily in the functionally-defined and anatomically-distinct ON sublamina, with few reported in the OFF sublamina. We used freeze-fracture replica immunogold labeling and confocal microscopy to quantitatively analyze the morphologies and distributions of neuronal gap junctions in the IPL of adult rat and mouse retina. Under "baseline" conditions (photopic illumination/general anesthesia), 649 neuronal gap junctions immunogold-labeled for connexin36 were identified in rat IPL, of which 375 were photomapped to OFF vs. ON sublaminae. In contrast to previous reports, the volume-density of gap junctions was equally abundant in both sublaminae. Five distinctive morphologies of gap junctions were identified: conventional crystalline and non-crystalline "plaques" (71% and 3%), plus unusual "string" (14%), "ribbon" (7%) and "reticular" (2%) forms. Plaque and reticular gap junctions were distributed throughout the IPL. However, string and ribbon gap junctions were restricted to the OFF sublamina, where they represented 48% of gap junctions in that layer. In string and ribbon junctions, curvilinear strands of connexons were dispersed over 5 to 20 times the area of conventional plaques having equal numbers of connexons. To define morphologies of gap junctions under different light-adaptation conditions, we examined an additional 1150 gap junctions from rats and mice prepared after 30 min of photopic, mesopic and scotopic illumination, with and without general anesthesia. Under these conditions, string and ribbon gap junctions remained abundant in the OFF sublamina and absent in the ON sublamina. Abundant gap junctions in the OFF sublamina of these two rodents with rod-dominant retinas revealed previously-undescribed but extensive pathways for inter-neuronal communication; and the wide dispersion of connexons in string and ribbon gap junctions suggests unique structural features of gap junctional coupling in the OFF vs. ON sublamina.

  8. EXPLOITATION OF FLOODPLAIN RESOURCES BY ADULT LARGESCALE SUCKER OF THE WILLAMETTE RIVER, OREGON

    EPA Science Inventory

    We tested two predictions of the flood-pulse concept on a large, temperate alluvial river that historically flooded an extensive fringing floodplain. We predicted adult largescale sucker, Catostomus macrocheilus, would: (1) migrate onto the floodplain during high water; and (2) e...

  9. Genetic response of a white sucker population to experimental whole lake acidification

    EPA Science Inventory

    Background/Question/MethodsLake acidification can strongly impact the structure and function of lake ecosystems, causing extirpation of some species while other organisms are able adapt to changing pH. We followed the genetics of a population of white sucker (Catostomus commerso...

  10. Infection by a black spot-causing species of Uvulifer and associated opercular alterations in fishes from a high-desert stream in Wyoming

    USGS Publications Warehouse

    Quist, M.C.; Bower, M.R.; Hubert, W.A.

    2007-01-01

    Black spot is a common disease syndrome of freshwater fishes. This study provides information on the rank of density of the black spot agent and opercular bone alterations associated with at least one digenean, Uvulifer sp., infecting native and non-native catostomids and cyprinids of the Upper Colorado River Basin. We evaluated the density rank of pigmented metacercariae and associated alterations in the operculum of the bluehead sucker Catostomus discobolus, flannelmouth sucker C. latipinnis, white sucker C. commersoni, catostomid hybrids, roundtail chub Gila robusta, and creek chub Semotilus atromaculatus, sampled from Muddy Creek, Wyoming, USA in 2003 or 2004. All fish species contained individuals that exhibited gross signs of the black spot agent. Bluehead and flannelmouth suckers had 100% prevalence of infection. Although the other suckers and chubs contained encysted metacercariae in at least one individual, the presence of pigmented metacercariae was not apparent (i.e. based on gross observations) in many individuals. Catostomids had higher densities of metacercariae than cyprinids, as shown by frequency distributions of density ranks. Opercular holes (i.e. holes that completely penetrated the opercle and were in direct association with the pigment associated metacercariae) and pockets (depressions on the external surface of the opercle associated with metacercariae) were abundant among catostomids but rare among cyprinids. ?? Inter-Research 2007.

  11. Infection by a black spot-causing species of Uvulifer and associated opercular alterations in fishes from a high-desert stream in Wyoming.

    PubMed

    Quist, Michael C; Bower, Michael R; Hubert, Wayne A

    2007-12-13

    Black spot is a common disease syndrome of freshwater fishes. This study provides information on the rank of density of the black spot agent and opercular bone alterations associated with at least one digenean, Uvulifer sp., infecting native and non-native catostomids and cyprinids of the Upper Colorado River Basin. We evaluated the density rank of pigmented metacercariae and associated alterations in the operculum of the bluehead sucker Catostomus discobolus, flannelmouth sucker C. latipinnis, white sucker C. commersoni, catostomid hybrids, roundtail chub Gila robusta, and creek chub Semotilus atromaculatus, sampled from Muddy Creek, Wyoming, USA in 2003 or 2004. All fish species contained individuals that exhibited gross signs of the black spot agent. Bluehead and flannelmouth suckers had 100% prevalence of infection. Although the other suckers and chubs contained encysted metacercariae in at least one individual, the presence of pigmented metacercariae was not apparent (i.e. based on gross observations) in many individuals. Catostomids had higher densities of metacercariae than cyprinids, as shown by frequency distributions of density ranks. Opercular holes (i.e. holes that completely penetrated the opercle and were in direct association with the pigment associated metacercariae) and pockets (depressions on the external surface of the opercle associated with metacercariae) were abundant among catostomids but rare among cyprinids.

  12. Spatial and temporal use of a spawning site in the middle green river by wild and hatchery-reared razorback suckers

    USGS Publications Warehouse

    Modde, T.; Bowen, Z.H.; Kitcheyan, D.C.

    2005-01-01

    The population of endangered razorback suckers Xyrauchen texanus in the middle Green River (upper Colorado River basin) has declined during the last 40 years. The apparent cause for this decline is a lack of successful recruitment. This study used radiotelemetry to evaluate the ability of hatchery-reared razorback suckers to locate spawning areas where wild fish congregate during the ascending hydrographic limb of the snowmelt runoff. Hatchery-reared razorback suckers appeared to show similar reproductive behavior to wild fish. Both wild and hatchery-reared fish were found near the middle Green River spawning area between 1 and 25 May 2000. Hatchery fish occupied the same areas on the spawning site as wild fish, and remained on the spawning site during both nocturnal and diurnal hours. Males were more abundant on the spawning area than females, but the few females captured tended to stage away from the primary spawning area. The results from this study suggest hatchery-reared fish are capable of responding to natural cues that prompt spawning aggregations and are successful in locating existing spawning aggregations of wild fish. Given attention to stocking criteria, including genetic diversity and the size and time of stocking, the challenges of recovering razorback suckers will center on those factors that led to the population declines, particularly the survival of early life stages in off-channel habitats. ?? American Fisheries Society 2005.

  13. Lead-contaminated imported tamarind candy and children's blood lead levels.

    PubMed

    Lynch, R A; Boatright, D T; Moss, S K

    2000-01-01

    In 1999, an investigation implicated tamarind candy as the potential source of lead exposure for a child with a significantly elevated blood lead level (BLL). The Oklahoma City-County Health Department tested two types of tamarind suckers and their packaging for lead content. More than 50% of the tested suckers exceeded the US Food and Drug Administration (FDA) Level of Concern for lead in this type of product. The authors calculated that a child consuming one-quarter to one-half of either of the two types of suckers in a day would exceed the maximum FDA Provis onal Tolerable Intake for lead. High lead concentrations in the two types of wrappers suggested leaching as a potential source of contamination. The authors used the Environmental Protection Agency's Integrated Exposure Uptake Biokinetic (IEUBK) model to predict the effects of consumption of contaminated tamarind suckers on populat on BLLs. The IEUBK model predicted that consumption of either type of sucker at a rate of one per day would result in dramatic increases in mean BLLs for children ages 6-84 months in Oklahoma and in the percentage of children wth elevated BLLs (> or =10 micrograms per deciliter [microg/dL]). The authors conclude that consumption of these products represents a potential public health threat. In addition, a history of lead contamination in imported tamarind products suggests that import control measures may not be completely effective in preventing additional lead exposure.

  14. Patterns of Larval Sucker Emigration from the Sprague and Lower Williamson Rivers of the Upper Klamath Basin, Oregon, Prior to the Removal of Chiloquin Dam - 2006 Annual Report

    USGS Publications Warehouse

    Ellsworth, Craig M.; Tyler, Torrey J.; VanderKooi, Scott P.; Markle, Douglas F.

    2009-01-01

    In 2006, we collected larval Lost River sucker Deltistes luxatus (LRS), shortnose sucker Chasmistes brevirostris (SNS), and Klamath largescale sucker Catostomus snyderi (KLS) emigrating from spawning areas in the Williamson and Sprague Rivers. This work is part of a multi-year effort to characterize the relative abundance, drift timing, and length frequencies of larval suckers in this watershed prior to the removal of Chiloquin Dam on the lower Sprague River. Additional larval drift samples were collected from the Fremont Bridge on Lakeshore Drive on the south end of Upper Klamath Lake near its outlet to the Link River. Because of difficulties in distinguishing KLS larvae from SNS larvae, individuals identified as either of these two species were grouped together and reported as KLS-SNS in this report. We found that larval densities varied by site with the highest densities being collected at the most upstream site on the Sprague River at river kilometer (rkm) 108.0 near Beatty, Oregon (Beatty), and the most downstream sites near Chiloquin, Oregon; one site on the Sprague River at rkm 0.7 (Chiloquin) and the other site on the Williamson River at rkm 7.4 (Williamson). Larval catches were relatively small and sporadic at two other sites on the Sprague River located between Chiloquin and Beatty (Power Station at rkm 9.5 and Lone Pine at rkm 52.7) and one site on the Sycan River at rkm 4.7. Most larvae (79 percent) collected in 2006 were identified as LRS. More larvae and eggs were collected at Chiloquin than at any other site. The seasonal timing of larval drift varied by location; larvae generally were captured earlier at upstream sites than at downstream sites. Cumulative catch percentages of drifting larvae suggest that larval LRS emigrated earlier than KLS-SNS larvae at every site. Drift of LRS larvae at Beatty began 3 to 4 weeks earlier than at Chiloquin or Williamson. At Chiloquin, peak larval catches occurred 3 and 5 weeks after peak egg catches. The daily peak in larval drift at Chiloquin occurred approximately 1.5 to 2.0 hours after sunset. Nightly peak larval drift varied by location; larvae were captured earlier in the evening at sites closer to known spawning locations than sites farther away from these areas. The highest numerical catches of sucker-sized eggs were at Chiloquin indicating that this site is in close proximity to a spawning area. Numerical catches of older, more developed larval and juvenile suckers also were highest at Chiloquin. This may be due to the turbulent nature of this site, which could have swept larger fish into the drift. Proportional catches of older, more developed larval and juvenile suckers were highest at Sycan, Lone Pine, Power Station, and Fremont Bridge. This indicates these sites are located nearer to sucker nursery areas rather than spawning areas. Very few larval LRS were collected at Fremont Bridge at the south end of Upper Klamath Lake. Larval KLS-SNS densities at Fremont Bridge were the third highest of the seven sampling sites. Peak drift of larval KLS-SNS at Fremont Bridge occurred the week after peak drift of larval KLS-SNS at Williamson. Although inter-annual variation continues to appear in the larval drift data, our results continue to show consistent patterns of larval emigration in the drainage basin. In combination with data collected from the spawning movements and destinations of radio-tagged and PIT-tagged adult suckers, this larval drift data will provide a baseline standard by which to determine the effects of dam removal on the spawning distribution of endangered Klamath Basin suckers in the Sprague River.

  15. Assessment of Population Status for a White Sucker (Catostomus commersoni) Population Exposed to Bleached Kraft Pulp Mill Effluent

    EPA Science Inventory

    Credible ecological risk assessments often need to include analysis of population-level impacts. In the present study, a predictive model was developed to translate changes in the fecundity and the age structure of a breeding population of white sucker (Catostomus commersoni) co...

  16. Assessment of Population Status for a White Sucker (Catostomus commersoni) Population Exposed to Bleached Kraft Pulp Mill Effluent

    EPA Science Inventory

    A predictive model was developed to translate changes in the fecundity and the age structure of a breeding population of white sucker (Catostomus commersoni) collected in the field to alterations in population growth rate. Application of this density dependent population ...

  17. Bone characteristics and metal concentrations in white suckers (Catostomus commersoni) from one neutral and three acidified lakes in Maine

    USGS Publications Warehouse

    Hamilton, Steven J.; Haines, Terry A.

    1989-01-01

    The bone characteristics of white suckers, Catostomus commersoni, from four lakes in Maine were studied in relation to lake water quality and metal concentrations in fish. Green Lake had a neutral pH, high buffering capacity, and low aluminum concentrations, whereas the other three lakes had low pH, low buffering capacity, and elevated aluminum concentrations. The concentrations of aluminum in white suckers did not differ among the four lakes, but concentrations of cadmium, lead, and mercury were greater in fish from the three low-pH lakes. The vertebrae were weaker and more flexible in fish from the low-pH lakes than in those from neutral-pH Green Lake. The inferior mechanical properties of bone in fish from the low-pH lakes were probably a result of the significantly lower proline concentrations in collagen. Low pH of lake water or elevated whole-body concentrations of cadmium, lead, and mercury, either individually or combined, could have caused the inferior mechanical properties of bone of white suckers from the low-pH lakes.

  18. Tag loss and short-term mortality associated with passive integrated transponder tagging of juvenile Lost River suckers

    USGS Publications Warehouse

    Burdick, Summer M.

    2011-01-01

    Passive integrated transponder (PIT) tags are commonly used to mark small catostomids, but tag loss and the effect of tagging on mortality have not been assessed for juveniles of the endangered Lost River sucker Deltistes luxatus. I evaluated tag loss and short-term (34-d) mortality associated with the PIT tagging of juvenile Lost River suckers in the laboratory by using a completely randomized design and three treatment groups (PIT tagged, positive control, and control). An empty needle was inserted into each positive control fish, whereas control fish were handled but not tagged. Only one fish expelled its PIT tag. Mortality rate averaged 9.8 ± 3.4% (mean ± SD) for tagged fish; mortality was 0% for control and positive control fish. All tagging mortalities occurred in fish with standard lengths of 71 mm or less, and most of the mortalities occurred within 48 h of tagging. My results indicate that 12.45- × 2.02-mm PIT tags provide a viable method of marking juvenile Lost River suckers that are 72 mm or larger.

  19. Hydraulic tests with direct-push equipment

    USGS Publications Warehouse

    Butler, J.J.; Healey, J.M.; McCall, G.W.; Garnett, E.J.; Loheide, Steven P.

    2002-01-01

    The potential of direct-push technology for hydraulic characterization of saturated flow systems was investigated at a field site with a considerable degree of subsurface control. Direct-push installations were emplaced by attaching short lengths of screen (shielded and unshielded) to the bottom end of a tool string that was then advanced into the unconsolidated sediments. A series of constant-rate pumping tests were performed in a coarse sand and gravel aquifer using direct-push tool strings as observation wells. Very good agreement (within 4%) was found between hydraulic conductivity (K) estimates from direct-push installations and those from conventional wells. A program of slug tests was performed in direct-push installations using small-diameter adaptations of solid-slug and pneumatic methods. In a sandy silt interval of moderate hydraulic conductivity, K values from tests in a shielded screen tool were in excellent agreement (within 2%) with those from tests in a nearby well. In the coarse sand and gravel aquifer, K values were within 12% of those from multilevel slug tests at a nearby well. However, in the more permeable portions of the aquifer (K > 70 m/day), the smaller-diameter direct-push rods (0.016 m inner diameter [I.D.]) attenuated test responses, leading to an underprediction of K. In those conditions, use of larger-diameter rods (e.g., 0.038 m I.D.) is necessary to obtain K values representative of the formation. This investigation demonstrates that much valuable information can be obtained from hydraulic tests in direct-push installations. As with any type of hydraulic test, K estimates are critically dependent on use of appropriate emplacement and development procedures. In particular, driving an unshielded screen through a heterogeneous sequence will often lead to a buildup of low-K material that can be difficult to remove with standard development procedures.

  20. Lead-contaminated imported tamarind candy and children's blood lead levels.

    PubMed Central

    Lynch, R A; Boatright, D T; Moss, S K

    2000-01-01

    In 1999, an investigation implicated tamarind candy as the potential source of lead exposure for a child with a significantly elevated blood lead level (BLL). The Oklahoma City-County Health Department tested two types of tamarind suckers and their packaging for lead content. More than 50% of the tested suckers exceeded the US Food and Drug Administration (FDA) Level of Concern for lead in this type of product. The authors calculated that a child consuming one-quarter to one-half of either of the two types of suckers in a day would exceed the maximum FDA Provis onal Tolerable Intake for lead. High lead concentrations in the two types of wrappers suggested leaching as a potential source of contamination. The authors used the Environmental Protection Agency's Integrated Exposure Uptake Biokinetic (IEUBK) model to predict the effects of consumption of contaminated tamarind suckers on populat on BLLs. The IEUBK model predicted that consumption of either type of sucker at a rate of one per day would result in dramatic increases in mean BLLs for children ages 6-84 months in Oklahoma and in the percentage of children wth elevated BLLs (> or =10 micrograms per deciliter [microg/dL]). The authors conclude that consumption of these products represents a potential public health threat. In addition, a history of lead contamination in imported tamarind products suggests that import control measures may not be completely effective in preventing additional lead exposure. PMID:11354337

  1. Tumours in white suckers from Lake Michigan tributaries: Pathology and prevalence

    USGS Publications Warehouse

    Blazer, Vicki S.; Walsh, H.L.; Braham, R.P.; Hahn, C. M.; Mazik, P.; McIntyre, P.B.

    2016-01-01

    The prevalence and histopathology of neoplastic lesions were assessed in white suckerCatostomus commersonii captured at two Lake Michigan Areas of Concern (AOCs), the Sheboygan River and Milwaukee Estuary. Findings were compared to those observed at two non-AOC sites, the Root and Kewaunee rivers. At each site, approximately 200 adult suckers were collected during their spawning migration. Raised skin lesions were observed at all sites and included discrete white spots, mucoid plaques on the body surface and fins and large papillomatous lesions on lips and body. Microscopically, hyperplasia, papilloma and squamous cell carcinoma were documented. Liver neoplasms were also observed at all sites and included both hepatocellular and biliary tumours. Based on land use, the Kewaunee River was the site least impacted by human activities previously associated with fish tumours and had significantly fewer liver neoplasms when compared to the other sites. The proportion of white suckers with liver tumours followed the same patterns as the proportion of urban land use in the watershed: the Milwaukee Estuary had the highest prevalence, followed by the Root, Sheboygan and Kewaunee rivers. The overall skin neoplasm (papilloma and carcinoma) prevalence did not follow the same pattern, although the percentage of white suckers with squamous cell carcinoma exhibited a similar relationship to land use. Testicular tumours (seminoma) were observed at both AOC sites but not at the non-AOC sites. Both skin and liver tumours were significantly and positively associated with age but not sex.

  2. Characterization of plasma vitellogenin and sex hormone concentrations during the annual reproductive cycle of the endangered razorback sucker

    USGS Publications Warehouse

    Hinck, Jo Ellen; Papoulias, Diana M.; Annis, Mandy L.; Tillitt, Donald E.; Marr, Carrie; Denslow, Nancy D.; Kroll, Kevin J.; Nachtmann, Jason

    2011-01-01

    Population declines of the endangered razorback sucker Xyrauchen texanus in the Colorado River basin have been attributed to predation by and competition with nonnative fishes, habitat alteration, and dam construction. The reproductive health and seasonal variation of the reproductive end points of razorback sucker populations are currently unknown. Using nonlethal methods, we characterized the plasma hormonal fluctuations of reproductively mature female and male razorback suckers over a 12-month period in a hatchery by measuring their vitellogenin (VTG) and three sex hormones: 17β-estradiol (E2), testosterone (T), and 11-ketotestosterone (KT). Fish were identified as reproductive or nonreproductive based on their body weight, VTG, and sex hormone profiles. In reproductive females, the E2 concentration increased in the fall and winter, and increases in T and VTG concentrations were generally associated with the spawning period. Mean T concentrations were consistently greater in reproductive females than in nonreproductive females, but this pattern was even more pronounced during the spawning period (spring). Consistently low T concentrations (<3 ng/mL) in adult females during the spawning period may indicate reproductive impairment. In reproductive males, spring increases in KT and T concentrations were associated with spawning; concentrations of E2 (<0.48 ng/mL) and VTG (<0.001 mg/mL) were low in males throughout the study. In addition, the E2 : KT ratio and T were the best metrics by which to distinguish female from male adult razorback suckers throughout the year. These metrics of reproductive health and condition may be particularly important to recovery efforts of razorback suckers given that the few remaining wild populations are located in a river where water quality and quantity issues are well documented. In addition to the size, age, and recruitment information currently considered in the recovery goals of this endangered species, reproductive end points could be included as recovery metrics with which to monitor seasonal trends and determine whether repatriated populations are cycling naturally.

  3. Effects of Chiloquin Dam on spawning distribution and larval emigration of Lost River, shortnose, and Klamath largescale suckers in the Williamson and Sprague Rivers, Oregon

    USGS Publications Warehouse

    Martin, Barbara A.; Hewitt, David A.; Ellsworth, Craig M.

    2013-01-01

    Chiloquin Dam was constructed in 1914 on the Sprague River near the town of Chiloquin, Oregon. The dam was identified as a barrier that potentially inhibited or prevented the upstream spawning migrations and other movements of endangered Lost River (Deltistes luxatusChasmistes brevirostris) suckers, as well as other fish species. In 2002, the Bureau of Reclamation led a working group that examined several alternatives to improve fish passage at Chiloquin Dam. Ultimately it was decided that dam removal was the best alternative and the dam was removed in the summer of 2008. The U.S. Geological Survey conducted a long-term study on the spawning ecology of Lost River, shortnose, and Klamath largescale suckers (Catostomus snyderi) in the Sprague and lower Williamson Rivers from 2004 to 2010. The objective of this study was to evaluate shifts in spawning distribution following the removal of Chiloquin Dam. Radio telemetry was used in conjunction with larval production data and detections of fish tagged with passive integrated transponders (PIT tags) to evaluate whether dam removal resulted in increased utilization of spawning habitat farther upstream in the Sprague River. Increased densities of drifting larvae were observed at a site in the lower Williamson River after the dam was removed, but no substantial changes occurred upstream of the former dam site. Adult spawning migrations primarily were influenced by water temperature and did not change with the removal of the dam. Emigration of larvae consistently occurred about 3-4 weeks after adults migrated into a section of river. Detections of PIT-tagged fish showed increases in the numbers of all three suckers that migrated upstream of the dam site following removal, but the increases for Lost River and shortnose suckers were relatively small compared to the total number of fish that made a spawning migration in a given season. Increases for Klamath largescale suckers were more substantial. Post-dam removal monitoring only included 2 years with below average river discharge during the spawning season; data from years with higher flows may provide a different perspective on the effects of dam removal on the spawning migrations of the two endangered sucker species.

  4. Cooperative particle motion in complex (dusty) plasmas

    NASA Astrophysics Data System (ADS)

    Zhdanov, Sergey; Morfill, Gregor

    2014-05-01

    Strongly coupled complex (dusty) plasmas give us a unique opportunity to go beyond the limits of continuous media and study various generic processes occurring in liquids or solids at the kinetic level. A particularly interesting and challenging topic is to study dynamic cooperativity at local and intermediate scales. As an important element of self-organization, cooperative particle motion is present in many physical, astrophysical and biological systems. As a rule, cooperative dynamics, bringing to life 'abnormal' effects like enhanced diffusion, self-dragging, or self-propelling of particles, hold aspects of 'strange' kinetics. The synergy effects are also important. Such kind of cooperative behavior was evidenced for string-like formations of colloidal rods, dynamics of mono- and di-vacancies in 2d colloidal crystals. Externally manipulated 'dust molecules' and self-assembled strings in driven 3d particle clusters were other noticeable examples. There is a certain advantage to experiment with complex plasmas merely because these systems are easy to manipulate in a controllable way. We report on the first direct observation of microparticle cooperative movements occurring under natural conditions in a 2d complex plasma.

  5. Depth and Diameter of the Parent Roots of Aspen Root Suckers

    Treesearch

    Robert E. Farmer

    1962-01-01

    Studies of the Populus tremuloides root system by Day (1944), Sandberg (1951) and Barnes (1959) have all shown lateral roots extending as much as 30 feet from tree base. These roots may branch extensively and sometimes exhibit an "undulating" growth habit. According to the above authors, suckers occur on the segments of these lateral roots...

  6. Dynamics of aspen root biomass and sucker production following fire

    Treesearch

    Roy A. Renkin; Don G. Despain

    2001-01-01

    Changes in preburn aspen root biomass 8 years following prescribed fire were analyzed for five experimental sites distributed across a moisture gradient. Total root biomass decreased across all sites but was proportionately greater in xeric than mesic sites. Response of post-burn aspen suckers to ungulate browsing varied according to site and treatment. Browsing...

  7. Neuroethology: self-recognition helps octopuses avoid entanglement.

    PubMed

    Crook, Robyn J; Walters, Edgar T

    2014-06-02

    How an octopus performs complex movements of its eight sucker-studded arms without entanglement has been a mystery. A new study has found that self-recognition of the octopus's skin by its suckers inhibits reflexive grasping of its own arms, simplifying the mechanisms needed to generate intricate arm behavior. Copyright © 2014 Elsevier Ltd. All rights reserved.

  8. Status and trends of adult Lost River (Deltistes luxatus) and shortnose (Chasmistes brevirostris) sucker populations in Upper Klamath Lake, Oregon, 2014

    USGS Publications Warehouse

    Hewitt, David A.; Janney, Eric C.; Hayes, Brian S.; Harris, Alta C.

    2015-10-02

    Despite relatively high survival in most years, we conclude that both species have experienced substantial decreases in the abundance of spawning adults because losses from mortality have not been balanced by recruitment of new individuals. Although capture-recapture data indicate substantial recruitment of new individuals into the spawning populations for SNS and river spawning LRS in some years, size data do not corroborate these estimates. As a result, the status of the endangered sucker populations in Upper Klamath Lake remains worrisome, especially for shortnose suckers. Our monitoring program provides a robust platform for estimating vital population parameters, evaluating the status of the populations, and assessing the effectiveness of conservation and recovery efforts.

  9. Host specificity of Lepeophtheirus crassus (Wilson and Bere) (Copepoda: Caligidae) parasitic on the marlin sucker Remora osteochir (Cuvier) in the Atlantic Ocean.

    PubMed

    Ho, Ju-shey; Collete, Bruce B; Madinabeitia, Ione

    2006-10-01

    Three species of remoras--Remora brachyptera (Lowe), Remora osteochir (Cuvier), and Remora remora (Linnaeus)--were collected from 4 species of billfishes--Istiophorus platypterus (Shaw), Makaira nigricans Lacepéde, Tetrapturus albidus Poey, and Tetrapturus pfluegeri Robins and de Sylva--on board a Japanese long-liner Shoyo Maru during her cruise in 2002 across the Atlantic. However, only the marlin sucker (R. osteochir) was found to carry a parasitic copepod, Lepeophtheirus crassus (Wilson and Bere, 1936). Although 12 species of parasitic copepods have been reported from billfishes around the world ocean, none of them is L. crassus. Thus, L. crassus is considered a parasite specific to the marlin sucker.

  10. Reproductive health indicators of fishes from Pennsylvania watersheds: association with chemicals of emerging concern.

    PubMed

    Blazer, V S; Iwanowicz, D D; Walsh, H L; Sperry, A J; Iwanowicz, L R; Alvarez, D A; Brightbill, R A; Smith, G; Foreman, W T; Manning, R

    2014-10-01

    Fishes were collected at 16 sites within the three major river drainages (Delaware, Susquehanna, and Ohio) of Pennsylvania. Three species were evaluated for biomarkers of estrogenic/antiandrogenic exposure, including plasma vitellogenin and testicular oocytes in male fishes. Smallmouth bass Micropterus dolomieu, white sucker Catostomus commersonii, and redhorse sucker Moxostoma species were collected in the summer, a period of low flow and low reproductive activity. Smallmouth bass were the only species in which testicular oocytes were observed; however, measurable concentrations of plasma vitellogenin were found in male bass and white sucker. The percentage of male bass with testicular oocytes ranged from 10 to 100%, with the highest prevalence and severity in bass collected in the Susquehanna drainage. The percentage of males with plasma vitellogenin ranged from 0 to 100% in both bass and sucker. Biological findings were compared with chemical analyses of discrete water samples collected at the time of fish collections. Estrone concentrations correlated with testicular oocytes prevalence and severity and with the percentage of male bass with vitellogenin. No correlations were noted with the percentage of male sucker with vitellogenin and water chemical concentrations. The prevalence and severity of testicular oocytes in bass also correlated with the percent of agricultural land use in the watershed above a site. Two sites within the Susquehanna drainage and one in the Delaware were immediately downstream of wastewater treatment plants to compare results with upstream fish. The percentage of male bass with testicular oocytes was not consistently higher downstream; however, severity did tend to increase downstream.

  11. Reproductive health indicators of fishes from Pennsylvania watersheds: association with chemicals of emerging concern

    USGS Publications Warehouse

    Blazer, V.S.; Iwanowicz, D.D.; Walsh, H.L.; Sperry, A.J.; Iwanowicz, L.R.; Alvarez, D.A.; Brightbill, R.A.; Smith, G.; Foreman, W.T.; Manning, R.

    2014-01-01

    Fishes were collected at 16 sites within the three major river drainages (Delaware, Susquehanna, and Ohio) of Pennsylvania. Three species were evaluated for biomarkers of estrogenic/antiandrogenic exposure, including plasma vitellogenin and testicular oocytes in male fishes. Smallmouth bass Micropterus dolomieu, white sucker Catostomus commersonii, and redhorse sucker Moxostoma species were collected in the summer, a period of low flow and low reproductive activity. Smallmouth bass were the only species in which testicular oocytes were observed; however, measurable concentrations of plasma vitellogenin were found in male bass and white sucker. The percentage of male bass with testicular oocytes ranged from 10 to 100 %, with the highest prevalence and severity in bass collected in the Susquehanna drainage. The percentage of males with plasma vitellogenin ranged from 0 to 100 % in both bass and sucker. Biological findings were compared with chemical analyses of discrete water samples collected at the time of fish collections. Estrone concentrations correlated with testicular oocytes prevalence and severity and with the percentage of male bass with vitellogenin. No correlations were noted with the percentage of male sucker with vitellogenin and water chemical concentrations. The prevalence and severity of testicular oocytes in bass also correlated with the percent of agricultural land use in the watershed above a site. Two sites within the Susquehanna drainage and one in the Delaware were immediately downstream of wastewater treatment plants to compare results with upstream fish. The percentage of male bass with testicular oocytes was not consistently higher downstream; however, severity did tend to increase downstream.

  12. Effects of ultraviolet-B radiation on fish: Histologic comparison of a UVB-sensitive and a UVB-tolerant species

    USGS Publications Warehouse

    Blazer, V.S.; Fabacher, D.L.; Little, E.E.; Ewing, M.S.; Kocan, K.M.

    1997-01-01

    Lahontan cutthroat trout Oncorhynchus clarki henshawi were sensitive to simulated solar ultraviolet-B radiation (UVB) and exhibited grossly visible signs of sunburn upon exposure. Razorback suckers Xyrauchen texanus, however, were tolerant to simulated solar UVB and showed no grossly visible signs of exposure. Cutthroat trout also had considerably less of an unidentified, possibly photoprotective, substance in the skin than did razorback suckers. In all attempt to characterize the cellular response to simulated solar UVB exposure in the skin of these two species, we examined sections from UVB-exposed fish by light and electron microscopy. Cutthroat trout showed grossly visible signs of sunburn by 48 h. Histologic observations included a sloughing of the mucous cells, necrosis and edema in the epidermis and dermis, and, in some cases, secondary fungal infections. Razorback suckers did not show any visible signs of sunburn during 72 h of experimental exposure. Histologic analyses revealed that cell necrosis had occurred, but the severe necrosis and sloughing noted in cutthroat trout was not observed. An increase in epidermal thickness, apparently due to hypertrophy and hyperplasia of large PAS-negative cells, occurred in the razorback suckers. These cells contained a large central region of low electron density and appeared to be club cells. In some, extensive interdigitation of the electron-lucent cytoplasm with adjacent epithelial cell margins occurred. Near the surface of the epidermis these cells were larger and the interface with epithelial cells lacked complex interdigitation. These cells may contain the substance that appears to protect razorback suckers against UV-B radiation.

  13. Health of white sucker within the St. Louis River area of concern associated with habitat usage as assessed using stable isotopes

    USGS Publications Warehouse

    Blazer, V.S.; Hoffman, J.; Walsh, H.L.; Braham, R.P.; Hahn, C.; Collins, P.; Jorgenson, Z.; Ledder, T.

    2014-01-01

    In Spring 2011, 200 adult white sucker were collected in four areas of the St. Louis River area of concern (AOC), located in Minnesota and Wisconsin, USA. The areas included the upper AOC as a reference area, the upper estuary, St. Louis Bay and Superior Bay. Grossly visible abnormalities were documented and preserved for microscopic analyses, as were five to eight representative pieces of liver tissue. A piece of dorsal muscle was preserved for stable isotope analyses and otoliths removed for age determination. The incidence of raised skin lesions (mucoid plaques) was high (31 %), however, microscopically only 4.5 % of the white suckers had neoplasia (papillomas). The remaining lesions were epidermal hyperplasia. Superior Bay had the lowest percentage of skin/lip lesions (10 %), while St. Louis Bay had the highest (44 %). St. Louis Bay also had the highest incidence of skin neoplasms (12 %). No hepatocellular neoplasms were documented, however bile duct tumors were observed in 4.5 % of the suckers. Foci of cellular alteration were observed in fish from all sites except the upper AOC. Stable isotope data indicated that most of the suckers relied on the St. Louis River AOC for the majority (>75 %) of their diet, indicating they were resident within the AOC and not in Lake Superior. The amount of diet obtained from the upper estuary was a significant predictor of skin lesion incidence. Hence, habitat use within the AOC appears to be an important risk factor for skin and possibly, liver lesions.

  14. Proteomic profiles of white sucker (Catostomus commersonii) sampled from within the Thunder Bay Area of Concern reveal up-regulation of proteins associated with tumor formation and exposure to environmental estrogens.

    PubMed

    Simmons, Denina B D; Bols, Niels C; Duncker, Bernard P; McMaster, Mark; Miller, Jason; Sherry, James P

    2012-02-07

    White sucker (Catostomus commersonii) sampled from the Thunder Bay Area of Concern were assessed for health using a shotgun approach to compile proteomic profiles. Plasma proteins were sampled from male and female fish from a reference location, an area in recovery within Thunder Bay Harbour, and a site at the mouth of the Kaministiquia River where water and sediment quality has been degraded by industrial activities. The proteins were characterized using reverse-phase liquid chromatography tandem to a quadrupole-time-of-flight (LC-Q-TOF) mass spectrometer and were identified by searching in peptide databases. In total, 1086 unique proteins were identified. The identified proteins were then examined by means of a bioinformatics pathway analysis to gain insight into the biological functions and disease pathways that were represented and to assess whether there were any significant changes in protein expression due to sampling location. Female white sucker exhibited significant (p = 0.00183) site-specific changes in the number of plasma proteins that were related to tumor formation, reproductive system disease, and neurological disease. Male fish plasma had a significantly different (p < 0.0001) number of proteins related to neurological disease and tumor formation. Plasma concentrations of vitellogenin were significantly elevated in females from the Kaministiquia River compared to the Thunder Bay Harbour and reference sites. The protein expression profiles indicate that white sucker health has benefited from the remediation of the Thunder Bay Harbour site, whereas white sucker from the Kaministiquia River site are impacted by ongoing contaminant discharges.

  15. Health of white sucker within the St. Louis River area of concern associated with habitat usage as assessed using stable isotopes.

    PubMed

    Blazer, V S; Hoffman, J; Walsh, H L; Braham, R P; Hahn, C; Collins, P; Jorgenson, Z; Ledder, T

    2014-03-01

    In Spring 2011, 200 adult white sucker were collected in four areas of the St. Louis River area of concern (AOC), located in Minnesota and Wisconsin, USA. The areas included the upper AOC as a reference area, the upper estuary, St. Louis Bay and Superior Bay. Grossly visible abnormalities were documented and preserved for microscopic analyses, as were five to eight representative pieces of liver tissue. A piece of dorsal muscle was preserved for stable isotope analyses and otoliths removed for age determination. The incidence of raised skin lesions (mucoid plaques) was high (31 %), however, microscopically only 4.5 % of the white suckers had neoplasia (papillomas). The remaining lesions were epidermal hyperplasia. Superior Bay had the lowest percentage of skin/lip lesions (10 %), while St. Louis Bay had the highest (44 %). St. Louis Bay also had the highest incidence of skin neoplasms (12 %). No hepatocellular neoplasms were documented, however bile duct tumors were observed in 4.5 % of the suckers. Foci of cellular alteration were observed in fish from all sites except the upper AOC. Stable isotope data indicated that most of the suckers relied on the St. Louis River AOC for the majority (>75 %) of their diet, indicating they were resident within the AOC and not in Lake Superior. The amount of diet obtained from the upper estuary was a significant predictor of skin lesion incidence. Hence, habitat use within the AOC appears to be an important risk factor for skin and possibly, liver lesions.

  16. Health of white sucker within the St. Louis River area of concern associated with habitat usage as assessed using stable isotopes

    EPA Science Inventory

    In Spring 2011, 200 adult white sucker were collected in four areas of the St. Louis River Area of Concern (AOC), located in Minnesota and Wisconsin, USA. The areas included the upper AOC as a reference area, the upper estuary, St. Louis Bay and Superior Bay. Grossly visible abno...

  17. Correlation of gene expression and contaminat concentrations in wild largescale suckers: a field-based study

    USGS Publications Warehouse

    Christiansen, Helena E.; Mehinto, Alvine C.; Yu, Fahong; Perry, Russell W.; Denslow, Nancy D.; Maule, Alec G.; Mesa, Matthew G.

    2014-01-01

    Toxic compounds such as organochlorine pesticides (OCs), polychlorinated biphenyls (PCBs), and polybrominated diphenyl ether flame retardants (PBDEs) have been detected in fish, birds, and aquatic mammals that live in the Columbia River or use food resources from within the river. We developed a custom microarray for largescale suckers (Catostomus macrocheilus) and used it to investigate the molecular effects of contaminant exposure on wild fish in the Columbia River. Using Significance Analysis of Microarrays (SAM) we identified 72 probes representing 69 unique genes with expression patterns that correlated with hepatic tissue levels of OCs, PCBs, or PBDEs. These genes were involved in many biological processes previously shown to respond to contaminant exposure, including drug and lipid metabolism, apoptosis, cellular transport, oxidative stress, and cellular chaperone function. The relation between gene expression and contaminant concentration suggests that these genes may respond to environmental contaminant exposure and are promising candidates for further field and laboratory studies to develop biomarkers for monitoring exposure of wild fish to contaminant mixtures found in the Columbia River Basin. The array developed in this study could also be a useful tool for studies involving endangered sucker species and other sucker species used in contaminant research.

  18. A revision of the haploporinae nicoll, 1914 (digenea: haploporidae) from mullets (mugilidae): two new haploporine genera and a key to the genera of the subfamily.

    PubMed

    Blasco-Costa, Isabel; Montero, Francisco E; Gibson, David I; Balbuena, Juan Antonio; Kostadinova, Aneta

    2009-03-01

    Two new haploporine genera are established for parasites of mullets. Ragaia n. g. is erected for R. lizae n. sp. from Liza ramado in the Ebro Delta on the Mediterranean Coast of Spain. This new genus is distinguished by the unique combination of the following characters: a strongly muscular ventral sucker which is twice as large as the oral sucker; a large, muscular hermaphroditic sac similar in length to the ventral sucker; a saccular, thick-walled internal seminal vesicle which is larger than the external seminal vesicle; and the ovary and vitellarium located rather close to the posterior extremity. Pseudodicrogaster n. g. is erected to accommodate Dicrogaster japonica Machida, 1996, as P. japonica (Machida, 1996) n. comb., a parasite of Mugil cephalus L. off Fukaura, Japan. This genus is recognised on the basis of: the tubular condition of both the internal and external seminal vesicles, the latter being much shorter than the former; the sucker ratio; the massive pyriform hermaphroditic sac; the location of the testis; and the presence of two eye-spots in developed miracidia. A key to the nine recognised genera of the Haploporinae is presented.

  19. White sucker Catostomus commersonii respond to conspecific and sea lamprey Petromyzon marinus alarm cues but not potential predator cues

    USGS Publications Warehouse

    Jordbro, Ethan J.; Di Rocco, Richard T.; Imre, Istvan; Johnson, Nicholas; Brown, Grant E.

    2016-01-01

    Recent studies proposed the use of chemosensory alarm cues to control the distribution of invasive sea lamprey Petromyzon marinus populations in the Laurentian Great Lakes and necessitate the evaluation of sea lamprey chemosensory alarm cues on valuable sympatric species such as white sucker. In two laboratory experiments, 10 replicate groups (10 animals each) of migratory white suckers were exposed to deionized water (control), conspecific whole-body extract, heterospecific whole-body extract (sea lamprey) and two potential predator cues (2-phenylethylamine HCl (PEA HCl) and human saliva) during the day, and exposed to the first four of the above cues at night. White suckers avoided the conspecific and the sea lamprey whole-body extract both during the day and at night to the same extent. Human saliva did not induce avoidance during the day. PEA HCl did not induce avoidance at a higher concentration during the day, or at night at the minimum concentration that was previously shown to induce maximum avoidance by sea lamprey under laboratory conditions. Our findings suggest that human saliva and PEA HCl may be potential species-specific predator cues for sea lamprey.

  20. Standard weight (Ws) equations for four rare desert fishes

    USGS Publications Warehouse

    Didenko, A.V.; Bonar, Scott A.; Matter, W.J.

    2004-01-01

    Standard weight (Ws) equations have been used extensively to examine body condition in sport fishes. However, development of these equations for nongame fishes has only recently been emphasized. We used the regression-line-percentile technique to develop standard weight equations for four rare desert fishes: flannelmouth sucker Catostomus latipinnis, razorback sucker Xyrauchen texanus, roundtail chub Gila robusta, and humpback chub G. cypha. The Ws equation for flannelmouth suckers of 100-690 mm total length (TL) was developed from 17 populations: log10Ws = -5.180 + 3.068 log10TL. The Ws equation for razorback suckers of 110-885 mm TL was developed from 12 populations: log 10Ws = -4.886 + 2.985 log10TL. The W s equation for roundtail chub of 100-525 mm TL was developed from 20 populations: log10Ws = -5.065 + 3.015 log10TL. The Ws equation for humpback chub of 120-495 mm TL was developed from 9 populations: log10Ws = -5.278 + 3.096 log 10TL. These equations meet criteria for acceptable standard weight indexes and can be used to calculate relative weight, an index of body condition.

  1. Distributions of PCB congeners and homologues in white sucker and coho salmon from Lake Michigan

    USGS Publications Warehouse

    Stapanian, Martin A.; Madenjian, Charles P.; Batterman, Stuart A.; Chernyak, Sergei M.; Edwards, William H.; McIntyre, Peter B.

    2018-01-01

    We tested the hypothesis of the proportion of higher chlorinated biphenyl (PCB) congeners increasing with increasing trophic level by comparing the respective PCB homologue distributions in an omnivore, white sucker (Catostomus commersoni), and a top predator, coho salmon (Oncorhynchus kisutch), from Lake Michigan. Adult females had the same congener and homologue proportions of total PCB concentration (ΣPCB) as adult males in both species. Hexachlorinated congeners comprised the largest proportion (32%) found in white sucker, followed by heptachlorinated (21%) and octochlorinated (18%) congeners. In contrast, pentachlorinated congeners comprised the largest proportion (33%) of ΣPCB found in coho salmon, followed by hexachlorinated (26%) and tetrachlorinated (24%) congeners. Coho salmon contained significantly higher proportions of tri-, tetra-, and pentachlorinated congeners, whereas white sucker contained significantly higher proportions of hexa- through decachlorinated congeners. Our results were opposite of the hypothesis of greater degree of PCB chlorination with increasing trophic level, and supported the contention that the PCB congener proportions in fish depends mainly on diet, and does not necessarily reflect the trophic level of the fish. Our results also supported the contention that diets do not vary between the sexes in most fish populations.

  2. Fish abundance and population stability in a reservoir tailwater and an unregulated headwater stream

    USGS Publications Warehouse

    Jacobs, K.E.; Swink, W.D.

    1983-01-01

    Fish abundance and population stability were compared in the tailwater and in an unregulated tributary of Barren River Lake, a flood control reservoir in south central Kentucky. Fish abundance was greater in the tailwater near the dam and was dominated by three species common in the reservoir: gizzard shad (Dorosoma cepedianum), bluegills (Lepomis macrochirus), and white crappies (Pomoxis annularis). Three riverine suckers were less abundant in the tailwater than in the unregulated stream: northern hog suckers (Hypentelium nigricans), black redhorse (Moxostoma duquesnei), and golden redhorse (Moxostoma erythrurum). The fish populations in the tailwater, particularly common carp (Cyprinus carpio), northern hog suckers, black redhorse, and golden redhorse, were less stable than those in the unregulated stream. Population stability is defined as the extent to which fish remain in a stream section. This study suggests that the occurrence of reservoir species in the tailwater was the result of fish passage from the reservoir during high discharges in fall and winter. Reservoir operations (altered flow, low summer water temperature, and poor summer water quality) probably were responsible for the unstable populations of common carp and riverine suckers in the tailwater.

  3. Concentrations of cadmium, Cobalt, Lead, Nickel, and Zinc in Blood and Fillets of Northern Hog Sucker (Hypentelium nigricans) from streams contaminated by lead-Zinc mining: Implications for monitoring

    USGS Publications Warehouse

    Schmitt, C.J.; Brumbaugh, W.G.; May, T.W.

    2009-01-01

    Lead (Pb) and other metals can accumulate in northern hog sucker (Hypentelium nigricans) and other suckers (Catostomidae), which are harvested in large numbers from Ozark streams by recreational fishers. Suckers are also important in the diets of piscivorous wildlife and fishes. Suckers from streams contaminated by historic Pb-zinc (Zn) mining in southeastern Missouri are presently identified in a consumption advisory because of Pb concentrations. We evaluated blood sampling as a potentially nonlethal alternative to fillet sampling for Pb and other metals in northern hog sucker. Scaled, skin-on, bone-in "fillet" and blood samples were obtained from northern hog suckers (n = 75) collected at nine sites representing a wide range of conditions relative to Pb-Zn mining in southeastern Missouri. All samples were analyzed for cadmium (Cd), cobalt (Co), Pb, nickel (Ni), and Zn. Fillets were also analyzed for calcium as an indicator of the amount of bone, skin, and mucus included in the samples. Pb, Cd, Co, and Ni concentrations were typically higher in blood than in fillets, but Zn concentrations were similar in both sample types. Concentrations of all metals except Zn were typically higher at sites located downstream from active and historic Pb-Zn mines and related facilities than at nonmining sites. Blood concentrations of Pb, Cd, and Co were highly correlated with corresponding fillet concentrations; log-log linear regressions between concentrations in the two sample types explained 94% of the variation for Pb, 73-83% of the variation for Co, and 61% of the variation for Cd. In contrast, relations for Ni and Zn explained <12% of the total variation. Fillet Pb and calcium concentrations were correlated (r = 0.83), but only in the 12 fish from the most contaminated site; concentrations were not significantly correlated across all sites. Conversely, fillet Cd and calcium were correlated across the range of sites (r = 0.78), and the inclusion of calcium in the fillet-to-blood relation explained an additional 12% of the total variation in fillet Cd. Collectively, the results indicate that blood sampling could provide reasonably accurate and precise estimates of fillet Pb, Co, and Cd concentrations that would be suitable for identifying contaminated sites and for monitoring, but some fillet sampling might be necessary at contaminated sites for establishing consumption advisories. ?? 2009 US Government.

  4. Rod gripper, changer, and storage system

    NASA Technical Reports Server (NTRS)

    Benson, Mark; Demi, Todd; Mcneill, Robert; Waldo, Keith; Afghan, Alex; Oliver, Jim

    1989-01-01

    A rod changer and storage design is presented for the lunar deep drill apparatus to be used in conjunction with the Skitter walking platform. The design must take into account all of the lunar environment and working conditions. Some of these are: (1) the moon has one sixth the gravity of earth; (2) temperature gradients can range from about -170 to 265 C; (3) because of the high transportation costs, the design must be as light as possible; and (4) the process must be remotely operated (from earth or satellite) and must be automated. Because of Skitter's multiple degree of freedom movement, the design will utilize Skitter's movement to locate an implement and transport it from the rack to the drill string. The implement will be gripped by a thumb and two finger device, identified through an electronic sensing device on the thumb, and transported from the rack to the footplate and back from the footplate to the rack. The major designs discussed in this report have been broken down into three major areas: (1) gripper design (linear transport mechanism); (2) indexing system; and (3) rack design.

  5. Controlling Hazel, Aspen Suckers, and Mountain Maple with Picloram

    Treesearch

    Donald A. Perala

    1971-01-01

    Tests showed that picloram/2,4-D mixture was equal to or superior to 2,4-D alone or a 2,4,5-D/2,4,5-T mixture in controlling hazel, aspen suckers, and mountain maple for reforestation purposes. Survival of red pine planted 9 months after treatment was not influenced by residual soil effects of picloram. However, foliar application contributed to mortality of...

  6. Passive restoration augments active restoration in deforested landscapes: the role of root suckering adjacent to planted stands of Acacia koa

    Treesearch

    Paul G. Scowcroft; Justin T. Yeh

    2013-01-01

    Active forest restoration in Hawaii’s Hakalau Forest National Wildlife Refuge has produced a network of Acacia koa tree corridors and islands in deforested grasslands. Passive restoration by root suckering has potential to expand tree cover and close gaps between planted stands. This study documents rates of encroachment into grassland, clonal...

  7. Maintaining population persistence in the face of an extremely altered hydrograph: implications for three sensitive fishes in a tributary of the Green River, Utah

    USGS Publications Warehouse

    Bottcher, Jared L.

    2009-01-01

    The ability of an organism to disperse to suitable habitats, especially in modified and fragmented systems, determines individual fitness and overall population viability. The bluehead sucker (Catostomus discobolus), flannelmouth sucker (Catostomus latipinnis), and roundtail chub (Gila robusta) are three species native to the upper Colorado River Basin that now occupy only 50% of their historic range. Despite these distributional declines, populations of all three species are present in the San Rafael River, a highly regulated tributary of the Green River, Utah, providing an opportunity for research. Our goal was to determine the timing and extent of movement, habitat preferences, and limiting factors, ultimately to guide effective management and recovery of these three species. In 2007-2008, we sampled fish from 25 systematically selected, 300-m reaches in the lower 64 km of the San Rafael River, spaced to capture the range of species, life-stages, and habitat conditions present. We implanted all target species with a passive integrated transponder (PIT) tag, installed a passive PIT tag antennae, and measured key habitat parameters throughout each reach and at the site of native fish capture. We used random forest modeling to identify and rank the most important abiotic and biotic predictor variables, and reveal potential limiting factors in the San Rafael River. While flannelmouth sucker were relatively evenly distributed within our study area, highest densities of roundtail chub and bluehead sucker occurred in isolated, upstream reaches characterized by complex habitat. In addition, our movement and length-frequency data indicate downstream drift of age-0 roundtail chub, and active upstream movement of adult flannelmouth sucker, both from source populations, providing the lower San Rafael River with colonists. Our random forest analysis highlights the importance of pools, riffles, and distance-to-source populations, suggesting that bluehead sucker and roundtail chub are habitat limited in the lower San Rafael River. These results suggest management efforts should focus on diversifying habitat, maintaining in-stream flow, and removing barriers to movement.

  8. Habitat use and population characteristics of potentially spawning shovelnose sturgeon Scaphirhynchus platorynchus (Rafinesque, 1820), blue sucker ( Cycleptus elongatus (Lesueur, 1817), and associated species in the lower Wisconsin River, USA

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Lyons, John D.; Walchak, D.; Haglund, J.

    The goal of this study was to compare the possible locations, timing, and characteristics of potentially spawning shovelnose sturgeon ( Scaphirhynchus platorynchus), blue sucker ( Cycleptus elongatus), and associated species during the spring of 2007-2015 in the 149-km-long lower Wisconsin River, Wisconsin, USA, a large, shallow, sand-dominated Mississippi River tributary. A 5-km index station of two pairs of rocky shoals surrounded by sandy areas was electrofished for shovelnose sturgeon and blue sucker in a standardized fashion a total of 40 times from late March through mid- June, the presumed spawning period. On one date in 2008 and two dates inmore » 2012, all rocky shoals and adjacent sandy areas in the lowermost 149 km of the river were also electrofished for both species. Shovelnose sturgeon and blue sucker appeared to spawn in the limited rocky areas of the river along with at least four other species: mooneye ( Hiodon tergisus), quillback ( Carpiodes cyprinus), smallmouth buffalo ( Ictiobus bubalus), and shorthead redhorse ( Moxostoma macrolepidotum), usually at depths of 0.8-2.0 m and surface velocities of 0.4–1.0 m/s. However, apparently spawning shovelnose sturgeon were found only on mid-channel cobble and coarse gravel shoals within a single 7-km segment that included the 5-km index station, whereas apparently spawning blue suckers were encountered on these same shoals but also more widely throughout the river on eroding bluff shorelines of bedrock and boulder and on artificial boulder wing dams and shoreline rip-rap. Both species showed evidence of homing to the same mid-channel shoal complexes across years.« less

  9. Habitat use and population characteristics of potentially spawning shovelnose sturgeon Scaphirhynchus platorynchus (Rafinesque, 1820), blue sucker ( Cycleptus elongatus (Lesueur, 1817), and associated species in the lower Wisconsin River, USA

    DOE PAGES

    Lyons, John D.; Walchak, D.; Haglund, J.; ...

    2016-11-07

    The goal of this study was to compare the possible locations, timing, and characteristics of potentially spawning shovelnose sturgeon ( Scaphirhynchus platorynchus), blue sucker ( Cycleptus elongatus), and associated species during the spring of 2007-2015 in the 149-km-long lower Wisconsin River, Wisconsin, USA, a large, shallow, sand-dominated Mississippi River tributary. A 5-km index station of two pairs of rocky shoals surrounded by sandy areas was electrofished for shovelnose sturgeon and blue sucker in a standardized fashion a total of 40 times from late March through mid- June, the presumed spawning period. On one date in 2008 and two dates inmore » 2012, all rocky shoals and adjacent sandy areas in the lowermost 149 km of the river were also electrofished for both species. Shovelnose sturgeon and blue sucker appeared to spawn in the limited rocky areas of the river along with at least four other species: mooneye ( Hiodon tergisus), quillback ( Carpiodes cyprinus), smallmouth buffalo ( Ictiobus bubalus), and shorthead redhorse ( Moxostoma macrolepidotum), usually at depths of 0.8-2.0 m and surface velocities of 0.4–1.0 m/s. However, apparently spawning shovelnose sturgeon were found only on mid-channel cobble and coarse gravel shoals within a single 7-km segment that included the 5-km index station, whereas apparently spawning blue suckers were encountered on these same shoals but also more widely throughout the river on eroding bluff shorelines of bedrock and boulder and on artificial boulder wing dams and shoreline rip-rap. Both species showed evidence of homing to the same mid-channel shoal complexes across years.« less

  10. Comparison of the precision of age estimates generated from fin rays, scales, and otoliths of Blue Sucker

    USGS Publications Warehouse

    Acre, Matthew R.; Alejandrez, Celeste; East, Jessica; Massure, Wade A.; Miyazono, S.; Pease, Jessica E.; Roesler, Elizabeth L.; Williams, H.M.; Grabowski, Timothy B.

    2017-01-01

    Evaluating the precision of age estimates generated by different readers and different calcified structures is an important part of generating reliable estimations of growth, recruitment, and mortality for fish populations. Understanding the potential loss of precision associated with using structures harvested without sacrificing individuals, such as scales or fin rays, is particularly important when working with imperiled species, such as Cycleptus elongatus (Blue Sucker). We collected otoliths (lapilli), scales, and the first fin rays of the dorsal, anal, pelvic, and pectoral fins of 9 Blue Suckers. We generated age estimates from each structure by both experienced (n = 5) and novice (n = 4) readers. We found that, independent of the structure used to generate the age estimates, the mean coefficient of variation (CV) of experienced readers was approximately 29% lower than that of novice readers. Further, the mean CV of age estimates generated from pectoral-fin rays, pelvic-fin rays, and scales were statistically indistinguishable and less than those of dorsal-fin rays, anal-fin rays, and otoliths. Anal-, dorsal-, and pelvic-fin rays and scales underestimated age compared to otoliths, but age estimates from pectoral-fin rays were comparable to those from otoliths. Skill level, structure, and fish total-length influenced reader precision between subsequent reads of the same aging structure from a particular fish. Using structures that can be harvested non-lethally to estimate the age of Blue Sucker can provide reliable and reproducible results, similar to those that would be expected from using otoliths. Therefore, we recommend the use of pectoral-fin rays as a non-lethal method to obtain age estimates for Blue Suckers.

  11. Chemical and Ecological Health of White Sucker (Catostomus Commersoni) in Rock Creek Park, Washington, D.C., 2003-04

    USGS Publications Warehouse

    Miller, Cherie V.; Weyers, Holly S.; Blazer, Vicki; Freeman, Mary E.

    2006-01-01

    Several classes of chemicals that are known or suspected contaminants were found in bed sediment in Rock Creek, including polyaromatic hydrocarbons (PAHs), phthalate esters, organochlorine pesticides, dioxins and furans, trace metals and metalloids (mercury, arsenic, cadmium, chromium, cobalt, copper, lead, nickel, silver, and zinc), and polychlorinated biphenyls (total PCBs and selected aroclors). Concentrations of many of these chemicals consistently exceeded threshold or chronic-effects guidelines for the protection of aquatic life and often exceeded probable effects levels (PELs). Exceedance of PELs was dependent on the amount of total organic carbon in the sediments. Concurrent with the collection of sediment-quality data, white sucker (Catostomus commersoni) were evaluated for gross-external and internal-organ anomalies, whole-body burdens of chemical contaminants, and gut contents to determine prey. The histopathology of internal tissues of white sucker was compared to contaminant levels in fish tissue and bed sediment. Gut contents were examined to determine preferential prey and thus potential pathways for the bioaccumulation of chemicals from bed sediments. Male and female fish were tested separately. Lesions and other necroses were observed in all fish collected during both years of sample collection, indicating that fish in Rock Creek have experienced some form of environmental stress. No direct cause and effect was determined for chemical exposure and compromised fish health, but a substantial weight of evidence indicates that white sucker, which are bottom-feeding fish and low-order consumers in Rock Creek, are experiencing some reduction in vitality, possibly due to immunosuppression. Abnormalities observed in gonads of both sexes of white sucker and observations of abnormal behavior during spawning indicated some interruption in reproductive success.

  12. Chemical and ecological health of white sucker (Catostomus Commersoni) in Rock Creek Park, Washington, D.C., 2003?04

    USGS Publications Warehouse

    Miller, C.V.; Weyers, H.S.; Blazer, V.S.; Freeman, M.E.

    2006-01-01

    Several classes of chemicals that are known or suspected contaminants were found in bed sediment in Rock Creek, including polyaromatic hydrocarbons (PAHs), phthalate esters, organochlorine pesticides, dioxins and furans, trace metals and metalloids (mercury, arsenic, cadmium, chromium, cobalt, copper, lead, nickel, silver, and zinc), and polychlorinated biphenyls (total PCBs and selected aroclors). Concentrations of many of these chemicals consistently exceeded thresholdor chronic-effects guidelines for the protection of aquatic life and often exceeded probable effects levels (PELs). Exceedance of PELs was dependent on the amount of total organic carbon in the sediments. Concurrent with the collection of sediment-quality data, white sucker (Catostomus commersoni) were evaluated for gross-external and internal-organ anomalies, whole-body burdens of chemical contaminants, and gut contents to determine prey. The histopathology of internal tissues of white sucker was compared to contaminant levels in fish tissue and bed sediment. Gut contents were examined to determine preferential prey and thus potential pathways for the bioaccumulation of chemicals from bed sediments. Male and female fish were tested separately. Lesions and other necroses were observed in all fish collected during both years of sample collection, indicating that fish in Rock Creek have experienced some form of environmental stress. No direct cause and effect was determined for chemical exposure and compromised fish health, but a substantial weight of evidence indicates that white sucker, which are bottom-feeding fish and low-order consumers in Rock Creek, are experiencing some reduction in vitality, possibly due to immunosuppression. Abnormalities observed in gonads of both sexes of white sucker and observations of abnormal behavior during spawning indicated some interruption in reproductive success.

  13. Cibola High Levee Pond Annual Report 2003. Interim Report

    USGS Publications Warehouse

    Mueller, G.A.; Carpenter, J.; Marsh, P.C.; Minckley, C.O.

    2003-01-01

    Bonytail and razorback sucker have once again spawned and produced swim-up larvae in Cibola High Levee Pond (CHLP). CHLP continues to support annual recruitment of bonytail while recent razorback sucker recruitment remains elusive. Thus far, razorbacks have experienced intermittent years of spawning success. Both native species were observed spawning on, or near, the riprap on the river levee. Razorbacks spawned from late January until mid-March over gravel and large cobble along the levee toe (2-3 m depth) and bonytail spawned along the levee shoreline during mid-April. Razorback suckers rapidly fin during the reproductive act, which flushes fines from the substrate and leaves gravel relatively clean. Bonytail on the other hand, appear to spawn over or on substrate that has been disturbed by beaver activity. Substrate scour or disturbance appears to be an important factor in spawning site selectiona?|

  14. Behaviors of southwestern native fishes in response to introduced catfish predators

    USGS Publications Warehouse

    Ward, David L.; Figiel, Chester R.

    2013-01-01

    Native fishes reared in hatcheries typically suffer high predation mortality when stocked into natural environments. We evaluated the behavior of juvenile bonytail Gila elegans, roundtail chub Gila robusta, razorback sucker Xyrauchen texanus, and Sonora sucker Catostomus insignis in response to introduced channel catfish Ictalurus punctatus and flathead catfish Pylodictis olivaris. Our laboratory tests indicate these species did not inherently recognize catfish as a threat, but they can quickly (within 12 h) change their behavior in response to a novel predator paired with the sight and scent of a dead conspecific. Chubs appear to avoid predation by swimming away from the threat, whereas suckers reduced movement. Effects of antipredator conditioning on survival of fish reared in hatcheries is unknown; however, our results suggest some native fish can be conditioned to recognize introduced predators, which could increase poststocking survival.

  15. Basal area growth for aspen suckers under simulated browsing on Cedar Mountain, southern Utah, western United States of America

    Treesearch

    K. Tshireletso; J. C. Malechek; D. L. Bartos

    2010-01-01

    The objective of the study was to determine the effects of season and intensity of clipping using simulated browsing on suckers' (Populus tremulaides Michx.) basal area growth on Cedar Mountain, Southern Utah, Western United States of America. Three randomly selected stands measuring 70 m x 70 m were clear-felled in mid-July, 2005, and fenced. Simulated browsing...

  16. A new species of Malmiana (Oligochaeta: Hirudinida: Piscicolidae) from tidepool fishes in northern California.

    PubMed

    Burreson, Eugene M; Kalman, Julianne E

    2006-02-01

    Malmiana buthi n. sp. is described from the body and fins of the fluffy sculpin, Oligocottus snyderi, the tidepool sculpin, Oligocottus maculosus, and the woolly sculpin, Clinocottus analis, collected from tidepools at Horseshoe Cove on the Bodega Marine Reserve in Sonoma County, California. Prevalence of the leech was 32.6% on live-caught O. snyderi; mean intensity on O. snyderi was 3.3 leeches per fish, with a range from 1 to 7. The leech is not known to exceed 8 mm total length. The body is smooth, lacking papillae, tubercles, or pulsatile vesicles. Two pairs of crescentiform eyes are present on the oral sucker, and 1 pair of punctiform eyes occurs on the second annulus of the trachelosome. The caudal sucker has 14 small punctiform ocelli spaced evenly around the margin. The last 9 segments of the urosome have pairs of large punctiform ocelli both dorsally and ventrally. Body and caudal sucker pigmentation is uniformly reddish brown dorsally and ventrally with segental, lateral, unpigmented areas on both the urosome and trachelosome; pigmentation on the oral sucker is in the form of a cross. There are 5 pairs of testisacs; accessory gland cells on the atrial cornua and vector tissue are absent.

  17. A new cercaria and metacercaria of Acanthoparyphium (Echinostomatidae) found in an intertidal snail Zeacumantus subcarinatus (Batillaridae) from New Zealand.

    PubMed

    Martorelli, Sergio R; Poulin, Robert; Mouritsen, Kim N

    2006-09-01

    A new 23-collar-spined cercaria and metacercaria are described from intertidal molluscs of the coast of New Zealand. The new cercaria found emerging from the mud snails Zeacumantus subcarinatus (Sowerby) (Prosobranchia: Batillariidae) is characterized mainly by the number and arrangement of the cephalic glands, the size of the suckers, and the size and number of the collar spines. The cercaria encysts in the cockle Austrovenus stutchburyi (Wood) (Bivalvia: Veneridae) which lives in sympatry with the first intermediate host throughout New Zealand. Laboratory infections of cockles by cercariae from naturally infected snails resulted in metacercariae identical to those found in naturally infected cockles. The main features of the cercaria and metacercaria are the presence of a reniform collar with 23 spines, two pairs of small cephalic glands at the oral sucker level, another two pairs of much longer ones posterior to the pharynx, and the excretory vesicle Y-shaped with the main collecting canals extending to the anterior level of the ventral sucker. Each of the main collecting canals had 10-11 pairs of bilateral diverticula between the anterior edge of the ventral sucker and anterior body end. A brief discussion of its possible life cycle and ecology is also provided.

  18. Petasiger islandicus n. sp. (Digenea: Echinostomatidae) in the horned grebe Podiceps auritus (L.) (Aves: Podicipedidae) from Iceland.

    PubMed

    Kostadinova, Aneta; Skirnisson, Karl

    2007-11-01

    Petasiger islandicus n. sp. is described and figured from a demographically isolated population of the horned grebe Podiceps auritus auritus (L.) in Lake Mývatn (Iceland). This new species belongs to the group of species with 19 collar spines which possess a large elongate-oval cirrus-sac, well-developed pars prostatica and massive bulb-like cirrus. Within this group, P. islandicus appears most similar to P. oschmarini Kostadinova & Gibson, 1998, a form with similar body dimensions described from the same host, but differs in having a larger head collar, collar spines, oral sucker, pharynx, testes and sucker-width ratio, and a smaller cirrus-sac, cirrus and eggs. Two Nearctic species resemble P. islandicus in general morphology but differ as follows: P. pseudoneocomense Bravo-Hollis, 1969 has a larger body and collar width, notably shorter collar spines, smaller testes and sucker-width ratio, and a shorter but much wider cirrus-sac which is also smaller relative to the ventral sucker and almost entirely anterior to it; and P. caribbensis Nassi, 1980 has a smaller body, shorter collar spines and a seminal vesicle which is small in relation to the cirrus-sac, vitelline fields reaching anteriorly to the level of the genital pore and the intestinal bifurcation is located more anteriorly.

  19. A method for sampling microbial aerosols using high altitude balloons.

    PubMed

    Bryan, N C; Stewart, M; Granger, D; Guzik, T G; Christner, B C

    2014-12-01

    Owing to the challenges posed to microbial aerosol sampling at high altitudes, very little is known about the abundance, diversity, and extent of microbial taxa in the Earth-atmosphere system. To directly address this knowledge gap, we designed, constructed, and tested a system that passively samples aerosols during ascent through the atmosphere while tethered to a helium-filled latex sounding balloon. The sampling payload is ~ 2.7 kg and comprised of an electronics box and three sampling chambers (one serving as a procedural control). Each chamber is sealed with retractable doors that can be commanded to open and close at designated altitudes. The payload is deployed together with radio beacons that transmit GPS coordinates (latitude, longitude and altitude) in real time for tracking and recovery. A cut mechanism separates the payload string from the balloon at any desired altitude, returning all equipment safely to the ground on a parachute. When the chambers are opened, aerosol sampling is performed using the Rotorod® collection method (40 rods per chamber), with each rod passing through 0.035 m3 per km of altitude sampled. Based on quality control measurements, the collection of ~ 100 cells rod(-1) provided a 3-sigma confidence level of detection. The payload system described can be mated with any type of balloon platform and provides a tool for characterizing the vertical distribution of microorganisms in the troposphere and stratosphere. Copyright © 2014 Elsevier B.V. All rights reserved.

  20. Assessment of the St. Louis River AOC fish tumors and other ...

    EPA Pesticide Factsheets

    The Fish Tumors and Other Deformities Beneficial Use Impairment (BUI) was listed as one of nine BUIs at the time the St. Louis River AOC was designated in 1987. At the time, no formal studies had been conducted to estimate the prevalence of either fish tumors or deformities. To assess the current status of fish tumors and deformities in the AOC, adult white suckers (Catostomus commersonii) were sampled during the spawning period in May during 2011, 2013 and 2015. We measured each fish, determined its sex and age (sagittal otolith), sampled dorsal muscle tissue for carbon and nitrogen stable isotope ratios to determine river versus lake residency, and conducted a necropsy-based assessment to document grossly visible abnormalities. Pieces of any observable abnormalities were preserved for subsequent histological analyses. A total of 622 white sucker were surveyed from 2011 through 2015. Of these, 27 (4.3%) white suckers had skin neoplasms, all of which were papillomas. A total of 29 (4.7%) of the white suckers had liver neoplasms. Sex and age were significantly associated with skin neoplasia incidence, whereas only age was significantly associated with liver neoplasia incidence. Neither neoplasia type was significantly associated with habitat use, indicating that neoplasia incidence did not change with increasing feeding in the AOC relative to Lake Superior. Further, there was not a significant difference in skin or liver neoplasia incidence between migratory whit

  1. Retention of Ancestral Genetic Variation Across Life-Stages of an Endangered, Long-Lived Iteroparous Fish.

    PubMed

    Carson, Evan W; Turner, Thomas F; Saltzgiver, Melody J; Adams, Deborah; Kesner, Brian R; Marsh, Paul C; Pilger, Tyler J; Dowling, Thomas E

    2016-11-01

    As with many endangered, long-lived iteroparous fishes, survival of razorback sucker depends on a management strategy that circumvents recruitment failure that results from predation by non-native fishes. In Lake Mohave, AZ-NV, management of razorback sucker centers on capture of larvae spawned in the lake, rearing them in off-channel habitats, and subsequent release ("repatriation") to the lake when adults are sufficiently large to resist predation. The effects of this strategy on genetic diversity, however, remained uncertain. After correction for differences in sample size among groups, metrics of mitochondrial DNA (mtDNA; number of haplotypes, N H , and haplotype diversity, H D ) and microsatellite (number of alleles, N A , and expected heterozygosity, H E ) diversity did not differ significantly between annual samples of repatriated adults and larval year-classes or among pooled samples of repatriated adults, larvae, and wild fish. These findings indicate that the current management program thus far maintained historical genetic variation of razorback sucker in the lake. Because effective population size, N e , is closely tied to the small census population size (N c = ~1500-3000) of razorback sucker in Lake Mohave, this population will remain at risk from genetic, as well as demographic risk of extinction unless N c is increased substantially. © The American Genetic Association 2016. All rights reserved. For permissions, please e-mail: journals.permissions@oup.com.

  2. [Function of the acetabulum of digenetic trematodes, as exemplified by Dicrocoelium dendriticum].

    PubMed

    Neuhaus, W

    1985-01-01

    The suckers of animals adhere to the substratum either in air or in aqueous fluids. The varying compressibility of these media causes differences in function, the principles of which are described. The ventral arch of the acetabulum of Dicrocoelium dendriticum, like the remaining body, is limited by the integument, basal lamina and skin muscles. The dorsal arch covers a basal lamina, which is close to a plexus of diagonal, longitudinal and circular muscles. The radical muscles, attached at the ventral basal lamina by thin connective tissue, continue in relatively thick contractile fibers, which split up into several fibrils, which also attached by thin connective tissue at the dorsal basal lamina. In this way the tension of the muscles is likewise distributed over the dorsal and ventral arches of the acetabulum. After contact with the substratum the sucker creates a partial vacuum and attachment by means of the pressure of the radial muscles against the wall of the hole. Because of the fluid content of the hole, the volume does not change much. The dorsal arch of the sucker withstands the pressure of the radical muscles, because its surface area is six times greater than that of the ventral arch and consequently the load is six times less. The sucker, covered with basal lamina, has a constant volume; its layer of muscles resists deformation and supports the stability of the arch.

  3. Factors influencing poststocking dispersal of razorback sucker

    USGS Publications Warehouse

    Mueller, G.A.; Marsh, P.C.; Foster, D.; Ulibarri, M.; Burke, T.

    2003-01-01

    Efforts to reintroduce razorback suckers Xyrauchen texanus to specific river reaches have been plagued by downstream drift and poor survival, which have been attributed to stress, disorientation, predation, and poor conditioning. Poststocking dispersal of eight test groups (15 fish each) of razorback suckers was examined for 28 d with telemetry equipment. Fish were released in three different locations in the Colorado River basin of Utah, Arizona, and Nevada: (1) a 65,000-ha reservoir, (2) a small (<1-ha) backwater, and (3) a large (30-ha) backwater on the mainstem river. At each location, subgroups were released immediately (reference) or held to acclimate them to the site (3–7 d) before release. Two of four subgroups for the large-backwater test were preconditioned to flow. Dispersal from the stocking sites was rapid and declined with time for all tests, as fish appeared to seek and find cover. Downstream drift was most pronounced (x = 69.5 km) from the small backwater and significantly (Kruskal–Wallis test, P < 0.01) greater than either the reservoir (x = 3.7 km) or large-backwater sites (x = 7.7 km). Site-acclimation tests were inconclusive, but downstream movement was significantly (Wilcoxon two-sample test, Z = −2.298, P < 0.01) less for fish preconditioned to flow (x = 1.9 km) compared with pond-reared fish (x = 7.7 km). We concluded that poststocking dispersal may decrease if razorback suckers are preconditioned to flow.

  4. Genetic variation of naturally growing olive trees in Israel: from abandoned groves to feral and wild?

    PubMed

    Barazani, Oz; Keren-Keiserman, Alexandra; Westberg, Erik; Hanin, Nir; Dag, Arnon; Ben-Ari, Giora; Fragman-Sapir, Ori; Tugendhaft, Yizhar; Kerem, Zohar; Kadereit, Joachim W

    2016-12-13

    Naturally growing populations of olive trees are found in the Mediterranean garrigue and maquis in Israel. Here, we used the Simple Sequence Repeat (SSR) genetic marker technique to investigate whether these represent wild var. sylvestris. Leaf samples were collected from a total of 205 trees at six sites of naturally growing olive populations in Israel. The genetic analysis included a multi-locus lineage (MLL) analysis, Rousset's genetic distances, Fst values, private alleles, other diversity values and a Structure analysis. The analyses also included scions and suckers of old cultivated olive trees, for which the dominance of one clone in scions (MLL1) and a second in suckers (MLL7) had been shown earlier. The majority of trees from a Judean Mts. population and from one population from the Galilee showed close genetic similarity to scions of old cultivated trees. Different from that, site-specific and a high number of single occurrence MLLs were found in four olive populations from the Galilee and Carmel which also were genetically more distant from old cultivated trees, had relatively high genetic diversity values and higher numbers of private alleles. Whereas in two of these populations MLL7 (and partly MLL1) were found in low frequency, the two other populations did not contain these MLLs and were very similar in their genetic structure to suckers of old cultivated olive trees that originated from sexual reproduction. The genetic distinctness from old cultivated olive trees, particularly of one population from Galilee and one from Carmel, suggests that trees at these sites might represent wild var. sylvestris. The similarity in genetic structure of these two populations with the suckers of old cultivated trees implies that wild trees were used as rootstocks. Alternatively, trees at these two sites may be remnants of old cultivated trees in which the scion-derived trunk died and was replaced by suckers. However, considering landscape and topographic environment at the two sites this second interpretation is less likely.

  5. Age, growth, and maturity of the longnose sucker Catostomus catostomus, of western Lake Superior

    USGS Publications Warehouse

    Bailey, Merryll M.

    1969-01-01

    Studies of age, growth, and maturity were based on 1760 fish collected in western Lake Superior in 1964-65. The body:scale relation was curvilinear and the curve had an intercept of 1.65 inches on the length axis. The weight increased as the 2.85 power of the length. Some fish formed an annulus before May 18 in 1965; all had completed annuli by late September. Longnose suckers grew 3.6 inches the 1st year, reached 12 inches in the 6th year, and 18 inches in the 11th year. Fish from Pikes Bay grew faster than those from Gull Island Shoal. Over 6 years were required for weight to reach 1 lb and nearly 10 years to reach 2 lb. Minimum length at maturity was 10.5 inches for males and 11.5 inches for females. The youngest mature male belonged to age-group IV and the youngest mature female to age-group V. All males were mature at 14.5-14.9 inches (age-group VIII) and all females at 15.0-15.4 inches (age-group IX). Finclipped longnose suckers returned to spawn in the Brule River in successive years. One fish returned to spawn in 4 successive years. Many of the fish were not recaptured until 2 or 3 years after marking. The time of the Brule River spawning migration depended more on water temperature than on length of day. The average water temperature during the peak of the spawning runs of 1958-64 was 55.4 F. Larval suckers apparently spend little time in the Brule River and adjacent streams and drift downstream to the lake soon after hatching. The number of eggs in the ovaries of eight suckers ranged from 14 to 35 thousand and averaged 24 thousand for fish 13.9-17.7 inches long.

  6. Chronic toxicity and hazard assessment of an inorganic mixture simulating irrigation drainwater to razorback sucker and bonytail

    USGS Publications Warehouse

    Hamilton, Steven J.; Buhl, Kevin J.; Bullard, Fern A.; Little, Edward E.

    2000-01-01

    We conducted two 90 day chronic toxicity studies with two endangered fish, razorback sucker and bonytail. Swim-up larvae were exposed in a reconstituted water simulating the middle Green River. The toxicant mixture simulated the environmental ratio and concentrations of inorganics reported in a Department of the Interior study for the mouth of Ashley Creek on the Green River, and was composed of nine elements. The mixture was tested at 1X, 2X, 4X, 8X, and 16X where X was the measured environmental concentration (2 μg/L arsenic, 630 μg/L boron, 10 μg/L copper, 5 μg/L molybdenum, 51 μg/L selenate, 8 μg/L selenite, 33 μg/L uranium, 2 μg/L vanadium, and 20 μg/L zinc). Razorback sucker had reduced survival after 60 days exposure to the inorganic mixture at 8X, whereas growth was reduced after 30 and 60 days at 2X and after 90 days at 4X. Bonytail had reduced survival after 30 days exposure at 16X, whereas growth was reduced after 30, 60, and 90 days at 8X. Swimming performance of razorback sucker and bonytail were reduced after 60 and 90 days of exposure at 8X. Whole-body residues of copper, selenium, and zinc increased in a concentration-response manner and seemed to be regulated at 90 days of exposure at 4X and lower treatments for razorback sucker, and at 8X and lower for bonytail. Adverse effects occurred in fish with whole-body residues of copper, selenium, and zinc similar to those causing similar effects in other fish species. Comparison of adverse effect concentrations with measured environmental concentrations showed a high hazard to the two endangered fish. Irrigation activities may be a contributing factor to the decline of these endangered fishes in the middle Green River. 

  7. Chronic toxicity of un-ionized ammonia to early life-stages of endangered Colorado pikeminnow (Ptychocheilus lucius) and razorback sucker (Xyrauchen texanus) compared to the surrogate fathead minnow (Pimephales promelas)

    USGS Publications Warehouse

    Fairchild, J.F.; Allert, A.L.; Sappington, L.C.; Waddell, B.

    2005-01-01

    Ammonia-contaminated groundwater enters the Upper Colorado River from beneath the abandoned Moab Uranium Mill Tailings Pile near Moab, Utah. This reach of the Upper Colorado River was designated as critical habitat for four endangered fish species because it is one of the few existing areas with known spawning and rearing habitats. Un-ionized ammonia (NH3) concentrations frequently exceed 1.00 mg/L in backwaters adjacent to the tailings pile, which exceeds the Utah 30-d average chronic water quality criterion for un-ionized ammonia (0.07 mg/L NH3; temperature 20??C; pH 8.2) by a factor of more than 10. However, there is little published information regarding the sensitivity of endangered fishes to ammonia. We conducted 28-d static renewal studies with post-swim-up larvae to determine the relative sensitivity of Colorado pikeminnow (Ptychocheilus lucius), razorback sucker (Xyrauchen texanus), and the standard surrogate fathead minnow (Pimephales promelas) to NH3. Chronic values (ChVs) for mortality and growth were determined as the geometric mean of the no observed effect concentration and the lowest observed effect concentration based on analysis of variance. The ChVs for growth of fathead minnow, Colorado pikeminnow, and razorback sucker were 0.43, 0.40, and 0.67 mg/L NH3, respectively. The ChVs for mortality of fathead minnow, Colorado pikeminnow, and razorback sucker were 0.43, 0.70, and 0.67 mg/L NH3, respectively. Therefore, the ChVs for mortality and growth were similar for fathead minnow and razorback sucker; however, the ChV for growth was lower than the ChV for mortality for Colorado pikeminnow. Maximum likelihood regression was used to calculate 28-d lethal concentrations (LCx) for each species. The 28-d LC50, LC20, and LC1 values for fathead minnow were 0.69, 0.42, and 0.13 mg/L NH3, respectively. The 28-d LC50, LC20, and LC1 values for Colorado pikeminnow were 0.76, 0.61, and 0.38 mg/L NH3, respectively. The 28-d LC50, LC20, and LC1 values for razorback sucker were 0.54, 0.38, and 0.25 mg/L NH3, respectively. The fathead minnow, Colorado pikeminnow, and razorback sucker are relatively similar in sensitivity and rank at the 35th, 49th, and 31st percentiles, respectively, of the theoretical chronic fish sensitivity distributions for NH3. The existing water quality criteria for NH3, if met by remediation activities at the Moab site, would be protective of these endangered fishes even if fish sensitivity is based on the conservative LC1 value. ?? 2005 Springer Science+Business Media, Inc.

  8. Sex difference in PCB concentrations of a catostomid fish

    USGS Publications Warehouse

    Madenjian, Charles P.; Stevens, Andrew L.; Stapanian, Martin A.; Batterman, Stuart A.; Chernyak, Sergei M.; Menczer, Jordan E.; McIntyre, Peter B.

    2017-01-01

    Unraveling the complexities associated with the relative differences in contaminant concentrations between the sexes of mature fish may provide insights into important behavioral and physiological differences between the sexes of not just fish but higher vertebrates as well. Whole-fish polychlorinated biphenyl (PCB) concentrations were determined in 25 mature female white suckers (Catostomus commersoni) and 26 mature male white suckers caught during their spawning run in the Kewaunee River, a tributary to Lake Michigan. Total length and weight were measured for each fish, and age of each fish was estimated from thin-sectioned otoliths. PCB concentration significantly increased with increasing total length, weight, and age. Consequently, three analysis of covariance (ANCOVA) models were fitted to the data to assess the effect of sex on white sucker PCB concentration. Based on model averaging, estimates of mean PCB concentrations in female and male white suckers were 185 and 219 ng/g, respectively. Thus, males were 18% greater in PCB concentration than females. We conclude that this difference between the sexes was most likely mainly driven by a higher rate of energy expenditure in males compared with females. Greater energy expenditure, owing to greater swimming activity and a higher resting metabolic rate, resulted in a higher rate of food consumption, which in turn led to a greater rate of PCB accumulation. Higher whole-fish PCB concentration in males compared with females has now been shown in nine different fish species. Our study represented the first documentation of this type of sex difference in a catostomid fish.

  9. Interaction between juniper Juniperus communis L. and its fruit pest insects: Pest abundance, fruit characteristics and seed viability

    NASA Astrophysics Data System (ADS)

    García, Daniel

    1998-12-01

    The relationships between the fruit features of Juniperus communis and the presence of fruit pests were studied in Sierra Nevada, SE Spain. The abundance of two insect species — a pulp-sucking scale and a seed-predator wasp — was surveyed with respect both to fruit characteristics and to viability of seeds contained therein. Seed-predator pressure was not significantly related to any fruit characteristics; however, pulp suckers tended to be more abundant in plants with low pulp: seed ratios and high fruit-water content. In addition, fruits with high levels of pulp-sucker attack tended to have higher water content. A multi-factor ANOVA, considering the identity of the plant and the attack of the different pests as factors, showed that plant identity accounts for most of the variation in fruit characteristics. The viability of seeds tended to be lower in plants strongly attacked by both pests. Fruits attacked by seed predators showed significantly lower proportions of viable and unviable seeds than did unattacked fruits. Seed viability was also lower in those fruits heavily attacked by pulp suckers, but this pattern is strongly mediated by plant identity. Pest activity proved to be clearly associated with a direct decrease in juniper reproductive capacity. This loss involved a reduction of the viable-seed number, mainly related to the seed predator, as well as a reduction of fruit attractiveness to frugivorous dispersers, related to the pulp sucker.

  10. Megalophallus deblocki n. sp. (Digenea: Microphallidae) from Rostrhamus sociabilis (Vieillot) (Aves: Accipitridae) in Paraguay.

    PubMed

    Kostadinova, Aneta; Vaucher, Claude; Gibson, David I

    2006-02-01

    A new species in the little-known digenean fauna of Paraguayan birds is described in detail, especially in relation to its terminal genitalia. The new form, Megalophallus deblocki n. sp., is a microphallid from the intestine of the snail kite Rostrhamus sociabilis (Vieillot). It is differentiated in detail from its congeners, but differs mainly in having a substantially larger phallus (e.g. in relation to the ventral sucker), a smaller ventral sucker and smaller eggs. Comments are included on the functional morphology of the terminal genitalia.

  11. Algal toxins in Upper Klamath Lake, Oregon: Linking water quality to juvenile sucker health

    USGS Publications Warehouse

    VanderKooi, S.P.; Burdick, S.M.; Echols, K.R.; Ottinger, C.A.; Rosen, B.H.; Wood, T.M.

    2010-01-01

    As the lead science agency for the Department of Interior, the U.S. Geological Survey is actively involved in resource issues in the Klamath River basin. Activities include research projects on endangered Lost River and shortnose suckers, threatened coho salmon, groundwater resources, seasonal runoff forecasting, water quality in Upper Klamath Lake and the Klamath River, nutrient cycling in wetlands, and assessment of land idling programs to reduce water consumption. Many of these studies are collaborations with various partners including Department of Interior agencies, Indian Tribes, and State agencies.

  12. ABUNDANCE AND ULTRASTRUCTURAL DIVERSITY OF NEURONAL GAP JUNCTIONS IN THE OFF AND ON SUBLAMINAE OF THE INNER PLEXIFORM LAYER OF RAT AND MOUSE RETINA

    PubMed Central

    KAMASAWA, N.; FURMAN, C. S.; DAVIDSON, K. G. V.; SAMPSON, J. A.; MAGNIE, A. R.; GEBHARDT, B. R.; KAMASAWA, M.; YASUMURA, T.; ZUMBRUNNEN, J. R.; PICKARD, G. E.; NAGY, J. I.; RASH, J. E.

    2007-01-01

    Neuronal gap junctions are abundant in both outer and inner plexiform layers of the mammalian retina. In the inner plexiform layer (IPL), ultrastructurally-identified gap junctions were reported primarily in the functionally-defined and anatomically-distinct ON sublamina, with few reported in the OFF sublamina. We used freeze-fracture replica immunogold labeling and confocal microscopy to quantitatively analyze the morphologies and distributions of neuronal gap junctions in the IPL of adult rat and mouse retina. Under “baseline” conditions (photopic illumination/general anesthesia), 649 neuronal gap junctions immunogold-labeled for connexin36 were identified in rat IPL, of which 375 were photomapped to OFF vs. ON sublaminae. In contrast to previous reports, the volume-density of gap junctions was equally abundant in both sublaminae. Five distinctive morphologies of gap junctions were identified: conventional crystalline and non-crystalline “plaques” (71% and 3%), plus unusual “string” (14%), “ribbon” (7%) and “reticular” (2%) forms. Plaque and reticular gap junctions were distributed throughout the IPL. However, string and ribbon gap junctions were restricted to the OFF sublamina, where they represented 48% of gap junctions in that layer. In string and ribbon junctions, curvilinear strands of connexons were dispersed over 5 to 20 times the area of conventional plaques having equal numbers of connexons. To define morphologies of gap junctions under different light-adaptation conditions, we examined an additional 1150 gap junctions from rats and mice prepared after 30 min of photopic, mesopic and scotopic illumination, with and without general anesthesia. Under these conditions, string and ribbon gap junctions remained abundant in the OFF sublamina and absent in the ON sublamina. Abundant gap junctions in the OFF sublamina of these two rodents with rod-dominant retinas revealed previously-undescribed but extensive pathways for inter-neuronal communication; and the wide dispersion of connexons in string and ribbon gap junctions suggests unique structural features of gap junctional coupling in the OFF vs. ON sublamina. PMID:17010526

  13. Effects of introduced crayfish on selected native fishes of Arizona

    USGS Publications Warehouse

    Carpenter, J.

    2000-01-01

    The virile crayfish (Orconectes virilis), an aggressive polytrophic species, has been introduced into many Arizona streams. I investigated competition and predation between this crayfish and several native Arizona fishes. I conducted field experiments to assess competition for food between crayfish and fish, and laboratory experiments to examine competition for shelter and food, and predation. In Sabino Creek, I manipulated crayfish densities in isolated pools to examine effects of crayfish on growth, mortality, and recruitment of Gila chub (Gila intermedia). Regardless of crayfish density, Gila chub declined slightly in weight and condition. Mortality and recruitment did not differ between densities of crayfish. I examined crayfish effects on benthic macroinvertebrates, a submerged aquatic macrophyte and associated invertebrates, and three fish species in a small stream in the White Mountains by fencing eight stream sections to prevent movement. The three fishes were speckled dace (Rhinichthys osculus), Sonora sucker (Catostomus insignis), and desert sucker (C. clarki). Molluscs > 10 mm and macrophytes were less abundant at sites with a high density of crayfish than at sites with low crayfish densities. Insect diversity was lower in high- vs. low-density sites. No treatment effect was observed on growth or condition of individually marked fish. Short-term laboratory experiments demonstrated predatory interactions and competition for shelter between crayfish and Gila chub, desert sucker, and speckled dace. Crayfish used shelter more than fish, displaced fish from shelter, and frequently attacked fish. Fish never attacked crayfish, and only once displaced crayfish from shelter. In predation experiments, crayfish preyed upon all species, but preyed most heavily upon desert suckers. Fish never altered use of the water column in the presence of crayfish. Density manipulation experiments in a laboratory measured food competition between crayfish and two native fishes. Growth of Gila chub was less affected by crayfish than by increased density of Gila chub. Thus crayfish are not strong competitors with Gila chub for food. However, growth of flannelmouth sucker (Catostomus latipinnis) was negatively impacted by presence of crayfish. These laboratory experiments provide evidence that introduced crayfish can reduce fish growth by competition for food, and that native fishes are vulnerable to crayfish predation.

  14. Egg deposition by lithophilic-spawning fishes in the Detroit and Saint Clair Rivers, 2005–14

    USGS Publications Warehouse

    Prichard, Carson G.; Craig, Jaquelyn M.; Roseman, Edward F.; Fischer, Jason L.; Manny, Bruce A.; Kennedy, Gregory W.

    2017-03-14

    A long-term, multiseason, fish egg sampling program conducted annually on the Detroit (2005–14) and Saint Clair (2010–14) Rivers was summarized to identify where productive fish spawning habitat currently exists. Egg mats were placed on the river bottom during the spring and fall at historic spawning areas and candidate fish spawning habitat restoration sites throughout both rivers. Widespread evidence was found of lithophilic spawning by numerous native fish species, including walleye (Sander vitreus), lake whitefish (Coregonus clupeaformis), lake sturgeon (Acipenser fulvescens), suckers (Catostomidae spp.), and trout-perch (Percopsis omiscomaycus). Walleye, lake whitefish, and suckers spp. spawned in nearly every region of each river in all years on both reef and nonreef substrates. Lake sturgeon eggs were collected almost exclusively over constructed reefs. Catch-per-unit effort of walleye, lake whitefish, and sucker eggs was much greater in the Detroit River than in the Saint Clair River, while Saint Clair River sites supported the greatest collections of lake sturgeon eggs. Collections during this study of lake sturgeon eggs on man-made spawning reefs suggest that artificial reefs may be an effective tool for restoring fish populations in the Detroit and Saint Clair Rivers; however, the quick response of lake sturgeon to spawn on newly constructed reefs and the fact that walleye, lake whitefish, and sucker eggs were often collected over substrate with little interstitial space to protect eggs from siltation and predators suggests that lack of suitable spawning habitat may continue to limit reproduction of lithophilic-spawning fish species in the Saint Clair-Detroit River System.

  15. Response of fish population dynamics to mitigation activities in a large regulated river

    USGS Publications Warehouse

    Watkins, Carson J.; Ross, Tyler J.; Quist, Michael C.; Hardy, Ryan S.

    2017-01-01

    Extensive water development in large rivers has precipitated many negative ecological effects on native fish populations. Mitigation for such development often focuses on restoring biological integrity through remediation of the physical and chemical properties of regulated rivers. However, evaluating and defining the success of those programs can be difficult. We modeled the influence of mitigation-related environmental factors on growth and recruitment of two ecologically important native fish species (Largescale Sucker Catostomus macrocheilus and Mountain Whitefish Prosopium williamsoni) in the Kootenai River, Idaho. Artificial nutrient (phosphorus) addition best predicted the variability in annual growth of both species. Nutrient addition was positively related to Largescale Sucker growth but negatively related to Mountain Whitefish growth. The best model explained 82% of the annual variability in incremental growth for Largescale Suckers and 61% of the annual variability for Mountain Whitefish. Year-class strength of Largescale Suckers was not closely related to any of the environmental variables evaluated; however, year-class strength of Mountain Whitefish was closely associated with nutrient addition, discharge, and temperature. Most research has focused on biotic assemblages to evaluate the effects of mitigation activities on fishes, but there is an increased need to identify the influence of rehabilitation activities on fish population dynamics within those assemblages. Here, we demonstrate how fish growth can serve as an indicator of rehabilitation success in a highly regulated large river. Future fish restoration projects can likely benefit from a change in scope and from consideration of an evaluation framework involving the response of population rate functions to mitigation.

  16. Self-recognition mechanism between skin and suckers prevents octopus arms from interfering with each other.

    PubMed

    Nesher, Nir; Levy, Guy; Grasso, Frank W; Hochner, Binyamin

    2014-06-02

    Controlling movements of flexible arms is a challenging task for the octopus because of the virtually infinite number of degrees of freedom (DOFs) [1, 2]. Octopuses simplify this control by using stereotypical motion patterns that reduce the DOFs, in the control space, to a workable few [2]. These movements are triggered by the brain and are generated by motor programs embedded in the peripheral neuromuscular system of the arm [3-5]. The hundreds of suckers along each arm have a tendency to stick to almost any object they contact [6-9]. The existence of this reflex could pose significant problems with unplanned interactions between the arms if not appropriately managed. This problem is likely to be accentuated because it is accepted that octopuses are "not aware of their arms" [10-14]. Here we report of a self-recognition mechanism that has a novel role in motor control, restraining the arms from interfering with each other. We show that the suckers of amputated arms never attach to octopus skin because a chemical in the skin inhibits the attachment reflex of the suckers. The peripheral mechanism appears to be overridden by central control because, in contrast to amputated arms, behaving octopuses sometime grab amputated arms. Surprisingly, octopuses seem to identify their own amputated arms, as they treat arms of other octopuses like food more often than their own. This self-recognition mechanism is a novel peripheral component in the embodied organization of the adaptive interactions between the octopus's brain, body, and environment [15, 16]. Copyright © 2014 Elsevier Ltd. All rights reserved.

  17. Distribution and relative abundance of fishes in littoral areas of Chief Joseph Reservoir, Columbia River

    USGS Publications Warehouse

    Gadomski, Dena M.; Venditti, David A.; Robinson, T. Craig; Beeman, John W.; Maule, Alec G.

    2004-01-01

    We surveyed fish assemblages in littoral areas of Chief Joseph Reservoir of the upper Columbia River to aid in understanding this ecosystem. Fish distributions and abundances were examined during April-July 1999 in relation to environmental conditions in the reservoir. We also compared the fish assemblages in Chief Joseph reservoir in 1999 to a past study conducted during 1974-1975, and to assemblages in other areas of the Columbia River. During 67 hr of electrofishing and 78 beach seine hauls in Chief Joseph Reservoir, 7460 fishes representing 8 families were collected. The majority of the catch was native – northern pikeminnow; redside shiners; longnose, bridgelip, and largescale suckers; and sculpins. The most abundant introduced species was walleye, and one species, rainbow trout, was mostly of net-pen origin. Larger sizes of suckers and northern pikeminnow were most abundant in the upper reservoir, likely due to upstream spawning migrations. The lower reservoir contained greater abundances of smaller fishes, and this area had lower flows, smaller substrates, and more complex shorelines that offered these fishes refugia. Only adult suckers displayed significant differences in abundances related to substrate. The relative abundances of species appeared to have changed since the 1970s, when the dominant fishes were northern pikeminnow, peamouth, largescale suckers, and walleye. Fish assemblage differences between Chief Joseph Reservoir and lower Columbia River reservoirs were also evident due to the morphology of the reservoir, its more northerly location, and the lack of fish passage facilities at Chief Joseph Dam. Our study is one of the few descriptions of fishes in the upper Columbia Rivers.

  18. Advanced stages of embryonic development and cotylocidial morphogenesis in the intrauterine eggs of Aspidogaster limacoides Diesing, 1835 (Aspidogastrea), with comments on their phylogenetic implications.

    PubMed

    Świderski, Zdzisław; Poddubnaya, Larisa G; Gibson, David I; Młocicki, Daniel

    2012-06-01

    Ultrastructural aspects of the advanced embryonic development and cotylocidial morphogenesis of the aspidogastrean Aspidogaster limacoides are described. The posterior or distal regions of the uterus are filled with eggs containing larvae at advanced stages of morphogenesis and fully-formed cotylocidia. Various stages and organs of this larva are described in detail, including the aspects of the developing and fully-differentiated cotylocidium, the body wall (tegument and musculature), glandular regions and the protonephridial excretory system. Blastomere multiplication by means of mitotic divisions takes place simultaneously with the degeneration or apoptosis of some micromeres; this frequently observed characteristic is compared and discussed in relation to corresponding reports for other neodermatans. During the advanced stages of the embryonic development of A. limacoides, the vitelline syncytium disappears and the size of the embryo increases rapidly. Evident polarization of the differentiating larva was observed; towards one pole of the egg, cytodifferentiation of the mouth, surrounded by the oral sucker and cephalic glands, takes place, whereas, towards the opposite pole, differentiation of the posterior sucker (incipient ventral disc) occurs. The oral and posterior suckers are formed from numerous embryonic cells which have differentiated into myocytes. The central part of the oral sucker undergoes invagination and forms the future pharynx and intestine. Fully-developed cotylocidia of A. limacoides have a neodermatan type of tegument, flame cells and two types of glandular structures. These results suggest a sister relationship between the Aspidogastrea and the Digenea, although the systematic position of aspidogastreans in relation to other platyhelminth taxa remains somewhat equivocal.

  19. Selenium impacts on razorback sucker, Colorado River, Colorado I. Adults.

    PubMed

    Hamilton, Steven J; Holley, Kathy M; Buhl, Kevin J; Bullard, Fern A; Ken Weston, L; McDonald, Susan F

    2005-05-01

    Adult razorback sucker (Xyrauchen texanus) were exposed to various selenium concentrations in ponds and isolated river channels of the Colorado River near Grand Junction, CO, to determine effects on their growth and residue accumulation over an 11-month period. Adults at Horsethief ponds were fed a commercial diet, whereas fish at Adobe Creek channel and North Pond foraged on natural food items. Selenium concentrations at Horsethief were 2.2 microg/L in water, 0.1-1.4 microg/g in sediment, and 2.3-3.1 microg/g in food organisms (1.1 microg/g in commercial fish food), at Adobe Creek were 3.8 microg/L in water, 0.5-2.1 microg/g in sediment, and 4-56 microg/g in food organisms, and at North Pond were 9.5 microg/L in water, 7-55 microg/g in sediment, and 20-81 microg/g in food organisms. The selenium concentrations in muscle plugs from adults at Adobe Creek (11.7 microg/g, SD = 0.4, n = 6) and North Pond (16.6 microg/g, SD = 1.0, n = 6) were greater than at Horsethief (4.5 microg/g, SD = 0.2, n = 6). During a depuration period adults from Adobe Creek and North Pond lost 1-2% of their selenium burden in 32 days and 14-19% in 66 days. Selenium accumulated in razorback sucker above toxic thresholds reported in other studies, yet those residues were less than those reported in muscle plugs of 40% of wild razorback sucker caught in the Green River, Utah.

  20. Description of Brachylecithum mackoi n. sp. (Digenea: Dicrocoeliidae) from the European hedgehog, Erinaceus europaeus (Insectivora: Erinaceidae).

    PubMed

    Casanova, Joan Caries; Ribas, Alexis

    2004-08-01

    A new hepatic dicrocoeliid species, Brachylecithum mackoi n. sp. (Digenea, Dicrocoeliidae), is described from the European hedgehog, Erinaceus europaeus (L.) (Insectivora, Erinaceidae). An infected host was found in the Mediterranean island of Elba (Italy), and more than 60 individuals were isolated from the biliary ducts. The holotype and 55 paratypes were examined. Brachylecithum mackoi n. sp. differs from congeneric species found in mammal hosts by having well-developed lappets in the ventral sucker, a sloping uterus between anterior testis and acetabulum, no overlap between vitellaria, and metrical features in the body size, sucker diameters, cirrus sac, and size of eggs. The only other Brachylecithum species of erinaceids in Europe and Africa, Brachylecithum aetechini Dollfus, 1951, differs from the new species in the above-mentioned morphological characters, greater dimensions of the body, and oral sucker, pharynx, cirrus sac, and egg dimensions. The presence of B. mackoi n. sp. in Elba Island is discussed in the light of apparent host specificity of erinaceid dicrocoeliids and geographical distribution of Palearctic and Ethiopian Erinaceidae.

  1. Transforming growth factor-β1 expression in endangered age-0 shortnose suckers (Chasmistes brevirostris) from Upper Klamath Lake, OR relative to histopathology, meristic, spatial, and temporal data.

    PubMed

    Ottinger, Christopher A; Densmore, Christine L; Robertson, Laura S; Iwanowicz, Deborah D; VanderKooi, Scott P

    2016-02-01

    During July-September of 2008, 2009, and 2010 endangered age-0 juvenile shortnose suckers were sampled from Upper Klamath Lake, OR in a health evaluation that included the measurement of transforming growth factor - beta (TGF-β) expression in spleen in combination with a histopathology assessment. This analysis was performed to determine if the expression of this immuno-regulator could be used as a component of a larger health evaluation intended to identify potential risk-factors that may help to explain why very few of these fish survive to age-1. Potential associations between TGF-β1 expression, histopathological findings, meristic data as well as temporal and spatial data were evaluated using analysis-of-variance. In this analysis, the absence or presence of opercula deformity and hepatic cell necrosis were identified as significant factors in accounting for the variance in TGF-β1 expression observed in age-0 shortnose suckers (n = 122, squared multiple R = 0.989). Location of sample collection and the absence or presence of anchor worms (Lernaea spp.) were identified as significant cofactors. The actual mechanisms involved with these relationships have yet to be determined. The strength, however, of our findings support the concept of using TGF-β1 expression as part of a broader fish health assessment and suggests the potential for using additional immunologic measures in future studies. Specifically, our results indicate that the measure of TGF-β1 expression in age-0 shortnose sucker health assessments can facilitate the process of identifying disease risks that are associated with the documented lack of recruitment into the adult population. Published by Elsevier Ltd.

  2. High diet overlap between native small-bodied fishes and nonnative fathead minnow in the Colorado River, Grand Canyon, Arizona

    USGS Publications Warehouse

    Seegert, Sarah E. Zahn; Rosi-Marshall, Emma J.; Baxter, Colden V.; Kennedy, Theodore A.; Hall, Robert O.; Cross, Wyatt F.

    2014-01-01

    River regulation may mediate the interactions among native and nonnative species, potentially favoring nonnative species and contributing to the decline of native populations. We examined food resource use and diet overlap among small-bodied fishes in the Grand Canyon section of the Colorado River as a first step in evaluating potential resource competition. We compared the diets of the predominant small-bodied fishes (native Speckled Dace Rhinichthys osculus, juvenile Flannelmouth Sucker Catostomus latipinnis, and juvenile Bluehead Sucker C. discobolus, and nonnative Fathead Minnow Pimephales promelas) across seasons at four sites downstream of Glen Canyon Dam using nonmetric multidimensional scaling and Schoener's similarity index. The diets of these fishes included diatoms, amorphous detritus, aquatic invertebrates (especially simuliid and chironomid larvae), terrestrial invertebrates, and terrestrial vegetation. Diets varied with season and were affected by high turbidity. Fish consumed more amorphous detritus and terrestrial vegetation during the summer monsoon season (July–September), when turbidity was higher. The diets of all species overlapped, but there was large variation in the degree of overlap. The diets of juvenile suckers and Fathead Minnows were most similar, while Speckled Dace had relatively distinct diets. The differences took the form of higher proportions of diatoms and amorphous detritus in the diets of Bluehead Suckers and Fathead Minnows and higher proportions of simuliids and chironomids in those of Speckled Dace. If food resources are or become limiting, diet overlap suggests that competition may occur among native and nonnative species, which could have implications for the population dynamics of these fishes and for the management of the Colorado River ecosystem in Grand Canyon.

  3. Hazard evaluation of inorganics, singly and in mixtures, to Flannelmouth Sucker Catostomus latipinnis in the San Juan River, New Mexico

    USGS Publications Warehouse

    Hamilton, S.J.; Buhl, K.J.

    1997-01-01

    Larval flannelmouth sucker (Catostomus latipinnis) were exposed to arsenate, boron, copper, molybdenum, selenate, selenite, uranium, vanadium, and zinc singly, and to five mixtures of five to nine inorganics. The exposures were conducted in reconstituted water representative of the San Juan River near Shiprock, New Mexico. The mixtures simulated environmental ratios reported for sites along the San Juan River (San Juan River backwater, Fruitland marsh, Hogback East Drain, Mancos River, and McElmo Creek). The rank order of the individual inorganics, from most to least toxic, was: copper > zinc > vanadium > selenite > selenate > arsenate > uranium > boron > molybdenum. All five mixtures exhibited additive toxicity to flannelmouth sucker. In a limited number of tests, 44-day-old and 13-day-old larvae exhibited no difference in sensitivity to three mixtures. Copper was the major toxic component in four mixtures (San Juan backwater, Hogback East Drain, Mancos River, and McElmo Creek), whereas zinc was the major toxic component in the Fruitland marsh mixture, which did not contain copper. The Hogback East Drain was the most toxic mixture tested. Comparison of 96-h LC50values with reported environmental water concentrations from the San Juan River revealed low hazard ratios for arsenic, boron, molybdenum, selenate, selenite, uranium, and vanadium, moderate hazard ratios for zinc and the Fruitland marsh mixture, and high hazard ratios for copper at three sites and four environmental mixtures representing a San Juan backwater, Hogback East Drain, Mancos River, and McElmo Creek. The high hazard ratios suggest that inorganic contaminants could adversely affect larval flannelmouth sucker in the San Juan River at four sites receiving elevated inorganics.

  4. Transforming growth factor-β1 expression in endangered age-0 shortnose suckers (Chasmistes brevirostris) from Upper Klamath Lake, OR relative to histopathology, meristic, spatial, and temporal data

    USGS Publications Warehouse

    Ottinger, Christopher A.; Densmore, Christine L.; Robertson, Laura S.; Iwanowicz, Deborah; Vanderkooi, Scott P.

    2016-01-01

    During July – September of 2008, 2009, and 2010 endangered age-0 juvenile shortnose suckers were sampled from Upper Klamath Lake, OR in a health evaluation that included the measurement of transforming growth factor – beta (TGF-β) expression in spleen in combination with a histopathology assessment. This analysis was performed to determine if the expression of this immuno-regulator could be used as a component of a larger health evaluation intended to identify potential risk-factors that may help to explain why very few of these fish survive to age-1. Potential associations between TGF-β1 expression, histopathological findings, meristic data as well as temporal and spatial data were evaluated using analysis-of-variance. In this analysis, the absence or presence of opercula deformity and hepatic cell necrosis were identified as significant factors in accounting for the variance in TGF-β1 expression observed in age-0 shortnose suckers (n = 122, squared multiple R = 0.989). Location of sample collection and the absence or presence of anchor worms (Lernaea spp.) were identified as significant cofactors. The actual mechanisms involved with these relationships have yet to be determined. The strength, however, of our findings support the concept of using TGF-β1 expression as part of a broader fish health assessment and suggests the potential for using additional immunologic measures in future studies. Specifically, our results indicate that the measure of TGF-β1 expression in age-0 shortnose sucker health assessments can facilitate the process of identifying disease risks that are associated with the documented lack of recruitment into the adult population.

  5. A new species of Saturnius Manter, 1969 (Digenea: Hemiuridae) from Mediterranean mullet (Teleostei: Mugilidae).

    PubMed

    Marzoug, Douniazed; Rima, Mohamed; Boutiba, Zitouni; Georgieva, Simona; Kostadinova, Aneta; Pérez-del-Olmo, Ana

    2014-02-01

    A new hemiurid digenean, Saturnius gibsoni n. sp., is described from the stomach lining of Mugil cephalus L. off Oran, Mediterranean coast of Algeria. Characteristic morphological features of the new species include small size of the body which is comprised of six pseudosegments, small ventral sucker, weakly developed mound-shaped flange at the level of the ventral sucker, and eggs being large in relation to the size of the body. Saturnius gibsoni n. sp. resembles S. minutus Blasco-Costa, Pankov, Gibson, Balbuena, Raga, Sarabeev & Kostadinova, 2006 and two unidentified Saturnius spp. in the small size of the body and most metrical features. However, in spite of the presence of five transverse septa resulting in six pseudosegments and the range overlap of some metrical features, the ventral sucker in S. minutus is much larger, the ventral sucker muscular flange is more prominent, the last pseudosegment is narrower in relation to body width and more rounded, and the eggs are smaller (mean 21 × 10 vs 25 × 12 μm). Furthermore, the partial sequences of the 28S rRNA gene region (domains D1-D3; 1,195 nt) obtained from two isolates of S. gibsoni n. sp. differed by 11 nt (0.9%) from that of S. minutus. Both unidentified forms of Saturnius are clearly distinguishable from S. gibsoni n. sp. by the presence of six stout, transverse muscular septa, forming seven pseudosegments (vs five septa forming six pseudosegments). Bayesian inference analysis of partial 28S rDNA sequences based on a total of 15 species from the families Hemiuridae and Lecithasteridae depicted the Bunocotylinae Dollfus, 1950 as a strongly supported basal clade, with Bunocotyle progenetica (Markowski, 1936) as the closest sister taxon to Saturnius spp.

  6. The surface topography of Callorhynchocotyle callorhynchi (Manter, 1955) (Monogenea: Hexabothriidae), a parasite of the holocephalan fish Callorhinchus capensis.

    PubMed

    Poddubnaya, Larisa G; Reed, Cecile; Gibson, David I

    2015-09-01

    A scanning electron microscopical study, incorporating some transmission electron microscopical observations, was undertaken on the surface topography of the gill parasite Callorhynchocotyle callorhynchi (Manter, 1955) (Monogenea: Hexabothriidae) from the Cape elephant fish Callorhinchus capensis (Holocephali) off the western coast of South Africa. The study revealed the presence of several new characteristics for this species. These include the presence of regularly distributed, knob-shaped projections on the surface of the haptor, haptoral appendix and sucker peduncles measuring 0.2 μm and in concentrations of approximately 100 per 10 μm(2) and the existence of a ridge which bisects each sucker lumen, forming two different loculi. We also report, for the first time for any monogenean, the presence of 'true spines'; these occur on the luminal surface of the haptoral suckers and have all of the characteristics of the tegumental spines of digeneans, i.e. they are situated within the distal syncytial tegumental cytoplasm, rest on the basal plasma membrane, have a uniform structure and are covered apically by the tegumental surface plasma membrane. These spines are simple, straight and single-pointed. Under the scanning electron microscopy (SEM), within an area of 20 μm in diameter, 23 such spines were counted, but their concentration and arrangement varies in different regions of the sucker. At their base, they measure about 1.5 μm in width and reach approximately 2 μm in height above the general level of the tegument, but transmission electron microscopy (TEM) measurements of an entire spine indicate that they may reach 3 μm in total length. The presence of spines, possessing similar morphological characteristics in both basal polyopisthocotylean monogeneans and digeneans, represents another characteristic which may prove useful in understanding the evolutionary relationships within the Neodermata.

  7. Selenium impacts on razorback sucker, Colorado River, Colorado: I. Adults

    USGS Publications Warehouse

    Hamilton, S.J.; Holley, K.M.; Buhl, K.J.; Bullard, F.A.; Weston, L.K.; McDonald, S.F.

    2005-01-01

    Adult razorback sucker (Xyrauchen texanus) were exposed to various selenium concentrations in ponds and isolated river channels of the Colorado River near Grand Junction, CO, to determine effects on their growth and residue accumulation over an 11-month period. Adults at Horsethief ponds were fed a commercial diet, whereas fish at Adobe Creek channel and North Pond foraged on natural food items. Selenium concentrations at Horsethief were 2.2 μg/L in water, 0.1–1.4 μg/g in sediment, and 2.3–3.1 μg/g in food organisms (1.1 μg/g in commercial fish food), at Adobe Creek were 3.8 μg/L in water, 0.5–2.1 μg/g in sediment, and 4–56 μg/g in food organisms, and at North Pond were 9.5 μg/L in water, 7–55 μg/g in sediment, and 20–81 μg/g in food organisms. The selenium concentrations in muscle plugs from adults at Adobe Creek (11.7 μg/g, SD=0.4, n=6) and North Pond (16.6 μg/g, SD=1.0, n=6) were greater than at Horsethief (4.5 μg/g, SD=0.2, n=6). During a depuration period adults from Adobe Creek and North Pond lost 1–2% of their selenium burden in 32 days and 14–19% in 66 days. Selenium accumulated in razorback sucker above toxic thresholds reported in other studies, yet those residues were less than those reported in muscle plugs of 40% of wild razorback sucker caught in the Green River, Utah.

  8. Effects of lake surface elevation on shoreline-spawning Lost River Suckers

    USGS Publications Warehouse

    Burdick, Summer M.; Hewitt, David A.; Rasmussen, J.E.; Hayes, Brian; Janney, Eric; Harris, Alta C.

    2015-01-01

    We analyzed remote detection data from PIT-tagged Lost River Suckers Deltistes luxatus at four shoreline spawning areas in Upper Klamath Lake, Oregon, to determine whether spawning of this endangered species was affected by low water levels. Our investigation was motivated by the observation that the surface elevation of the lake during the 2010 spawning season was the lowest in 38 years. Irrigation withdrawals in 2009 that were not replenished by subsequent winter-spring inflows caused a reduction in available shoreline spawning habitat in 2010. We compared metrics of skipped spawning, movement among spawning areas, and spawning duration across 8 years (2006-2013) that had contrasting spring water levels. Some aspects of sucker spawning were similar in all years, including few individuals straying from the shoreline areas to spawning locations in lake tributaries and consistent effects of increasing water temperatures on the accumulation of fish at the spawning areas. During the extreme low water year of 2010, 14% fewer female and 8% fewer male suckers joined the shoreline spawning aggregation than in the other years. Both males and females visited fewer spawning areas within Upper Klamath Lake in 2010 than in other years, and the median duration at spawning areas in 2010 was at least 36% shorter for females and 20% shorter for males relative to other years. Given the imperiled status of the species and the declining abundance of the population in Upper Klamath Lake, any reduction in spawning success and egg production could negatively impact recovery efforts. Our results indicate that lake surface elevations above 1,262.3-1,262.5 m would be unlikely to limit the number of spawning fish and overall egg production.

  9. Threats, conservation strategies, and prognosis for suckers (Catostomidae) in North America: insights from regional case studies of a diverse family of non-game fishes

    USGS Publications Warehouse

    Cooke, Steven J.; Bunt, Christopher M.; Hamilton, Steven J.; Jennings, Cecil A.; Pearson, Micheal P.; Cooperman, Michael S.; Markle, Douglas F.

    2005-01-01

    Catostomid fishes are a diverse family of 76+ freshwater species that are distributed across North America in many different habitats. This group of fish is facing a variety of impacts and conservation issues that are somewhat unique relative to more economically valuable and heavily managed fish species. Here, we present a brief series of case studies to highlight the threats such as migration barriers, flow regulation, environmental contamination, habitat degradation, exploitation and impacts from introduced (non-native) species that are facing catostomids in different regions. Collectively, the case studies reveal that individual species usually are not threatened by a single, isolated factor. Instead, species in general face numerous stressors that threaten multiple stages of their life history. Several factors have retarded sucker conservation including widespread inabilities of field workers to distinguish some species, lack of basic natural history and ecological knowledge of life history, and the misconception that suckers are tolerant of degraded conditions and are of little social or ecological value. Without a specific constituent group lobbying for conservation of non-game fishes, all such species, including members of the catostomid family, will continue to face serious risks because of neglect, ignorance, and misunderstanding. We suggest that conservation strategies should incorporate research and education/outreach components. Other conservation strategies that would be effective for protecting suckers include freshwater protected areas for critical habitat, restoration of degraded habitat, and design of catostomid-friendly fish bypass facilities. We believe that the plight of the catostomids is representative of the threats facing many other non-game freshwater fishes with diverse life-history strategies globally.

  10. Identification, immunolocalization, and characterization analyses of an exopeptidase of papain superfamily, (cathepsin C) from Clonorchis sinensis.

    PubMed

    Liang, Pei; He, Lei; Xu, Yanquan; Chen, Xueqing; Huang, Yan; Ren, Mengyu; Liang, Chi; Li, Xuerong; Xu, Jin; Lu, Gang; Yu, Xinbing

    2014-10-01

    Cathepsin C is an important exopeptidase of papain superfamily and plays a number of great important roles during the parasitic life cycle. The amino acid sequence of cathepsin C from Clonorchis sinensis (C. sinensis) showed 54, 53, and 49% identities to that of Schistosoma japonicum, Schistosoma mansoni, and Homo sapiens, respectively. Phylogenetic analysis utilizing the sequences of papain superfamily of C. sinensis demonstrated that cathepsin C and cathepsin Bs came from a common ancestry. Cathepsin C of C. sinensis (Cscathepsin C) was identified as an excretory/secretory product by Western blot analysis. The results of transcriptional level and translational level of Cscathepsin C at metacercaria stage were higher than that at adult worms. Immunolocalization analysis indicated that Cscathepsin C was specifically distributed in the suckers (oral sucker and ventral sucker), eggs, vitellarium, intestines, and testis of adult worms. In the metacercaria, it was mainly detected on the cyst wall and excretory bladder. Combining with the results mentioned above, it implies that Cscathepsin C may be an essential proteolytic enzyme for proteins digestion of hosts, nutrition assimilation, and immune invasion of C. sinensis. Furthermore, it may be a potential diagnostic antigen and drug target against C. sinensis infection.

  11. Razorback sucker movements and habitat use in the San Juan River inflow, Lake Powell, Utah, 1995-1997

    USGS Publications Warehouse

    Karp, C.A.; Mueller, G.

    2002-01-01

    Seventeen subadult, hatchery-reared razorback suckers (Xyrauchen texanus; (x̄ = 456 mm total length) were implanted with sonic transmitters and tracked for 23 months in the lower 89.6 km of the San Juan River (San Juan arm of Lake Powell, Utah). Fish were released at 2 sites, and 9 made extensive up-and downstream movements (x = 47.8 km; contact was lost with 4, and 4 others presumably died or lost their transmitters). The San Juan arm is primarily inundated canyon; however, most fish contacts occurred in shallow coves and shoreline with thick stands of flooded salt cedar in the upper inflow area. Eight fish frequented the Piute Farms river/lake mixing zone, and at least 4 moved upstream into the San Juan River. Seven fish were found in 2 aggregations in spring (3 fish in Neskahi Bay in 1996 and 4 fish just downstream of Piute Farms in 1997), and these may have been associated with spawning activity. Continued presence of razorback suckers in the Piute Farms area and lower San Juan River suggests the San Juan inflow to Lake Powell could be used as an alternate stocking site for reintroduction efforts.

  12. Communication using eye roll reflective signalling

    USGS Publications Warehouse

    Flamarique, I.N.; Mueller, G.A.; Cheng, C.L.; Figiel, C.R.

    2007-01-01

    Body reflections in the ultraviolet (UV) are a common occurrence in nature. Despite the abundance of such signals and the presence of UV cones in the retinas of many vertebrates, the function of UV cones in the majority of taxa remains unclear. Here, we report on an unusual communication system in the razorback sucker, Xyrauchen texanus, that involves flash signals produced by quick eye rolls. Behavioural experiments and field observations indicate that this form of communication is used to signal territorial presence between males. The flash signal shows highest contrast in the UV region of fhe visual spectrum (??max???380 nm), corresponding to the maximum wavelength of absorption of the UV cone mechanism in suckers. Furthermore, these cones are restricted to the dorsal retina of the animal and the upwelling light background is such that their relative sensitivity would be enhanced by chromatic adaptation of the other cone mechanisms. Thus, the UV cones in the sucker have optimal characteristics (both in terms of absorbance and retinal topography) to constitute the main detectors of the flash signal. Our findings provide the first ecological evidence for restricted distribution of UV cones in the retina of a vertebrate. ?? 2007 The Royal Society.

  13. Meteorological Sensor Array (MSA)-Phase I. Volume 2 (Data Management Tool: Proof of Concept)

    DTIC Science & Technology

    2014-10-01

    directory of next hourly file to read *** utcString = CStr (CInt(utcString) + 1) utcString = String(2 - Len(utcString), Ŕ...hourly file to read *** utcString = CStr (CInt(utcString) + 1) utcString = String(2 - Len(utcString), Ŕ") & utcString

  14. Description of rare echinostomatid fluke Echinodollfusia longiuscula n. sp. (Trematoda: Echinostomatidae) parasitizing white stork with an amended diagnosis of the genus Echinodollfusia.

    PubMed

    Macko, J K; Spakulová, M

    2014-04-01

    A new species of digenean, Echinodollfusia longiuscula n. sp. (Trematoda: Echinostomatidae), is described and illustrated based on specimens collected from the intestine of the white stork Ciconia ciconia (L.) (Aves: Ciconiidae) from eastern Slovakia and dissected soon after its spring arrival from African wintering. The new species differs from the two existing congeners Echinodollfusia stenon (Dollfus, 1950) and Echinodollfusia bulgarica Vassilev (Sofia 6:327-338, 1958) clearly by the oesophagus bearing diverticula, the bipartite seminal vesicle and gonads situated entirely in anterior body half. The new species E. longiuscula n. sp. can also be distinguished by possessing the following combination of features: a slender body of 12.85-21.06-mm long and maximum of 581-855-μm wide; head collar with 43-47 small collar spines, dorsal in double row; oral sucker more than half the size as ventral sucker, small pharynx, oval and in tandem testes situated pre-equatorial, small ovary at the boundary of anterior third of the body and anteriormost extent of the vitelline fields just reaching the level of posterior margin of the ventral sucker. An amended generic diagnosis of little-known Echinodollfusia Skrjabin et Bashkirova (Osnovy Trematodologii 12:51-930, 1956) is also given here.

  15. Demographics of the spawning aggregations of four catostomid species in the Savannah River, South Carolina and Georgia, USA

    USGS Publications Warehouse

    Grabowski, T.B.; Ratterman, N.L.; Isely, J.J.

    2008-01-01

    Differences in the life history strategies employed by otherwise ecologically similar species of a fish assemblage may be an important factor in the coexistence of these species and is an essential consideration in the conservation and management of these assemblages. We collected scales to determine age and growth of four species of the catostomid assemblage (northern hogsucker Hypentelium nigricans, spotted sucker Minytrema melanops, notchlip redhorse Moxostoma collapsum and robust redhorse Moxostoma robustum) of the Savannah River, Georgia-South Carolina in spring 2004 and 2005. Robust redhorse was the largest species; reaching sexual maturity at an older age and growing faster as a juvenile than the other species. Spotted sucker did not achieve the same size as robust redhorse, but reached sexual maturity at younger ages. Notchlip redhorse was intermediate between the abovementioned two species in age at maturity and size. Northern hogsucker was the smallest species of the assemblage and reached the sexual maturity at the age of three. Both robust redhorse and spotted sucker were sexually dimorphic in size-at-age. The range of life history strategies employed by Savannah River catostomids encompasses the range of life history strategies exhibited within the family as a whole. ?? 2007 Blackwell Munksgaard.

  16. Dispersal of larval suckers at the Williamson River Delta, Upper Klamath Lake, Oregon, 2006-09

    USGS Publications Warehouse

    Wood, Tamara M.; Hendrixson, Heather A.; Markle, Douglas F.; Erdman, Charles S.; Burdick, Summer M.; Ellsworth, Craig M.; Buccola, Norman L.

    2012-01-01

    An advection/diffusion modeling approach was used to simulate the transport of larval suckers from spawning areas in the Williamson River, through the newly restored Williamson River Delta, to Upper Klamath Lake. The density simulations spanned the years of phased restoration, from 2006/2007 prior to any levee breaching, to 2008 when the northern part of the delta was reconnected to the lake, and 2009 when levees on both sides of the delta had been breached. Model simulation results from all four years were compared to field data using rank correlation. Spearman ρ correlation coefficients were usually significant and in the range 0.30 to 0.60, providing moderately strong validation of the model. The correlation coefficients varied with fish size class in a way that suggested that the model best described the distribution of smaller fish near the Williamson River channel, and larger fish away from the channel. When Lost River and shortnose/Klamath largescale suckers were simulated independently, the correlation results suggested that the model better described the transport and dispersal of the latter species. The incorporation of night-time-only drift behavior in the Williamson River channel neither improved nor degraded correlations with field data. The model showed that advection by currents is an important factor in larval dispersal.

  17. An occupancy-based quantification of the highly imperiled status of desert fishes of the southwestern United States.

    PubMed

    Budy, Phaedra; Conner, Mary M; Salant, Nira L; Macfarlane, William W

    2015-08-01

    Desert fishes are some of the most imperiled vertebrates worldwide due to their low economic worth and because they compete with humans for water. An ecological complex of fishes, 2 suckers (Catostomus latipinnis, Catostomus discobolus) and a chub (Gila robusta) (collectively managed as the so-called three species) are endemic to the U.S. Colorado River Basin, are affected by multiple stressors, and have allegedly declined dramatically. We built a series of occupancy models to determine relationships between trends in occupancy, local extinction, and local colonization rates, identify potential limiting factors, and evaluate the suitability of managing the 3 species collectively. For a historical period (1889-2011), top performing models (AICc) included a positive time trend in local extinction probability and a negative trend in local colonization probability. As flood frequency decreased post-development local extinction probability increased. By the end of the time series, 47% (95% CI 34-61) and 15% (95% CI 6-33) of sites remained occupied by the suckers and the chub, respectively, and models with the 2 species of sucker as one group and the chub as the other performed best. For a contemporary period (2001-2011), top performing (based on AICc ) models included peak annual discharge. As peak discharge increased, local extinction probability decreased and local colonization probability increased. For the contemporary period, results of models that split all 3 species into separate groups were similar to results of models that combined the 2 suckers but not the chub. Collectively, these results confirmed that declines in these fishes were strongly associated with water development and that relative to their historic distribution all 3 species have declined dramatically. Further, the chub was distinct in that it declined the most dramatically and therefore may need to be managed separately. Our modeling approach may be useful in other situations in which targeted data are sparse and conservation status and best management approach for multiple species are uncertain. © 2015 Society for Conservation Biology.

  18. Biorobotic adhesion in water using suction cups.

    PubMed

    Bandyopadhyay, Promode R; Hrubes, J Dana; Leinhos, Henry A

    2008-03-01

    Echeneid fish, limpets and octopi use suction cups for underwater adhesion. When echeneid fish use suckers to 'hitch a ride' on sharks (which have riblet-patterned skins), the apparent absence of any pump or plumbing may be an advantage over biorobotic suction cups. An intriguing question is: How do they achieve seemingly persistent leak-free contact at low energy cost over rough surfaces? The design features of their suckers are explored in a biorobotic context of adhesion in water over rough surfaces. We have carried out experiments to compare the release force and tenacity of man-made suction cups with those reported for limpets and echeneid fish. Applied tensile and shear release forces were monotonically increased until release. The effects of cup size and type, host surface roughness, curvature and liquid surface tension have been examined. The flow of water in the sharkskin-like host surface roughness has been characterized. The average tenacity is 5.28 N cm(-2) (sigma = 0.53 N cm(-2), N = 37) in the sub-ambient pressure range of 14.6-49.0 kPa, in man-made cups for monotonically increasing applied release force. The tenacity is lower for harmonically oscillating release forces. The dynamic structural interactions between the suction cup and the oscillating applied forcing are discussed. Inspired by the matching of sharkskin riblet topology in echeneid fish suckers, it was found that biorobotic sealed contact over rough surfaces is also feasible when the suction cup makes a negative copy of the rough host surface. However, for protracted, persistent contact, the negative topology would have to be maintained by active means. Energy has to be spent to maintain the negative host roughness topology to minute detail, and protracted hitch-riding on sharks for feeding may not be free for echeneid fish. Further work is needed on the mechanism and efficiency of the densely populated tiny actuators in the fish suckers that maintain leak-proof contact with minimal energy cost and the feasibility of their biorobotic replication.

  19. Efficacy of hydrogen peroxide in controlling mortality associated with saprolegniasis on walleye, white sucker, and paddlefish eggs

    USGS Publications Warehouse

    Gaikowski, M.P.; Rach, J.J.; Drobish, M.; Hamilton, J.; Harder, T.; Lee, L.A.; Moen, C.; Moore, A.

    2003-01-01

    The efficacy of hydrogen peroxide in controlling saprolegniasis on eggs of walleye Stizostedion vitreum, white sucker Catostomus commersoni, and paddlefish Polyodon spathula was evaluated at four private, state, and federal production hatcheries participating in an Investigational New Animal Drug efficacy study (experiment 1; walleyes) and in a laboratory-based miniature egg jar incubation system (experiment 2; walleyes, white suckers, and paddlefish). Naturally occurring fungal infestations (saprolegniasis) were observed on eggs in both experiments. Confirmatory diagnosis of infested eggs from one hatchery in experiment 1 identified the pathogen as Saprolegnia parasitica. During experiment 1, eggs were treated daily for 15 min with either 0, 500, or 750 mg/L of hydrogen peroxide, and one trial compared a 500-mg/L hydrogen peroxide treatment with a formalin treatment at 1,667 mg/L. Saprolegniasis infestation was observed in control egg jars, whereas treatment with either formalin or hydrogen peroxide virtually eliminated the infestation. Hydrogen peroxide treatments of 500 mg/L either increased egg hatch or were as effective as physical removal of infested eggs in controlling mortality. Although treatment with formalin at 1,667 mg/L significantly increased the percent eye-up of walleye eggs compared with that of those treated with hydrogen peroxide at 500 mg/L, the difference was only 1.9-2.6%. In experiment 2, noneyed eggs were treated for 15 min every other day with 0, 283, 565, or 1,130 mg/L of hydrogen peroxide until the viable eggs hatched. Saprolegniasis infestation engulfed most control eggs, whereas infestation of treated eggs was either reduced or not visible. Hydrogen peroxide significantly increased egg hatch for all three species tested in experiment 2. Although hydrogen peroxide treatments as low as 283 mg/L significantly increased walleye and white sucker hatch, treatments between 500 and 1,000 mg/L are more likely to be effective in production egg incubation systems.

  20. Re-evaluation of the species composition of Bashkirovitrema Skrjabin, 1944 (Digenea: Echinostomatidae), with the description of two new species of this genus and the proposal of Kostadinovatrema novaeguiniense n. g., n. sp.

    PubMed

    Dronen, Norman O

    2009-11-01

    A comparison of specimens previously identified as Bashkirovitrema incrassatum (Diesing, 1850) from the African or cape clawless otter Aonyx capensis, and the speckle-throated or spotted-throated otter Hydrictis maculicollis from the Old World with specimens and descriptions of B. incrassatum from the New World showed that those from Africa (Bashkirovitrema africanum n. sp.) had a more extensive distribution of the vitelline fields than either B. canadense n. sp. from the northern river otter Lontra canadensis and the American mink Neovison vison (North America) and B. incrassatum from the Neotropical river otter Lontra longicaudis (South America). B. africanum n. sp. further differs from B. canadense n. sp. by having a smaller body, shorter forebody, smaller oral sucker, longer cirrus, shorter intertesticular space, shorter post-testicular space and longer eggs. B. canadense n. sp. can be distinguished from B. incrassatum by having a longer body, longer forebody, smaller sucker ratio, smaller testes, greater distance between the ventral sucker and the ovary, shorter cirrus-sac, longer post-testicular space and narrower eggs. B. africanum n. sp. differs from B. incrassatum by having a smaller ventral sucker, larger ovary, shorter cirrus-sac, smaller intertesticular space, larger post-testicular space and longer eggs that are not as wide. Kostadinovatrema n. g., as represented by K. novaeguiniense n. sp., can be separated from species of Bashkirovitrema Skrjabin, 1944 by having a wider body width to length profile, a head collar that is narrower than the forebody and armed with 33 rather than 27 collar spines, an ovary that is further removed anteriorly from the testes, and a dorso-ventrally flattened hindbody that is nearly as broad as the forebody. The new genus differs from Hypoderaeum Dietz, 1909 and Moliniella Hübner, 1939 by having 33 head collar spines.

  1. Relationships between bald eagle productivity and dynamics of fish populations and fisheries in the Wisconsin waters of Lake Superior, 1983-1999

    USGS Publications Warehouse

    Hoff, Michael H.; Meyer, Michael W.; Van Stappen, Julie; Fratt, Thomas W.

    2004-01-01

    Bald eagle (Haliaeetus leucocephalus) abundance declined in the 1950s and 1960s along the Wisconsin waters of Lake Superior, and were nearly absent along Wisconsin's Lake Superior shoreline. The population began to increase again between 1980 and 1983, and since then bald eagles nesting on islands along Wisconsin's Lake Superior shoreline (i.e., Apostle Islands) reproduced at a lower rate than have those nesting along the mainland shoreline of the lake and inland. Recent research indicated that bioaccumulation of toxic chemicals in the aquatic food chain no longer limits bald eagle reproduction there, and that productivity at island nests was lower than at mainland nests and inland nests as the result of low food availability. Management agencies have sought models that accurately predict productivity and explain ecological relationships, but no satisfactory models had previously been developed. Modeling was conducted here to determine which factors best explained productivity variability. The Ricker stock-recruitment model derived from only the bivariate breeding pair and productivity data explained only 1% of the variability in productivity data. The functional relationship that explained the greatest amount of productivity variation (83%) included the number of breeding pairs, burbot (Lota lota) biomass, longnose sucker (Catostomus catostomus) biomass, and commercial harvest of nontarget fishes. Model results were interpreted to mean that productivity was positively affected by populations of burbot and longnose sucker, which are important prey items, and by commercial fishermen feeding nontarget fish to bald eagles. Harvest of nontarget fishes by tribal fishermen and burbot and longnose sucker populations have not tended to change during the entire study period, although the burbot population has declined since 1991. Therefore, bald eagle productivity is not predicted to increase unless burbot, longnose sucker, or other preferred prey of bald eagles increases in the Apostle Islands.

  2. Influence of weight and type of planting material on fruit quality and its heterogeneity in pineapple [Ananas comosus (L.) Merrill].

    PubMed

    Fassinou Hotegni, V Nicodème; Lommen, Willemien J M; Agbossou, Euloge K; Struik, Paul C

    2014-01-01

    Cultural practices can affect the quality of pineapple fruits and its variation. The objectives of this study were to investigate (a) effects of weight class and type of planting material on fruit quality, heterogeneity in quality and proportion and yield of fruits meeting European export standards, and (b) the improvement in quality, proportion and yield of fruits meeting export standards when flowering was induced at optimum time. Experiments were conducted in Benin with cvs Sugarloaf (a Perola type) and Smooth Cayenne. In cv. Sugarloaf, experimental factors were weight class of planting material (light, mixed, heavy) and time of flowering induction (farmers', optimum) (Experiment 1). In cv. Smooth Cayenne an additional experimental factor was the type of planting material (hapas, ground suckers, a mixture of the two) (Experiment 2). Fruits from heavy planting material had higher infructescence and fruit weights, longer infructescences, shorter crowns, and smaller crown: infructescence length than fruits from light planting material. The type of planting material in Experiment 2 did not significantly affect fruit quality except crown length: fruits from hapas had shorter crowns than those from ground suckers. Crops from heavy planting material had a higher proportion and yield of fruits meeting export standards than those from other weight classes in Experiment 1 only; also the type of planting material in Experiment 2 did not affect these variates. Heterogeneity in fruit quality was usually not reduced by selecting only light or heavy planting material instead of mixing weights; incidentally the coefficient of variation was significantly reduced in fruits from heavy slips only. Heterogeneity was also not reduced by not mixing hapas and ground suckers. Flowering induction at optimum time increased the proportion and yield of fruits meeting export standards in fruits from light and mixed slip weights and in those from the mixture of heavy hapas plus ground suckers.

  3. Influence of weight and type of planting material on fruit quality and its heterogeneity in pineapple [Ananas comosus (L.) Merrill

    PubMed Central

    Fassinou Hotegni, V. Nicodème; Lommen, Willemien J. M.; Agbossou, Euloge K.; Struik, Paul C.

    2015-01-01

    Cultural practices can affect the quality of pineapple fruits and its variation. The objectives of this study were to investigate (a) effects of weight class and type of planting material on fruit quality, heterogeneity in quality and proportion and yield of fruits meeting European export standards, and (b) the improvement in quality, proportion and yield of fruits meeting export standards when flowering was induced at optimum time. Experiments were conducted in Benin with cvs Sugarloaf (a Perola type) and Smooth Cayenne. In cv. Sugarloaf, experimental factors were weight class of planting material (light, mixed, heavy) and time of flowering induction (farmers', optimum) (Experiment 1). In cv. Smooth Cayenne an additional experimental factor was the type of planting material (hapas, ground suckers, a mixture of the two) (Experiment 2). Fruits from heavy planting material had higher infructescence and fruit weights, longer infructescences, shorter crowns, and smaller crown: infructescence length than fruits from light planting material. The type of planting material in Experiment 2 did not significantly affect fruit quality except crown length: fruits from hapas had shorter crowns than those from ground suckers. Crops from heavy planting material had a higher proportion and yield of fruits meeting export standards than those from other weight classes in Experiment 1 only; also the type of planting material in Experiment 2 did not affect these variates. Heterogeneity in fruit quality was usually not reduced by selecting only light or heavy planting material instead of mixing weights; incidentally the coefficient of variation was significantly reduced in fruits from heavy slips only. Heterogeneity was also not reduced by not mixing hapas and ground suckers. Flowering induction at optimum time increased the proportion and yield of fruits meeting export standards in fruits from light and mixed slip weights and in those from the mixture of heavy hapas plus ground suckers. PMID:25653659

  4. One-dimensional reduction of viscous jets. I. Theory

    NASA Astrophysics Data System (ADS)

    Pitrou, Cyril

    2018-04-01

    We build a general formalism to describe thin viscous jets as one-dimensional objects with an internal structure. We present in full generality the steps needed to describe the viscous jets around their central line, and we argue that the Taylor expansion of all fields around that line is conveniently expressed in terms of symmetric trace-free tensors living in the two dimensions of the fiber sections. We recover the standard results of axisymmetric jets and we report the first and second corrections to the lowest order description, also allowing for a rotational component around the axis of symmetry. When applied to generally curved fibers, the lowest order description corresponds to a viscous string model whose sections are circular. However, when including the first corrections, we find that curved jets generically develop elliptic sections. Several subtle effects imply that the first corrections cannot be described by a rod model since it amounts to selectively discard some corrections. However, in a fast rotating frame, we find that the dominant effects induced by inertial and Coriolis forces should be correctly described by rod models. For completeness, we also recover the constitutive relations for forces and torques in rod models and exhibit a missing term in the lowest order expression of viscous torque. Given that our method is based on tensors, the complexity of all computations has been beaten down by using an appropriate tensor algebra package such as xAct, allowing us to obtain a one-dimensional description of curved viscous jets with all the first order corrections consistently included. Finally, we find a description for straight fibers with elliptic sections as a special case of these results, and recover that ellipticity is dynamically damped by surface tension. An application to toroidal viscous fibers is presented in the companion paper [Pitrou, Phys. Rev. E 97, 043116 (2018), 10.1103/PhysRevE.97.043116].

  5. Pathogenesis of spring viremia of carp virus in emerald shiner Notropis atherinoides Rafinesque, fathead minnow Pimephales promelas Rafinesque and white sucker Catostomus commersonii (Lacepede).

    PubMed

    Misk, E; Garver, K; Nagy, E; Isaac, S; Tubbs, L; Huber, P; Al-Hussinee, L; Lumsden, J S

    2016-06-01

    Spring viremia of carp (SVC) is a reportable disease to the World Organization of Animal Health (OIE) as it is known to cause significant international economic impact. In Canada, the first and only isolation of SVC virus (SVCV) was in 2006, from common carp Cyprinus carpio L., at Hamilton Harbour, Lake Ontario. The susceptibility of fathead minnow Pimephales promelas Rafinesque, emerald shiner Notropis atherinoides Rafinesque and white sucker Catostomus commersonii (Lacepede) to intraperitoneal injection of the Canadian isolate (HHOcarp06) was evaluated using experimental infection, virus isolation, quantitative reverse transcription polymerase chain reaction (qRT-PCR), histopathology and immunohistochemistry (IHC). Emerald shiner and fathead minnow were most susceptible with 43 and 53% cumulative mortality, respectively, compared with koi at 33%. Quantitative RT-PCR demonstrated that koi had high viral loads throughout the experiment. At 34 days post-infection, SVCV was detected from sampled emerald shiner and white sucker in very low titre and was not detected from fathead minnow. Koi, fathead minnow and emerald shiner had gross lesions typical of SVC disease. The histopathological picture was mostly dominated by necrotic changes in kidney, spleen, liver, pancreas and intestine. IHC further confirmed SVCV infection, and staining was largely correlated with histological lesions. © 2015 John Wiley & Sons Ltd.

  6. Suckers for Science.

    ERIC Educational Resources Information Center

    Haugen, Heidi Helene

    2001-01-01

    Introduces an inquiry-based program on leeches that features five components: (1) engagement; (2) exploration; (3) explanation; (4) evaluation; and (5) extension/elaboration. Investigates the anatomy and environmental conditions of leeches. (YDS)

  7. String scattering amplitudes and deformed cubic string field theory

    NASA Astrophysics Data System (ADS)

    Lai, Sheng-Hong; Lee, Jen-Chi; Lee, Taejin; Yang, Yi

    2018-01-01

    We study string scattering amplitudes by using the deformed cubic string field theory which is equivalent to the string field theory in the proper-time gauge. The four-string scattering amplitudes with three tachyons and an arbitrary string state are calculated. The string field theory yields the string scattering amplitudes evaluated on the world sheet of string scattering whereas the conventional method, based on the first quantized theory brings us the string scattering amplitudes defined on the upper half plane. For the highest spin states, generated by the primary operators, both calculations are in perfect agreement. In this case, the string scattering amplitudes are invariant under the conformal transformation, which maps the string world sheet onto the upper half plane. If the external string states are general massive states, generated by non-primary field operators, we need to take into account carefully the conformal transformation between the world sheet and the upper half plane. We show by an explicit calculation that the string scattering amplitudes calculated by using the deformed cubic string field theory transform into those of the first quantized theory on the upper half plane by the conformal transformation, generated by the Schwarz-Christoffel mapping.

  8. Covariant open bosonic string field theory on multiple D-branes in the proper-time gauge

    NASA Astrophysics Data System (ADS)

    Lee, Taejin

    2017-12-01

    We construct a covariant open bosonic string field theory on multiple D-branes, which reduces to a non-Abelian group Yang-Mills gauge theory in the zero-slope limit. Making use of the first quantized open bosonic string in the proper time gauge, we convert the string amplitudes given by the Polyakov path integrals on string world sheets into those of the second quantized theory. The world sheet diagrams generated by the constructed open string field theory are planar in contrast to those of the Witten's cubic string field theory. However, the constructed string field theory is yet equivalent to the Witten's cubic string field theory. Having obtained planar diagrams, we may adopt the light-cone string field theory technique to calculate the multi-string scattering amplitudes with an arbitrary number of external strings. We examine in detail the three-string vertex diagram and the effective four-string vertex diagrams generated perturbatively by the three-string vertex at tree level. In the zero-slope limit, the string scattering amplitudes are identified precisely as those of non-Abelian Yang-Mills gauge theory if the external states are chosen to be massless vector particles.

  9. E(lementary)-strings in six-dimensional heterotic F-theory

    NASA Astrophysics Data System (ADS)

    Choi, Kang-Sin; Rey, Soo-Jong

    2017-09-01

    Using E-strings, we can analyze not only six-dimensional superconformal field theories but also probe vacua of non-perturabative heterotic string. We study strings made of D3-branes wrapped on various two-cycles in the global F-theory setup. We claim that E-strings are elementary in the sense that various combinations of E-strings can form M-strings as well as heterotic strings and new kind of strings, called G-strings. Using them, we show that emissions and combinations of heterotic small instantons generate most of known six-dimensional superconformal theories, their affinizations and little string theories. Taking account of global structure of compact internal geometry, we also show that special combinations of E-strings play an important role in constructing six-dimensional theories of D- and E-types. We check global consistency conditions from anomaly cancellation conditions, both from five-branes and strings, and show that they are given in terms of elementary E-string combinations.

  10. Gravitational Scattering Amplitudes and Closed String Field Theory in the Proper-Time Gauge

    NASA Astrophysics Data System (ADS)

    Lee, Taejin

    2018-01-01

    We construct a covariant closed string field theory by extending recent works on the covariant open string field theory in the proper-time gauge. Rewriting the string scattering amplitudes generated by the closed string field theory in terms of the Polyakov string path integrals, we identify the Fock space representations of the closed string vertices. We show that the Fock space representations of the closed string field theory may be completely factorized into those of the open string field theory. It implies that the well known Kawai-Lewellen-Tye (KLT) relations of the first quantized string theory may be promoted to the second quantized closed string theory. We explicitly calculate the scattering amplitudes of three gravitons by using the closed string field theory in the proper-time gauge.

  11. Hot string soup: Thermodynamics of strings near the Hagedorn transition

    NASA Astrophysics Data System (ADS)

    Lowe, David A.; Thorlacius, Lárus

    1995-01-01

    Above the Hagedorn energy density closed fundamental strings form a long string phase. The dynamics of weakly interacting long strings is described by a simple Boltzmann equation which can be solved explicitly for equilibrium distributions. The averge total number of long strings grows logarithmically with total energy in the microcanonical ensemble. This is consistent with calculations of the free single string density of states provided the thermodynamic limit is carefully defined. If the theory contains open strings the long string phase is suppressed.

  12. 7 CFR 51.597 - U.S. Grade A.

    Code of Federal Regulations, 2011 CFR

    2011-01-01

    ... celery of similar varietal characteristics which are fairly well developed and have fairly good heart..., freezing, suckers, growth cracks, hollow crown, pithy branches, seedstems, rust, cracked stem, other...

  13. Status and trends of adult Lost River (Deltistes luxatus) and shortnose (Chasmistes brevirostris) sucker populations in Upper Klamath Lake, Oregon, 2015

    USGS Publications Warehouse

    Hewitt, David A.; Janney, Eric C.; Hayes, Brian S.; Harris, Alta C.

    2017-07-21

    Executive SummaryData from a long-term capture-recapture program were used to assess the status and dynamics of populations of two long-lived, federally endangered catostomids in Upper Klamath Lake, Oregon. Lost River suckers (LRS; Deltistes luxatus) and shortnose suckers (SNS; Chasmistes brevirostris) have been captured and tagged with passive integrated transponder (PIT) tags during their spawning migrations in each year since 1995. In addition, beginning in 2005, individuals that had been previously PIT-tagged were re-encountered on remote underwater antennas deployed throughout sucker spawning areas. Captures and remote encounters during the spawning season in spring 2015 were incorporated into capture-recapture analyses of population dynamics. Cormack-Jolly-Seber (CJS) open population capture-recapture models were used to estimate annual survival probabilities, and a reverse-time analog of the CJS model was used to estimate recruitment of new individuals into the spawning populations. In addition, data on the size composition of captured fish were examined to provide corroborating evidence of recruitment. Separate analyses were done for each species and also for each subpopulation of LRS. Shortnose suckers and one subpopulation of LRS migrate into tributary rivers to spawn, whereas the other LRS subpopulation spawns at groundwater upwelling areas along the eastern shoreline of the lake. Characteristics of the spawning migrations in 2015, such as the effects of temperature on the timing of the migrations, were similar to past years.Capture-recapture analyses for the LRS subpopulation that spawns at the shoreline areas included encounter histories for 13,617 individuals, and analyses for the subpopulation that spawns in the rivers included 39,321 encounter histories. With a few exceptions, the survival of males and females in both subpopulations was high (greater than or equal to 0.86) between 1999 and 2013. Survival was notably lower for males from the rivers in 2000, 2006, and 2012. Survival probabilities were lower for males from the shoreline areas in 2002. Between 2001 and 2014, the abundance of males in the lakeshore spawning subpopulation decreased by at least 59 percent and the abundance of females decreased by at least 53 percent. By combining information from capture-recapture models and size composition data, we concluded that the abundance of both sexes in the river spawning subpopulation of LRS likely has decreased at rates similar to the rates for the lakeshore spawning subpopulation between 2002 and 2014. Capture-recapture analyses for SNS included encounter histories for 20,981 individuals. Most annual survival estimates between 2005 and 2009 were high (greater than 0.88), but both sexes of SNS experienced lower and more variable survival in 2001–04 and 2010–13. The best-case scenario for SNS, based on capture-recapture recruitment modeling, indicates that the abundance of males in the spawning population decreased by 77 percent and the abundance of females decreased by 74 percent between 2001 and 2014. Decreases in abundance for both sexes likely are greater than these estimates indicate. Despite relatively high survival in most years, we conclude that both species have experienced substantial decreases in the abundance of spawning adults because losses from mortality have not been balanced by recruitment of new individuals. The status of the endangered sucker populations in Upper Klamath Lake remains worrisome, especially for SNS.

  14. Native Fish Sanctuary Project - Sanctuary Development Phase, 2007 Annual Report

    USGS Publications Warehouse

    Mueller, Gordon A.

    2007-01-01

    Notable progress was made in 2007 toward the development of native fish facilities in the Lower Colorado River Basin. More than a dozen facilities are, or soon will be, online to benefit native fish. When this study began in 2005 no self-supporting communities of either bonytail or razorback sucker existed. Razorback suckers were removed from Rock Tank in 1997 and the communities at High Levee Pond had been compromised by largemouth bass in 2004. This project reversed that trend with the establishment of the Davis Cove native fish community in 2005. Bonytail and razorback sucker successfully produced young in Davis Cove in 2006. Bonytail successfully produced young in Parker Dam Pond in 2007, representing the first successful sanctuary established solely for bonytail. This past year, Three Fingers Lake received 135 large razorback suckers, and Federal and State agencies have agreed to develop a cooperative management approach dedicating a portion of that lake toward grow-out and (or) the establishment of another sanctuary. Two ponds at River's Edge Golf Course in Needles, California, were renovated in June and soon will be stocked with bonytail. Similar activities are taking place at Mohave Community College, Cerbat Cliffs Golf Course, Cibola High Levee Pond, Office Cove, Emerald Canyon Golf Course, and Bulkhead Cove. Recruitment can be expected as fish become sexually mature at these facilities. Flood-plain facilities have the potential to support 6,000 adult razorback suckers and nearly 20,000 bonytail if native fish management is aggressively pursued. This sanctuary project has assisted agencies in developing 15 native fish communities by identifying specific resource objectives for those sites, listing and prioritizing research opportunities and needs, and strategizing on management approaches through the use of resource-management plans. Such documents have been developed for Davis Cove, Cibola High Levee Pond, Parker Dam Pond, and Three Fingers Lake. We anticipate similar documents will be developed in the near future for River's Edge Golf Course Ponds, Office Cove, Emerald Canyon Golf Course Ponds, Bulkhead Cove, Mohave Community College, and Cerbat Cliffs Golf Course ponds as these facilities come on line or are developed in the future. The following report discusses the process that went into the development of these facilities. Sites were visited, assessed as to their suitability based on the control of nonnative predators, habitat suitability, conversion cost, logistics, geographical location, and willingness of landowners. They were then prioritized according to their suitability, cost, timely conversion, and willingness of landowners. Existing native fish facilities were included in this evaluation for their value in helping to determine physical and biological parameter ranges. This report describes the approaches that led to success, those leading to failure, and some of the biological, institutional, and management issues of implementing native fish sanctuary development.

  15. 7 CFR 51.598 - U.S. Grade B.

    Code of Federal Regulations, 2011 CFR

    2011-01-01

    ... celery of similar varietal characteristics which are fairly well developed, which are clean, well trimmed... by crater rot, brown stem, wilting, cutworms, freezing, suckers, growth cracks, hollow crown, pithy...

  16. Estimating geographic variation on allometric growth and body condition of Blue Suckers with quantile regression

    USGS Publications Warehouse

    Cade, B.S.; Terrell, J.W.; Neely, B.C.

    2011-01-01

    Increasing our understanding of how environmental factors affect fish body condition and improving its utility as a metric of aquatic system health require reliable estimates of spatial variation in condition (weight at length). We used three statistical approaches that varied in how they accounted for heterogeneity in allometric growth to estimate differences in body condition of blue suckers Cycleptus elongatus across 19 large-river locations in the central USA. Quantile regression of an expanded allometric growth model provided the most comprehensive estimates, including variation in exponents within and among locations (range = 2.88–4.24). Blue suckers from more-southerly locations had the largest exponents. Mixed-effects mean regression of a similar expanded allometric growth model allowed exponents to vary among locations (range = 3.03–3.60). Mean relative weights compared across selected intervals of total length (TL = 510–594 and 594–692 mm) in a multiplicative model involved the implicit assumption that allometric exponents within and among locations were similar to the exponent (3.46) for the standard weight equation. Proportionate differences in the quantiles of weight at length for adult blue suckers (TL = 510, 594, 644, and 692 mm) compared with their average across locations ranged from 1.08 to 1.30 for southern locations (Texas, Mississippi) and from 0.84 to 1.00 for northern locations (Montana, North Dakota); proportionate differences for mean weight ranged from 1.13 to 1.17 and from 0.87 to 0.95, respectively, and those for mean relative weight ranged from 1.10 to 1.18 and from 0.86 to 0.98, respectively. Weights for fish at longer lengths varied by 600–700 g within a location and by as much as 2,000 g among southern and northern locations. Estimates for the Wabash River, Indiana (0.96–1.07 times the average; greatest increases for lower weights at shorter TLs), and for the Missouri River from Blair, Nebraska, to Sioux City, Iowa (0.90–1.00 times the average; greatest decreases for lower weights at longer TLs), were examined in detail to explain the additional information provided by quantile estimates.

  17. Sediment Dynamics Affecting the Threatened Santa Ana Sucker in the Highly-modified Santa Ana River and Inset Channel, Southern California, USA

    NASA Astrophysics Data System (ADS)

    Minear, J. T.; Wright, S. A.

    2015-12-01

    In this study, we investigate the sediment dynamics of the low-flow channel of the Santa Ana River that is formed by wastewater discharges and contains some of the last remaining habitat of the Santa Ana Sucker (Catostomus santaanae). The Santa Ana River is a highly-modified river draining the San Bernardino Mountains and Inland Empire metropolitan area east of Los Angeles. Home to over 4 million people, the watershed provides habitat for the federally-threatened Santa Ana Sucker, which presently reside within the mainstem Santa Ana River in a reach supported by year-round constant discharges from water treatment plants. The nearly constant low-flow wastewater discharges and infrequent runoff events create a small, approximately 8 m wide, inset channel within the approximately 300 m wide mainstem channel that is typically dry except for large flood flows. The sediment dynamics within the inset channel are characterized by constantly evolving bed substrate and sediment transport rates, and occasional channel avulsions. The sediment dynamics have large influence on the Sucker, which rely on coarse-substrate (gravel and cobble) for their food production. In WY 2013 through the present, we investigated the sediment dynamics of the inset channel using repeat bathymetric and substrate surveys, bedload sampling, and discharge measurements. We found two distinct phases of the inset channel behavior: 1. 'Reset' flows, where sediment-laden mainstem discharges from upstream runoff events result in sand deposition in the inset channel or avulse the inset channel onto previously dry riverbed; and 2. 'Winnowing' flows, whereby the sand within the inset channel is removed by clear-water low flows from the wastewater treatment plant discharges. Thus, in contrast to many regulated rivers where high flows are required to flush fine sediments from the bed (for example, downstream from dams), in the Santa Ana River the low flows from wastewater treatment plants serve as the flushing flows that gradually remove sand and expose the coarser substrate. In effect, the inset channel is analogous to a flume subject to periodic sediment loading events from upstream (runoff events) with long periods of negligible upstream sediment supply between the events (wastewater discharges).

  18. Using spatial, seasonal, and diel drift patterns of larval Lost River suckers Deltistes luxatus (Cypriniformes: Catostomidae) and shortnose suckers Chasmistes brevirostris (Cypriniformes: Catostomidae) to help identify a site for a water withdrawal structure on the Williamson River, Oregon

    USGS Publications Warehouse

    Ellsworth, Craig M.; Tyler, Torrey J.; VanderKooi, Scott P.

    2010-01-01

    A small irrigation diversion dam near Chiloquin, Oregon, was removed and replaced with a pump station to improve fish passage for Lost River suckers (Deltistes luxatus) and shortnose suckers (Chasmistes brevirostris) entering the Sprague River on their spawning migrations. During the developmental phase of the pump station, a need was identified to better understand the larval drift characteristics of these endangered catostomids in order to reduce entrainment into the irrigation system. The spatial, seasonal, and diel distribution of drifting larvae was measured during the 2004 spawning season at two proposed sites on the Williamson River where the pump station could be located. Larval drift for both species coincided with the irrigation season making them subject to entrainment into the irrigation system. Drift occurred almost exclusively at night with larvae entering the drift at sunset and exiting the drift at sunrise. Nighttime larval densities were concentrated near the surface and at midchannel at both sites. Densities were generally greater on the side of mid-channel with greater flow. During early morning sampling we detected a general shift in larval drift from surface to subsurface drift. We also observed an increase in larval densities towards the shore opposite from the proposed pump station at the upper site whereas larval densities remained high at midchannel at the lower site. During daytime sampling, the few larvae that were collected were distributed throughout the water column at both pump sites. This study found that larvae drifting during all time periods were generally distributed further across the cross section, deeper in the water column, and closer to where the proposed water withdrawal structure would be built at the downstream site when compared to the upstream site. Recommendations were provided to locate the withdrawal facility at the upstream site and operate it in a manner such that larval entrainment would likely be minimized.

  19. Nearshore habitat and fish community associations of coaster brook trout in Isle Royale, Lake Superior

    USGS Publications Warehouse

    Gorman, O.T.; Moore, S.A.; Carlson, A.J.; Quinlan, H.R.

    2008-01-01

    We characterized the nearshore habitat and fish community composition of approximately 300 km of shoreline within and adjacent to the major embayments of Isle Royale, Lake Superior. Sampling yielded 17 species, of which 12 were widespread and represented a common element of the Lake Superior fish community, including cisco Coregonus artedi, lake whitefish C. clupeaformis, round whitefish Prosopium cylindraceum, lake trout Salvelinus namaycush, rainbow smelt Osmerus mordax, lake chub Couesius plumbeus, longnose sucker Catostomus catostomus, white sucker C. commersonii, trout-perch Percopsis omiscomaycus, ninespine stickleback Pungitius pungitius, burbot Lota lota, and slimy sculpin Cottus cognatus. The presence of brook trout S. fontinalis in an embayment was associated with the common species of the Isle Royale nearshore fish community, particularly cisco, longnose sucker, and round whitefish. However, brook trout were present in only five embayments and were common only in Tobin Harbor. Most Isle Royale embayments had broadly overlapping ranges of nearshore habitats. Within embayments, fish were distributed along a habitat gradient from less-protected rocky habitat near the mouth to highly protected habitat with mixed and finer substrates at the head. Embayments with brook trout had greater mean protection from the open lake, greater variation in depth, greater mean cover, and higher mean frequencies of large substrates (cobble, boulder, and bedrock). Within those embayments, brook trout were associated with habitat patches with higher mean frequencies of small substrates (particularly sand and coarse gravel). Within Tobin Harbor, brook trout were associated with midembayment habitat and species assemblages, especially those locations with a mixture of sand, gravel, and cobble substrates, an absence of bedrock, and the presence of round whitefish, white sucker, and trout-perch. Comparison of embayments with the model, Tobin Harbor, showed that six embayments without brook trout had very similar arrays of habitat. However, four embayments with brook trout had relatively different arrays of habitat from Tobin Harbor. These results suggest that there is potential for further recovery of brook trout populations across Isle Royale nearshore habitats. ?? Copyright by the American Fisheries Society 2008.

  20. Cosmic strings and superconducting cosmic strings

    NASA Technical Reports Server (NTRS)

    Copeland, Edmund

    1988-01-01

    The possible consequences of forming cosmic strings and superconducting cosmic strings in the early universe are discussed. Lecture 1 describes the group theoretic reasons for and the field theoretic reasons why cosmic strings can form in spontaneously broken gauge theories. Lecture 2 discusses the accretion of matter onto string loops, emphasizing the scenario with a cold dark matter dominated universe. In lecture 3 superconducting cosmic strings are discussed, as is a mechanism which leads to the formation of structure from such strings.

  1. Quaking aspen productivity recovers after repeated prescribed fire.

    Treesearch

    D. A. Perala

    1995-01-01

    Describes how quaking aspen (Populus tremuloides Michx.) stand recovered after logging, and logging and burning. Aspen suckering was profuse after each destructive episode but differences in stockability caused different yield trajectories.

  2. From decay to complete breaking: pulling the strings in SU(2) Yang-Mills theory.

    PubMed

    Pepe, M; Wiese, U-J

    2009-05-15

    We study {2Q+1} strings connecting two static charges Q in (2+1)D SU(2) Yang-Mills theory. While the fundamental {2} string between two charges Q=1/2 is unbreakable, the adjoint {3} string connecting two charges Q=1 can break. When a {4} string is stretched beyond a critical length, it decays into a {2} string by gluon pair creation. When a {5} string is stretched, it first decays into a {3} string, which eventually breaks completely. The energy of the screened charges at the ends of a string is well described by a phenomenological constituent gluon model.

  3. Subterranean barriers, methods, and apparatuses for forming, inspecting, selectively heating, and repairing same

    DOEpatents

    Nickelson, Reva A.; Sloan, Paul A.; Richardson, John G.; Walsh, Stephanie; Kostelnik, Kevin M.

    2009-04-07

    A subterranean barrier and method for forming same are disclosed, the barrier including a plurality of casing strings wherein at least one casing string of the plurality of casing strings may be affixed to at least another adjacent casing string of the plurality of casing strings through at least one weld, at least one adhesive joint, or both. A method and system for nondestructively inspecting a subterranean barrier is disclosed. For instance, a radiographic signal may be emitted from within a casing string toward an adjacent casing string and the radiographic signal may be detected from within the adjacent casing string. A method of repairing a barrier including removing at least a portion of a casing string and welding a repair element within the casing string is disclosed. A method of selectively heating at least one casing string forming at least a portion of a subterranean barrier is disclosed.

  4. 75 FR 30425 - Endangered and Threatened Species Permit Applications

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-06-01

    ..., during normal business hours at the U.S. Fish and Wildlife Service, 500 Gold Ave., SW., Room 6034... transport studies for Colorado pikeminnow (Ptychochelius lucius) and razorback sucker (Xyrauchen texanus) in...

  5. Development of wall climbing robot

    NASA Astrophysics Data System (ADS)

    Kojima, Hisao; Toyama, Ryousei; Kobayashi, Kengo

    1992-03-01

    A configuration design is presented for a wall-climbing robot with high payload which is capable of moving on diversified surfaces of walls including the wall surface to ceilings in every direction. A developed quadruped wall climbing robot, NINJYA-1, is introduced. NINJYA-1 is composed of legs based on a 3D parallel link mechanism and a VM (Valve-regulated Multiple) sucker which will be able to suck even if there are grooves and a small difference in level. A wall climbing robot which supports rescue operation at a high building using a VM sucker is also introduced. Finally, a wall climbing robot named Disk Rover with a disk-type magnetic wheel is shown. The wheel shape is calculated by FEM. The disk-type magnetic wheel has a force three times more powerful than the one heretofore in use.

  6. Random versus fixed-site sampling when monitoring relative abundance of fishes in headwater streams of the upper Colorado River basin

    USGS Publications Warehouse

    Quist, M.C.; Gerow, K.G.; Bower, M.R.; Hubert, W.A.

    2006-01-01

    Native fishes of the upper Colorado River basin (UCRB) have declined in distribution and abundance due to habitat degradation and interactions with normative fishes. Consequently, monitoring populations of both native and nonnative fishes is important for conservation of native species. We used data collected from Muddy Creek, Wyoming (2003-2004), to compare sample size estimates using a random and a fixed-site sampling design to monitor changes in catch per unit effort (CPUE) of native bluehead suckers Catostomus discobolus, flannelmouth suckers C. latipinnis, roundtail chub Gila robusta, and speckled dace Rhinichthys osculus, as well as nonnative creek chub Semotilus atromaculatus and white suckers C. commersonii. When one-pass backpack electrofishing was used, detection of 10% or 25% changes in CPUE (fish/100 m) at 60% statistical power required 50-1,000 randomly sampled reaches among species regardless of sampling design. However, use of a fixed-site sampling design with 25-50 reaches greatly enhanced the ability to detect changes in CPUE. The addition of seining did not appreciably reduce required effort. When detection of 25-50% changes in CPUE of native and nonnative fishes is acceptable, we recommend establishment of 25-50 fixed reaches sampled by one-pass electrofishing in Muddy Creek. Because Muddy Creek has habitat and fish assemblages characteristic of other headwater streams in the UCRB, our results are likely to apply to many other streams in the basin. ?? Copyright by the American Fisheries Society 2006.

  7. Population status, demography and habitat preferences of the threatened lipstick palm Cyrtostachys renda Blume in Kerumutan Reserve, Sumatra

    NASA Astrophysics Data System (ADS)

    Widyatmoko, Didik; Burgman, Mark A.; Guhardja, Edi; Mogea, Johanis P.; Walujo, Eko B.; Setiadi, Dede

    2005-09-01

    Population status and demography of a population of the threatened lipstick palm Cyrtostachys renda in a peat swamp ecosystem of Kerumutan Reserve, Sumatra (one of the largest remaining populations) was documented at 16 different sites, covering a wide range of forest and habitat types, vegetation associations, and population sizes. Population sizes were dominated by suckers comprising 89% of the total population. Individuals with stem heights between 0 and 4 m (47.5%), stem diameters between 4 and 10 cm (82.0%), and leaf scar numbers between 0 and 60 (69.2%) dominated. Ages of individuals were estimated and used to fit a curvilinear relationship between age and stem height. Wild plants reach reproductive maturity within 25-30 years, or when they have stem heights in excess of 2.0 m, or when they have 15-25 leaf scars. They can survive more than 80 years. Cultivated plants appear to reproduce earlier and produce more seeds than wild plants. Individual growth was plant size-dependent with the adult stage being the most productive. Higher mortality was experienced by suckers, especially in continuously waterlogged conditions and locations with dense canopies. Sucker growth was faster than seedling growth, an adaptation that may allow the species to cope with periodically waterlogged conditions. Population abundances varied with habitat types; well-drained areas were the most suitable habitat. To conserve the most important remaining populations of the lipstick palm, it is crucial to protect well-drained sites in Kerumutan Reserve.

  8. Age, growth, and reproductive biology of three catostomids from the Apalachicola River, Florida

    USGS Publications Warehouse

    Grabowski, Timothy B.; Young, S.P.; Isely, J.J.; Ely, Patrick C.

    2012-01-01

    Riverine catostomids can show a wide range of interspecific variation in life-history characteristics. Understanding these differences is an important consideration in evaluating the sensitivity of these fishes to disturbance and in formulating effective conservation strategies, particularly when dealing with an assemblage consisting of multiple species within a watershed. We collected Apalachicola redhorse Moxostoma n. sp. cf. poecilurum (n = 125), spotted sucker Minytrema melanops (n = 94), and quillback Carpiodes cyprinus (n = 94) to determine age, growth, and reproductive biology of spawning catostomids in the Apalachicola River, Florida, during 2007. Quillback was the smallest in total length at age; longest-lived; most fecund; and produced the smallest eggs. Apalachicola redhorse was the largest in body size; had an intermediate life span; and produced the fewest yet largest eggs. Spotted sucker was more similar to Apalachicola redhorse in most characteristics. Growth during ages 1-3 in all three species seemed to be negatively related to the proportion of observations of extreme flow, both high (Q90) and low (Q10), per year and a positive response in growth rate to high flows (>Q75 but < Q90). However, Apalachicola redhorse and spotted sucker growth was more sensitive to flow conditions than that of quillback. Our results suggest the life histories and ecological response of Apalachicola River catostomids to flow regulation are important components for developing strategies that incorporate the needs of these fishery resources into an ecosystem-based management approach.

  9. Informing New String Programmes: Lessons Learned from an Australian Experience

    ERIC Educational Resources Information Center

    Murphy, Fintan; Rickard, Nikki; Gill, Anneliese; Grimmett, Helen

    2011-01-01

    Although there are many examples of notable string programmes there has been relatively little comparative analysis of these programmes. This paper examines three benchmark string programmes (The University of Illinois String Project, The Tower Hamlets String Teaching Project and Colourstrings) alongside Music4All, an innovative string programme…

  10. [ital N]-string vertices in string field theory

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Bordes, J.; Abdurrahman, A.; Anton, F.

    1994-03-15

    We give the general form of the vertex corresponding to the interaction of an arbitrary number of strings. The technique employed relies on the comma'' representation of string field theory where string fields and interactions are represented as matrices and operations between them such as multiplication and trace. The general formulation presented here shows that the interaction vertex of [ital N] strings, for any arbitrary [ital N], is given as a function of particular combinations of matrices corresponding to the change of representation between the full string and the half string degrees of freedom.

  11. The Birth of String Theory

    NASA Astrophysics Data System (ADS)

    Cappelli, Andrea; Castellani, Elena; Colomo, Filippo; Di Vecchia, Paolo

    2012-04-01

    Part I. Overview: 1. Introduction and synopsis; 2. Rise and fall of the hadronic string G. Veneziano; 3. Gravity, unification, and the superstring J. H. Schwarz; 4. Early string theory as a challenging case study for philosophers E. Castellani; Part II. The Prehistory: The Analytic S-Matrix: 5. Introduction to Part II; 6. Particle theory in the sixties: from current algebra to the Veneziano amplitude M. Ademollo; 7. The path to the Veneziano model H. R. Rubinstein; 8. Two-component duality and strings P. G. O. Freund; 9. Note on the prehistory of string theory M. Gell-Mann; Part III. The Dual Resonance Model: 10. Introduction to Part III; 11. From the S-matrix to string theory P. Di Vecchia; 12. Reminiscence on the birth of string theory J. A. Shapiro; 13. Personal recollections D. Amati; 14. Early string theory at Fermilab and Rutgers L. Clavelli; 15. Dual amplitudes in higher dimensions: a personal view C. Lovelace; 16. Personal recollections on dual models R. Musto; 17. Remembering the 'supergroup' collaboration F. Nicodemi; 18. The '3-Reggeon vertex' S. Sciuto; Part IV. The String: 19. Introduction to Part IV; 20. From dual models to relativistic strings P. Goddard; 21. The first string theory: personal recollections L. Susskind; 22. The string picture of the Veneziano model H. B. Nielsen; 23. From the S-matrix to string theory Y. Nambu; 24. The analogue model for string amplitudes D. B. Fairlie; 25. Factorization in dual models and functional integration in string theory S. Mandelstam; 26. The hadronic origins of string theory R. C. Brower; Part V. Beyond the Bosonic String: 27. Introduction to Part V; 28. From dual fermion to superstring D. I. Olive; 29. Dual models with fermions: memoirs of an early string theorist P. Ramond; 30. Personal recollections A. Neveu; 31. Aspects of fermionic dual models E. Corrigan; 32. The dual quark models K. Bardakci and M. B. Halpern; 33. Remembering the dawn of relativistic strings J.-L. Gervais; 34. Early string theory in Cambridge: personal recollections C. Montonen; Part VI. The Superstring: 35. Introduction to Part VI; 36. Supersymmetry in string theory F. Gliozzi; 37. Gravity from strings: personal reminiscences of early developments T. Yoneya; 38. From the Nambu-Goto to the σ-model action L. Brink; 39. Locally supersymmetric action for superstring P. Di Vecchia; 40. Personal recollections E. Cremmer; 41. The scientific contributions of Joël Scherk J. H. Schwarz; Part VII. Preparing the String Renaissance: 42. Introduction to Part VII; 43. From strings to superstrings: a personal perspective M. B. Green; 44. Quarks, strings and beyond A. M. Polyakov; 45. The rise of the superstring theory A. Cappelli and F. Colomo; Appendices; Index.

  12. String resistance detector

    NASA Technical Reports Server (NTRS)

    Hall, A. Daniel (Inventor); Davies, Francis J. (Inventor)

    2007-01-01

    Method and system are disclosed for determining individual string resistance in a network of strings when the current through a parallel connected string is unknown and when the voltage across a series connected string is unknown. The method/system of the invention involves connecting one or more frequency-varying impedance components with known electrical characteristics to each string and applying a frequency-varying input signal to the network of strings. The frequency-varying impedance components may be one or more capacitors, inductors, or both, and are selected so that each string is uniquely identifiable in the output signal resulting from the frequency-varying input signal. Numerical methods, such as non-linear regression, may then be used to resolve the resistance associated with each string.

  13. String mediated phase transitions

    NASA Technical Reports Server (NTRS)

    Copeland, ED; Haws, D.; Rivers, R.; Holbraad, S.

    1988-01-01

    It is demonstrated from first principles how the existence of string-like structures can cause a system to undergo a phase transition. In particular, the role of topologically stable cosmic string in the restoration of spontaneously broken symmetries is emphasized. How the thermodynamic properties of strings alter when stiffness and nearest neighbor string-string interactions are included is discussed.

  14. Black string in dRGT massive gravity

    NASA Astrophysics Data System (ADS)

    Tannukij, Lunchakorn; Wongjun, Pitayuth; Ghosh, Suchant G.

    2017-12-01

    We present a cylindrically symmetric solution, both charged and uncharged, which is known as a black string solution to the nonlinear ghost-free massive gravity found by de Rham, Gabadadze, and Tolley (dRGT). This "dRGT black string" can be thought of as a generalization of the black string solution found by Lemos. Moreover, the dRGT black string solution includes other classes of black string solution such as the monopole-black string ones since the graviton mass contributes to the global monopole term as well as the cosmological-constant term. To investigate the solution, we compute mass, temperature, and entropy of the dRGT black string. We found that the existence of the graviton mass drastically affects the thermodynamics of the black string. Furthermore, the Hawking-Page phase transition is found to be possible for the dRGT black string as well as the charged dRGT black string. The dRGT black string solution is thermodynamically stable for r>r_c with negative thermodynamical potential and positive heat capacity while it is unstable for r

  15. Kite-flying: a unique but dangerous mode of electrical injury in children.

    PubMed

    Tiwari, V K; Sharma, D

    1999-09-01

    A retrospective study was conducted to evaluate the cause of a sudden rise in number of pediatric admissions with electrical injuries at our centre during the year 1998. In evaluating the cause, six out of twelve admissions were found to be related to kite-flying which is a popular sport during the months of June, July, August and September. In two out of six cases current travelled directly through the string of the kite. In two others, flame burns occured following ignition of clothing. Another patient had contact with wire through a metal rod. In the last case, arcing pulled the hand of the patient leading to direct contact with wire. The average burns size was approximately 31% body surface area (BSA), with all patients having burns over the palmar aspect of at least one hand. No patient required amputation for the injuries. In this article, attention has been focussed on the various modes of electrical injuries associated with kite-flying and some measures have been advised to avoid such accidents.

  16. Isotope effect in acetylene C2H2 and C2D2 rotations on Cu(001)

    NASA Astrophysics Data System (ADS)

    Shchadilova, Yulia E.; Tikhodeev, Sergei G.; Paulsson, Magnus; Ueba, Hiromu

    2014-04-01

    A comprehensive analysis of the elementary processes behind the scanning tunneling microscope controlled rotation of C2H2 and C2D2, isotopologues of a single acetylene molecule adsorbed on the Cu(001) surface, is given, with a focus on the isotope effects. With the help of density-functional theory we calculate the vibrational modes of C2H2 and C2D2 on Cu(001) and estimate the anharmonic couplings between them, using a simple strings-on-rods model. The probability of the elementary processes, nonlinear and combination band, is estimated using the Keldysh diagram technique. This allows us to clarify the main peculiarities and the isotope effects of the C2H2 and C2D2 on Cu(001) rotation, discovered in the pioneering work [B. C. Stipe et al., Phys. Rev. Lett. 81, 1263 (1998), 10.1103/PhysRevLett.81.1263], which have not been previously understood.

  17. The "Magic" String

    ERIC Educational Resources Information Center

    Hoover, Todd F.

    2010-01-01

    The "Magic" String is a discrepant event that includes a canister with what appears to be the end of two strings protruding from opposite sides of it. Due to the way the strings are attached inside the canister, it appears as if the strings can magically switch the way they are connected. When one string end is pulled, the observer's expectation…

  18. Automatic generation and analysis of solar cell IV curves

    DOEpatents

    Kraft, Steven M.; Jones, Jason C.

    2014-06-03

    A photovoltaic system includes multiple strings of solar panels and a device presenting a DC load to the strings of solar panels. Output currents of the strings of solar panels may be sensed and provided to a computer that generates current-voltage (IV) curves of the strings of solar panels. Output voltages of the string of solar panels may be sensed at the string or at the device presenting the DC load. The DC load may be varied. Output currents of the strings of solar panels responsive to the variation of the DC load are sensed to generate IV curves of the strings of solar panels. IV curves may be compared and analyzed to evaluate performance of and detect problems with a string of solar panels.

  19. Charged string loops in Reissner-Nordström black hole background

    NASA Astrophysics Data System (ADS)

    Oteev, Tursinbay; Kološ, Martin; Stuchlík, Zdeněk

    2018-03-01

    We study the motion of current carrying charged string loops in the Reissner-Nordström black hole background combining the gravitational and electromagnetic field. Introducing new electromagnetic interaction between central charge and charged string loop makes the string loop equations of motion to be non-integrable even in the flat spacetime limit, but it can be governed by an effective potential even in the black hole background. We classify different types of the string loop trajectories using effective potential approach, and we compare the innermost stable string loop positions with loci of the charged particle innermost stable orbits. We examine string loop small oscillations around minima of the string loop effective potential, and we plot radial profiles of the string loop oscillation frequencies for both the radial and vertical modes. We construct charged string loop quasi-periodic oscillations model and we compare it with observed data from microquasars GRO 1655-40, XTE 1550-564, and GRS 1915+105. We also study the acceleration of current carrying string loops along the vertical axis and the string loop ejection from RN black hole neighbourhood, taking also into account the electromagnetic interaction.

  20. Effect of Notched Strings on Tennis Racket Spin Performance: Ultrahigh-Speed Video Analysis of Spin Rate, Contact Time, and Post-Impact Ball Velocity

    NASA Astrophysics Data System (ADS)

    Kawazoe, Yoshihiko; Takeda, Yukihiro; Nakagawa, Masamichi

    While some tennis racket strings have more grip than others do, this does not guarantee that they will impart more spin to a tennis ball. Experiments with hand-held rackets are required to determine the longstanding question of how players can discern that different strings behave differently when laboratory tests indicate that they should play the same. In a previous study, we clarified the top-spin mechanism of a tennis racket by using high-speed video analysis on a tennis court for the first time. Furthermore, we improved it by using lubricated notched nylon strings. These experiments revealed that the more the main strings stretch and bend laterally, the more spin is imparted to the ball. This is due to the restoring force being parallel to the string face when the main strings spring back and the ball is released from the strings. Notched strings reduce the spin rate, but this can be effectively counteracted by employing lubricants. Furthermore, we found that imparting more spin reduces shock vibrations on the wrist during impact. The present study revealed that a ball has a 40% lower spin rate when hit with a racket with notched strings than with one with unnotched strings in the case of nylon (it had to be determined whether new strings or lubricated used strings give more spin). The experiments also showed that 30% more spin is imparted to a ball when the string intersections are lubricated by oil than when notched used nylon strings are used. Furthermore, we found that used natural gut notched strings reduced the spin rate by 70% compared to when new natural gut unnotched strings are used. We also investigated different top-spin behaviors obtained when professional and amateur tennis players hit a ball.

  1. Status and trends of adult Lost River (Deltistes luxatus) and shortnose (Chasmistes brevirostris) sucker populations in Upper Klamath Lake, Oregon, 2017

    USGS Publications Warehouse

    Hewitt, David A.; Janney, Eric C.; Hayes, Brian S.; Harris, Alta C.

    2018-04-24

    Executive SummaryData from a long-term capture-recapture program were used to assess the status and dynamics of populations of two long-lived, federally endangered catostomids in Upper Klamath Lake, Oregon. Lost River suckers (LRS; Deltistes luxatus) and shortnose suckers (SNS; Chasmistes brevirostris) have been captured and tagged with passive integrated transponder (PIT) tags during their spawning migrations in each year since 1995. In addition, beginning in 2005, individuals that had been previously PIT-tagged were re-encountered on remote underwater antennas deployed throughout sucker spawning areas. Captures and remote encounters during the spawning season in spring 2016 were incorporated into capture-recapture analyses of population dynamics.Cormack-Jolly-Seber (CJS) open population capture-recapture models were used to estimate annual survival probabilities, and a reverse-time analog of the CJS model was used to estimate recruitment of new individuals into the spawning populations. In addition, data on the size composition of captured fish were examined to provide corroborating evidence of recruitment. Model estimates of survival and recruitment were used to derive estimates of changes in population size over time and to determine the status of the populations through 2015. Separate analyses were done for each species and also for each subpopulation of LRS. Shortnose suckers and one subpopulation of LRS migrate into tributary rivers to spawn, whereas the other LRS subpopulation spawns at groundwater upwelling areas along the eastern shoreline of the lake.Capture-recapture analyses indicated that with a few exceptions, the survival of males and females in both Lost River sucker subpopulations was high (greater than 0.88) from 1999 to 2015. Survival was notably lower for males from the river in 2000, 2006, and 2012, and for the shoreline areas in 2002. From 2001 to 2015, the abundance of males in the lakeshore spawning subpopulation decreased by at least 64 percent and the abundance of females decreased by at least 56 percent. Capture-recapture models suggested that the abundance of both sexes in the river spawning subpopulation of LRS had increased substantially since 2006; increases were mostly due to large estimated recruitment events in 2006 and 2008. We know that the estimates in 2006 are substantially biased in favor of recruitment because of a sampling issue. We are skeptical of the magnitude of recruitment indicated by the 2008 estimates as well because (1) few small individuals that would indicate the presence of new recruits were captured in that year, and (2) recapture probabilities in recruitment models based on just physical recaptures of fish were lower than desired for robust inferences from capture-recapture models. If we assume instead that little or no recruitment occurred for this subpopulation, the abundance of both sexes in the river spawning subpopulation likely has decreased at rates similar to the rates for the lakeshore spawning subpopulation from 2002 to 2015.Shortnose suckers experienced lower and more variable annual survival than either LRS subpopulation. Annual survival of both sexes was relatively low in 2003, 2004, 2010, and 2012. In addition, female survival was low in 1999 and 2000 while male survival was low in 2002. Survival estimate precision in early years of the study; however, are poor. Capture-recapture models and size composition data indicate that recruitment of new individuals into the SNS spawning population was trivial from 2001 to 2005. Models indicate that more than 10 percent of the population was new recruits in a number of more recent years. As a result, capture-recapture modeling suggests that the abundance of adult spawning SNS was relatively stable from 2006 to 2010. We are skeptical of the estimated recruitment in 2006 because of the known sampling issue. We also are skeptical of the estimated recruitment in other recent years because few small individuals that would indicate the presence of new recruits were captured in any of those years, and recapture probabilities in recruitment models were low. The best-case scenario for SNS, based on capture-recapture recruitment modeling, indicates that the abundance of males in the spawning population decreased by 78 percent and the abundance of females decreased by 77 percent from 2001 to 2015. Decreases in abundance for both sexes are likely greater than these estimates indicate.Despite relatively high survival in most years, we conclude that both species have experienced substantial decreases in the abundance of spawning adults because losses from mortality have not been balanced by recruitment of new individuals. Although capture-recapture data indicate substantial recruitment of new individuals into the spawning populations for SNS and river spawning LRS in some years, size data do not corroborate these estimates. As a result, the status of the endangered sucker populations in Upper Klamath Lake remains distressed, especially for SNS. Our monitoring program provides a robust platform for estimating vital population parameters, evaluating the status of the populations, and assessing the effectiveness of conservation and recovery efforts.

  2. Syntactic transfer in artificial grammar learning.

    PubMed

    Beesley, T; Wills, A J; Le Pelley, M E

    2010-02-01

    In an artificial grammar learning (AGL) experiment, participants were trained with instances of one grammatical structure before completing a test phase in which they were required to discriminate grammatical from randomly created strings. Importantly, the underlying structure used to generate test strings was different from that used to generate the training strings. Despite the fact that grammatical training strings were more similar to nongrammatical test strings than they were to grammatical test strings, this manipulation resulted in a positive transfer effect, as compared with controls trained with nongrammatical strings. It is suggested that training with grammatical strings leads to an appreciation of set variance that aids the detection of grammatical test strings in AGL tasks. The analysis presented demonstrates that it is useful to conceptualize test performance in AGL as a form of unsupervised category learning.

  3. Effects of Wastewater Discharges on Endocrine and Reproductive Function of Western Mosquitofish (Gambusia spp.) and Implications for the Threatened Santa Ana Sucker (Catostomus santaanae)

    USGS Publications Warehouse

    Jenkins, Jill A.; Goodbred, Steven L.; Olivier, Heather M.; Draugelis-Dale, Rassa O.; Alvarez, David A.

    2009-01-01

    The Santa Ana River (SAR) in southern California is impacted by effluents from wastewater treatment plants (WWTP), which are sources of organic wastewater compounds (OWCs) and urban runoff. The Santa Ana River is one of only three river basins supporting native populations of the federally listed Santa Ana sucker (Catostomus santaanae) at the time the fish was included on the list 2000. In 2004 and 2005, a U.S. Geological Survey and U.S. Fish and Wildlife Service study was undertaken to determine if the threatened Santa Ana sucker was potentially exposed to OWCs and endocrine disrupting compounds (EDCs) in the SAR by using the western mosquitofish (Gambusia affinis) as a surrogate fish model. Four Santa Ana River sites were chosen along a gradient of proximity to WWTP effluents: (1) a point source of tertiary treated wastewater effluent (TTWE), (2) Rialto Drain (just below a WWTP), (3) Prado Dam (11 kilometers [km] below WWTPs), and (4) Sunnyslope Creek (no WWTP but having urban runoff influence). A reference site having no WWTPs or urban runoff, Thousand Palms, was also sampled. Chemical analyses of passive sampler extracts results showed that 15 OWCs and EDCs were detected in water from the Santa Ana River sites. Many of these compounds contributed to activity from an estrogenic in-vitro assay that showed a significant potential for impacting endocrine and reproductive systems compared to the 25 organochlorine compounds detected in aquatic biota. The site showing compounds having highest influence on sex steroid hormone activities was the point source for TTWE. Sex steroid hormone levels, secondary sex characteristics, organosomatic indices, and sperm quality parameters indicated impairment of endocrine and reproductive function of male western mosquitofish in the Santa Ana River. Exposure to EDCs and consequent impairment in mosquitofish followed the gradient of proximity to WWTP effluents, where the most significant effects were found at TTWE point source and Rialto Drain, followed by Prado Dam and Sunnyslope Creek. Each of these sites is suitable habitat for the Santa Ana sucker, especially Sunnyslope Creek and Rialto Drain where juveniles reside. Various OWCs and EDCs were detected at each Santa Ana River site, although one specific compound or group of compounds could not be singled out as a causative factor. Di (2-ethylhexyl) phthalate was strongly negatively correlated with testosterone in male mosquitofish. One group of potent environmental estrogens that likely contributed to endocrine and reproductive impairment are the natural and synthetic estrogen hormones, especially ethinyl estradiol; however, this compound was not targeted in these investigations. The multiple lines of evidence for impaired reproductive and endocrine function in western mosquitofish due to OWCs and EDCs from the Santa Ana River can be used to identify potential problems for the Santa Ana sucker inhabiting the same and nearby sites.

  4. An Ada/SQL (Structured Query Language) Application Scanner.

    DTIC Science & Technology

    1988-03-01

    Digital ...8217 (" DIGITS "), 46 new STRING’ ("DO"), new STRING’ ("ELSE"), new STRING’ ("ELSIF"), new STRING’ ("END"), new STRING’ ("ENTRY"), new STRING’ ("EXCEPTION...INTEGERPRINT; generic type NUM is digits <>; package FLOATPRINT is package txtprts.ada 18 prcdr PR (FL inFL %YE LINE n LINTYPE UNCLASSIFIED procedure

  5. Pitch glide effect induced by a nonlinear string-barrier interaction

    NASA Astrophysics Data System (ADS)

    Kartofelev, Dmitri; Stulov, Anatoli; Välimäki, Vesa

    2015-10-01

    Interactions of a vibrating string with its supports and other spatially distributed barriers play a significant role in the physics of many stringed musical instruments. It is well known that the tone of the string vibrations is determined by the string supports, and that the boundary conditions of the string termination may cause a short-lasting initial fundamental frequency shifting. Generally, this phenomenon is associated with the nonlinear modulation of the stiff string tension. The aim of this paper is to study the initial frequency glide phenomenon that is induced only by the string-barrier interaction, apart from other possible physical causes, and without the interfering effects of dissipation and dispersion. From a numerical simulation perspective, this highly nonlinear problem may present various difficulties, not the least of which is the risk of numerical instability. We propose a numerically stable and a purely kinematic model of the string-barrier interaction, which is based on the travelling wave solution of the ideal string vibration. The model is capable of reproducing the motion of the vibrating string exhibiting the initial fundamental frequency glide, which is caused solely by the complex nonlinear interaction of the string with its termination. The results presented in this paper can expand our knowledge and understanding of the timbre evolution and the physical principles of sound generation of numerous stringed instruments, such as lutes called the tambura, sitar and biwa.

  6. Constraint Reasoning Over Strings

    NASA Technical Reports Server (NTRS)

    Koga, Dennis (Technical Monitor); Golden, Keith; Pang, Wanlin

    2003-01-01

    This paper discusses an approach to representing and reasoning about constraints over strings. We discuss how many string domains can often be concisely represented using regular languages, and how constraints over strings, and domain operations on sets of strings, can be carried out using this representation.

  7. Materials science: How to suck like an octopus

    NASA Astrophysics Data System (ADS)

    Wilker, Jonathan J.

    2017-06-01

    Rubber sheets that reversibly bind and release substrates have been made by copying a subtlety in the shape of octopus suckers. The findings reveal how macro-scale biological structures can influence function. See Letter p.396

  8. Detection of DNA methylation changes in micropropagated banana plants using methylation-sensitive amplification polymorphism (MSAP).

    PubMed

    Peraza-Echeverria, S; Herrera-Valencia, V A.; Kay, A -J.

    2001-07-01

    The extent of DNA methylation polymorphisms was evaluated in micropropagated banana (Musa AAA cv. 'Grand Naine') derived from either the vegetative apex of the sucker or the floral apex of the male inflorescence using the methylation-sensitive amplification polymorphism (MSAP) technique. In all, 465 fragments, each representing a recognition site cleaved by either or both of the isoschizomers were amplified using eight combinations of primers. A total of 107 sites (23%) were found to be methylated at cytosine in the genome of micropropagated banana plants. In plants micropropagated from the male inflorescence explant 14 (3%) DNA methylation events were polymorphic, while plants micropropagated from the sucker explant produced 8 (1.7%) polymorphisms. No DNA methylation polymorphisms were detected in conventionally propagated banana plants. These results demonstrated the usefulness of MSAP to detect DNA methylation events in micropropagated banana plants and indicate that DNA methylation polymorphisms are associated with micropropagation.

  9. Revision and New Species of Vasotrema Stunkard, 1926 (Digenea: Schistosomatoidea): Turtle Blood Flukes of North American Softshell Turtles (Testudines: Trionychidae: Apalone spp.).

    PubMed

    Roberts, Jackson R; Bullard, Stephen A

    2017-10-01

    Gulf Coast spiny softshell turtles, Apalone spinifera aspera (Agassiz, 1857) (Testudines: Trionychidae) from Canoe Lake (33°47'56.16″N, 86°29'25.02″W; Springville, Alabama) and Round Lake (32°41'50.91″N, 87°14'30.39″W; Perry Lakes State Park, Marion, Alabama), were infected by V. robustum Stunkard, 1928 , Vasotrema longitestis Byrd, 1939 , and Vasotrema rileyae n. sp. The new species differs from its congeners by having papillate suckers, a short testis, an ovary dextral to the oviduct, and a pre-ovarian genital pore that is lateral to the ventral sucker. We studied the newly collected specimens and museum specimens of all congeners to revise the diagnosis of Vasotrema Stunkard, 1926 and redescribe and provide an updated dichotomous key to all species of the genus.

  10. Competition for food between an introduced crayfish and two fishes endemic to the Colorado River basin

    USGS Publications Warehouse

    Carpenter, J.

    2005-01-01

    Crayfish are not native to the Colorado River basin (CRB), however they are now established in portions of the mainstem and in many tributaries. I used density manipulation experiments in a laboratory setting to determine intra- and interspecific competition for food between Orconectes virilis, an aggressive polytrophic crayfish now common in the CRB, and two native fishes: Gila chub, Gila intermedia, and flannelmouth sucker, Catostomus latipinnis. I tested each fish species in separate trials. Growth of Gila chub decreased when animal densities increased, however they were more affected by intraspecific competition than by crayfish presence. In contrast, growth of flannelmouth suckers was more affected by crayfish than by intraspecific competition. Crayfish growth was not significantly altered by presence of either fish. Crayfish thus reduced fish growth by competition for food, but the effect differed markedly between the two species.

  11. Redescription of Synthesium pontoporiae n. comb. with notes on S. tursionis and S. seymouri n. comb. (Digenea: Brachycladiidae Odhner, 1905).

    PubMed

    Marigo, Juliana; Vicente, Ana Carolina Paulo; Valente, Ana Luisa Schifino; Measures, Lena; Santos, Cláudia Portes

    2008-04-01

    Synthesium pontoporiae n. comb. is redescribed, together with Synthesium tursionis and Synthesium seymouri n. comb.; the parasites were obtained from stranded and accidentally caught cetaceans. The sucker ratio (ratio between widths of the oral and ventral suckers) in S. pontoporiae was 1:1.8-3.0 (mean 1:2.2); in S. tursionis was 1:0.8-1.2; and in S. seymouri was 1:0.5-0.7. Synthesium pontoporiae differed from its congeners by additional diagnostic characters, including: oval to lobed testes; small cirrus with pyriform proximal region and flexible, tubular distal region formed by evagination of ejaculatory duct; and vitellarium in small follicles extending from the level of the seminal vesicle to the posterior extremity of the body and not forming dendritic radial bunches. Data on the morphology of adult S. pontoporiae and S. tursionis were inferred from confocal laser microscopical observations.

  12. The physics of unwound and wound strings on the electric guitar applied to the pitch intervals produced by tremolo/vibrato arm systems.

    PubMed

    Kemp, Jonathan A

    2017-01-01

    The physics of wound and unwound strings on the electric guitar are presented here, and the pitch intervals produced by the movements of a Fender Stratocaster tremolo unit are explained. Predicted changes in pitch sensitivity of different strings are given, and experimentally verified, for changes in saddle height, the distance of string free to move behind the nut and ratio of diameters/masses of the core and windings of wound strings. Also, it is shown that changes to the gauge of strings (assuming the string tension is sufficient for linear behaviour and in absence of changes to other construction details) don't alter the pitch intervals produced by a given angle of tremolo arm use assuming the instrument is set up with the same sounding pitches and starting bridge angle. It is demonstrated that it not possible to equalise the relative sensitivity of unwound steel stings on a Fender Stratocaster type tremolo unit through string construction techniques. The ratio of core to winding mass in the string, on the other hand, was found to be a very powerful design parameter for choosing the sensitivity of the string to tremolo arm use and standard pitch bends. For instance, the pitch intervals produced by operation of tremolo arm for wound strings may be made to approximately match that for one of the unwound strings if they share very similar core gauges (assuming the winding masses are chosen to give approximately the same tension at their sounding pitches). Such a design, only available currently by custom order, also delivers the optimum equalisation in sensitivity of strings for standard string bends (due to these also being produced by altering the length of the string to generate changes in tension and therefore pitch).

  13. Entanglement branes in a two-dimensional string theory

    DOE PAGES

    Donnelly, William; Wong, Gabriel

    2017-09-20

    What is the meaning of entanglement in a theory of extended objects such as strings? To address this question we consider the spatial entanglement between two intervals in the Gross-Taylor model, the string theory dual to two-dimensional Yang-Mills theory at large N. The string diagrams that contribute to the entanglement entropy describe open strings with endpoints anchored to the entangling surface, as first argued by Susskind. We develop a canonical theory of these open strings, and describe how closed strings are divided into open strings at the level of the Hilbert space. Here, we derive the modular Hamiltonian for themore » Hartle-Hawking state and show that the corresponding reduced density matrix describes a thermal ensemble of open strings ending on an object at the entangling surface that we call an entanglement brane, or E-brane.« less

  14. String-driven inflation

    NASA Technical Reports Server (NTRS)

    Turok, Neil

    1988-01-01

    It is argued that, in fundamental string theories, as one traces the universe back in time a point is reached when the expansion rate is so fast that the rate of string creation due to quantum effects balances the dilution of the string density due to the expansion. One is therefore led into a phase of constant string density and an exponentially expanding universe. Fundamental strings therefore seem to lead naturally to inflation.

  15. p-adic string theories provide lattice Discretization to the ordinary string worldsheet.

    PubMed

    Ghoshal, Debashis

    2006-10-13

    A class of models called p-adic strings is useful in understanding the tachyonic instability of string theory. These are found to be empirically related to the ordinary strings in the p-->1 limit. We propose that these models provide discretization for the string worldsheet and argue that the limit is naturally thought of as a continuum limit in the sense of the renormalization group.

  16. Self-organization in a system of binary strings with spatial interactions

    NASA Astrophysics Data System (ADS)

    Banzhaf, W.; Dittrich, P.; Eller, B.

    1999-01-01

    We consider an artificial reaction system whose components are binary strings. Upon encounter, two binary strings produce a third string which competes for storage space with the originators. String types or species can only survive when produced in sufficient numbers. Spatial interactions through introduction of a topology and rules for distance-dependent reactions are discussed. We observe various kinds of survival strategies of binary strings.

  17. p-adic String Theories Provide Lattice Discretization to the Ordinary String Worldsheet

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Ghoshal, Debashis

    2006-10-13

    A class of models called p-adic strings is useful in understanding the tachyonic instability of string theory. These are found to be empirically related to the ordinary strings in the p{yields}1 limit. We propose that these models provide discretization for the string worldsheet and argue that the limit is naturally thought of as a continuum limit in the sense of the renormalization group.

  18. Progress report for a research program in theoretical high energy physics

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Feldman, D.; Fried, H.M.; Jevicki, A.

    This year's research has dealt with: superstrings in the early universe; the invisible axion emissions from SN1987A; quartic interaction in Witten's superstring field theory; W-boson associated multiplicity and the dual parton model; cosmic strings and galaxy formation; cosmic strings and baryogenesis; quark flavor mixing; p -- /bar p/ scattering at TeV energies; random surfaces; ordered exponentials and differential equations; initial value and back-reaction problems in quantum field theory; string field theory and Weyl invariance; the renormalization group and string field theory; the evolution of scalar fields in an inflationary universe, with and without the effects of gravitational perturbations; cosmic stringmore » catalysis of skyrmion decay; inflation and cosmic strings from dynamical symmetry breaking; the physic of flavor mixing; string-inspired cosmology; strings at high-energy densities and complex temperatures; the problem of non-locality in string theory; string statistical mechanics; large-scale structures with cosmic strings and neutrinos; the delta expansion for stochastic quantization; high-energy neutrino flux from ordinary cosmic strings; a physical picture of loop bremsstrahlung; cylindrically-symmetric solutions of four-dimensional sigma models; large-scale structure with hot dark matter and cosmic strings; the unitarization of the odderon; string thermodynamics and conservation laws; the dependence of inflationary-universe models on initial conditions; the delta expansion and local gauge invariance; particle physics and galaxy formation; chaotic inflation with metric and matter perturbations; grand-unified theories, galaxy formation, and large-scale structure; neutrino clustering in cosmic-string-induced wakes; and infrared approximations to nonlinear differential equations. 17 refs.« less

  19. An investigation of enhanced mercury bioaccumulation in fish from offshore feeding.

    PubMed

    Chételat, John; Cloutier, Louise; Amyot, Marc

    2013-08-01

    We investigated the dietary pathways of mercury transfer in the food web of Morency Lake (Canada) to determine the influence of carbon source and habitat use on mercury bioaccumulation in fish. Whole-body concentrations of methylmercury (MeHg) were significantly different in four fish species (white sucker, brown bullhead, pumpkinseed and smallmouth bass) and increased with both trophic position and greater feeding on offshore (versus littoral) carbon. An examination of fish gut contents and the depth distribution of invertebrates in Morency Lake showed that smallmouth bass and brown bullhead were supplementing their littoral diet with the consumption of either opossum shrimp (Mysis diluviana) or profundal amphipods in offshore waters. The zooplanktivore Mysis had significantly higher MeHg concentrations than zooplankton and benthic invertebrates, and it was an elevated source of MeHg to smallmouth bass. In contrast, profundal amphipods consumed by brown bullhead did not have higher MeHg concentrations than littoral amphipods. Instead, partitioning of benthic invertebrate resources likely explains the greater MeHg bioaccumulation in brown bullhead, associated with offshore feeding of amphipods. White sucker and brown bullhead had a similar trophic position but white sucker consumed more chironomids, which had one-third the MeHg concentration of amphipods. Our findings suggest that offshore feeding in a lake can affect fish MeHg bioaccumulation via two different processes: (1) the consumption of MeHg-enriched pelagic prey, or (2) resource partitioning of benthic primary consumers with different MeHg concentrations. These observations on the mechanisms of habitat-specific bioaccumulation highlight the complexity of MeHg transfer through lake food webs.

  20. Artyfechinostomum malayanum: Metacercariae Encysted in Pila sp. Snails Purchased from Phnom Penh, Cambodia

    PubMed Central

    Sohn, Woon-Mok; Yong, Tai-Soon; Eom, Keeseon S.; Sinuon, Muth; Jeoung, Hoo-Gn; Chai, Jong-Yil

    2017-01-01

    The metacercariae of Artyfechinostomum malayanum (Leiper, 1911) Mendheim, 1943 were discovered in Pila sp. snails purchased from a market in Phnom Penh, Cambodia. They were isolated from the snails using the artificial digestion technique and were orally fed to 2 hamsters, 1 rat, and 2 mice to obtain the adult flukes. The metacercariae were round, 145–165 μm in diameter, having a cyst wall of 6–10 μm in thickness, a head collar and collar spines, and characteristic features of excretory granules. Adult flukes were recovered in the small intestines of the animals at days 14 and 32 post infection and were morphologically observed using a light microscope and a scanning electron microscope. They were plump or elongated, ventrally curved, 6.0–8.1×1.6–2.0 mm in size, and characterized by the head collar bearing 43 collar spines, including 5 end group ones on each side, a long cirrus sac extending beyond the posterior margin of the ventral sucker, a submedian ovary, and 2 deeply lobed testes. Eggs in uteri were operculate, ovoid to ellipsoid, and 120–135×68–75 μm in size. In scanning electron microscopy, the head collar was prominent with collar spines looking like horns. Scale-like tegumental spines were densely distributed on the ventral surface between the head collar and ventral sucker. Sensory papillae were distributed mainly on the tegument around suckers. By this study, it has been first confirmed that the life cycle of A. malayanum exists in Cambodia. PMID:28719961

  1. Gonadotropin-Releasing hormones in the brain and pituitary of the white sucker

    USGS Publications Warehouse

    Robinson, T. Craig; Tobet, Stuart A.; Chase, Cindy; Waldron, Travis; Sower, Stacia A.

    2000-01-01

    The present study investigated GnRH forms within the brain of a representative of the order Cypriniformes, the white sucker, Catostomus commersoni, using HPLC, RIA, andimmunocytochemistry. Several immunoreactive (ir) GnRH forms were identified in the brain of the white sucker by chromatography and radioimmunoassay, including ir-salmon GnRH, ir-lamprey GnRH-I and -III, and ir-chicken GnRH-II. Results from immunocytochemical studies were consistent with multiple GnRH forms distributed in different patterns, particularly for fibers. Neuronal perikarya containing ir-salmon GnRH and ir-lamprey-like GnRH were found laterally within the preoptic area and rostralhypothalamus. Cells containing exclusively ir-salmon GnRH appeared slightly more rostrally, but in the same region. Fibers containing ir-salmon GnRH and ir-lamprey-like GnRH were seen throughout the caudal telencephalon and extended into thediencephalon, toward the pituitary. Fibers containing ir-chicken-II-like GnRH were also seen in the caudal telencephalon, but were concentrated more dorsally in the diencephalon. Within the pituitary, fibers containing ir-salmon GnRH and ir-lamprey-like GnRH entered the neurohypophysis, but differed in their destinations. Fibers containing ir-salmon GnRH remained within the neurohypophysis, while fibers containing ir-lamprey-like GnRH targeted adenohypophyseal tissue. These findings are consistent with the hypothesis that multiple GnRH forms with multiple functions exist within the brain and pituitary of teleosts and provide further evidence of a lamprey-like GnRH within an early evolved teleost species.

  2. Effects of summer drawdown on the fishes and larval chironomids in Beulah Reservoir, Oregon

    USGS Publications Warehouse

    Rose, Brien P.; Mesa, Matthew G.

    2013-01-01

    Summer drawdown of Beulah Reservoir, Oregon, could adversely affect fish and invertebrate production, limit sport fishing opportunities, and hinder the recovery of threatened species. To assess the impacts of drawdown, we sampled fish and Chironomidae larvae in Beulah Reservoir in the springs of 2006 to 2008. The reservoir was reduced to 68% of full pool in 2006 and to run-of-river level in 2007. From spring 2006 to spring 2007, the catch per unit effort (CPUE) of fyke nets decreased significantly for dace [Rhinichthys spp.] and northern pikeminnow [Ptychocheilus oregonensis], increased significantly for suckers [Catastomus spp.] and white crappies [Pomoxis nigromaculatus], and was similar for redside shiners [Richardsonius balteatus]. CPUE of gillnets either increased significantly or remained similar depending on genera, and the size structure of redside shiners, suckers, and white crappies changed appreciably. From 2007 to 2008, the CPUE of northern pikeminnow, redside shiners, suckers, and white crappies decreased significantly depending on gear and the size structure of most fishes changed. Springtime densities of chironomid larvae in the water column were significantly higher in 2006 than in 2008, but other comparisons were similar. The densities of benthic chironomids were significantly lower in substrates that were frequently dewatered compared to areas that were partially or usually not dewatered. Individuals from frequently dewatered areas were significantly smaller than those from other areas and the densities of benthic chironomids in 2008 were significantly lower than other years. Summer drawdown can reduce the catch and alter the size structure of fishes and chironomid larvae in Beulah Reservoir.

  3. Identifying Preservation and Restoration Priority Areas for Desert Fishes in an Increasingly Invaded World

    NASA Astrophysics Data System (ADS)

    Pool, Thomas K.; Strecker, Angela L.; Olden, Julian D.

    2013-03-01

    A commonly overlooked aspect of conservation planning assessments is that wildlife managers are increasingly focused on habitats that contain non-native species. We examine this management challenge in the Gila River basin (150,730 km2), and present a new planning strategy for fish conservation. By applying a hierarchical prioritization algorithm to >850,000 fish records in 27,181 sub-watersheds we first identified high priority areas (PAs) termed "preservation PAs" with high native fish richness and low non-native richness; these represent traditional conservation targets. Second, we identified "restoration PAs" with high native fish richness that also contained high numbers of non-native species; these represent less traditional conservation targets. The top 10 % of preservation and restoration PAs contained common native species (e.g., Catostomus clarkii, desert sucker; Catostomus insignis, Sonora sucker) in addition to native species with limited distributions (i.e., Xyrauchen texanus, razorback sucker; Oncorhynchus gilae apache, Apache trout). The top preservation and restoration PAs overlapped by 42 %, indicating areas with high native fish richness range from minimally to highly invaded. Areas exclusively identified as restoration PAs also encompassed a greater percentage of native species ranges than would be expected by the random addition of an equivalent basin area. Restoration PAs identified an additional 19.0 and 26.6 % of the total ranges of two federally endangered species— Meda fulgida (spikedace) and Gila intermedia (Gila chub), respectively, compared to top preservation PAs alone—despite adding only 5.8 % of basin area. We contend that in addition to preservation PAs, restoration PAs are well suited for complementary management activities benefiting native fishes.

  4. Demographics and run timing of adult Lost River (Deltistes luxatus) and short nose (Chasmistes brevirostris) suckers in Upper Klamath Lake, Oregon, 2009

    USGS Publications Warehouse

    Hewitt, David A.; Hayes, Brian S.; Janney, Eric C.; Harris, Alta C.; Koller, Justin P.; Johnson, Mark A.

    2011-01-01

    Data from a long-term capture-recapture program were used to assess the status and dynamics of populations of two long-lived, federally endangered catostomids in Upper Klamath Lake, Oregon. Lost River suckers (Deltistes luxatus) and shortnose suckers (Chasmistes brevirostris) have been captured and tagged with passive integrated transponder (PIT) tags during their spawning migrations in each year since 1995. In addition, beginning in 2005, individuals that had been previously PIT-tagged were reencountered on remote underwater antennas deployed throughout the spawning areas. Captures and remote encounters during spring 2009 were used to describe the spawning migrations in that year and also were incorporated into capture-recapture analyses of population dynamics over the last decade. Cormack-Jolly-Seber (CJS) open population capture-recapture models were used to estimate annual survival probabilities, and a reverse-time analog of the CJS model was used to estimate recruitment of new individuals into the spawning populations. In addition, data on the size composition of captured fish was examined for any additional evidence of recruitment. Survival and recruitment estimates were combined to estimate changes in population size over time and to determine the status of the populations through 2007. Separate analyses were conducted for each species and also for each subpopulation of Lost River suckers (LRS). One subpopulation of LRS migrates into tributaries to spawn, similar to shortnose suckers (SNS), whereas the other subpopulation spawns at upwelling areas along the eastern shoreline of the lake. In 2009, we captured and tagged 781 LRS at four shoreline areas and recaptured an additional 638 individuals that had been tagged in previous years. Across all four areas, the remote antennas detected 6,056 individual LRS during the spawning season. Spawning activity peaked in April and most individuals were encountered at Sucker Springs and Cinder Flats. In the Williamson River, we captured and tagged 3,008 LRS and 287 SNS, and recaptured 271 LRS and 81 SNS that had been tagged in previous years. Remote antennas that spanned the river downstream of the tributary spawning areas detected a total of 12,509 LRS and 5,023 SNS. Most LRS passed upstream in mid-April when water temperatures were rising and near or greater than 10°C. In contrast, peaks in upstream passage of SNS occurred in late April and early May when water temperatures were rising and near or greater than 12°C. Finally, an additional 1,569 LRS and 1,794 SNS were captured in trammel net sampling at prespawn staging areas in the northeastern portion of the lake. Of these, 209 of the LRS and 452 of the SNS had been PIT-tagged in previous years. For LRS, encounter histories showed that nearly all of the fish captured at the staging areas were members of the subpopulation that spawns in the tributaries.Capture-recapture analyses for the LRS subpopulation that spawns at the shoreline areas included encounter histories for more than 9,000 individuals, and analyses for the subpopulation that spawns in the tributaries included more than 14,000 encounter histories. With a few exceptions, the survival of males and females in both subpopulations was high (>0.9) between 1999 and 2007. Notably lower survival occurred for both sexes from the tributaries in 2000, for males from the shoreline areas in 2002, and for males from the tributaries in 2006. Recruitment of new individuals into either spawning population was trivial in all years between 2002 and 2007. Over that period, the abundance of males in the lakeshore spawning subpopulation declined by 44–53 percent and the abundance of females declined by 25–38 percent. Similarly, the abundance of males in the tributary spawning subpopulation declined by as much as 39 percent and the abundance of females declined by as much as 33 percent. Capture-recapture analyses for SNS included encounter histories for more than 12,000 individuals. The majority of annual survival estimates between 2001 and 2007 were high (>0.8), but SNS experienced more years of low survival than either LRS subpopulation. The survival of both sexes was particularly low in both 2001 and 2004, and male survival also was somewhat low in 2002 and 2006. Similar to LRS, recruitment of new individuals into the spawning population was trivial in all years between 2001 and 2007. Over that period, the abundance of male SNS declined by 58–80 percent and the abundance of females declined by 52–73 percent. Despite relatively high survival in most years, both species have experienced substantial declines in the abundance of spawning fish because losses from mortality have not been balanced by recruitment of new individuals. Indeed, all populations appear to be largely comprised of fish that were present in the late 1990s and early 2000s. As a result, the status of the endangered sucker populations in Upper Klamath Lake remains worrisome, and the situation is most dire for shortnose suckers. Survival analyses show that the two species do not necessarily experience poor survival in the same years and that poor survival on an annual scale is not predictable from fish die-offs observed in the summer and fall. Future analyses will explore the connections between annual sucker survival and environmental factors of interest, such as water quality and disease. Our monitoring program provides a robust platform for estimating vital population parameters, evaluating the status of the populations, and assessing the effectiveness of conservation and recovery efforts.

  5. String Formatting Considered Harmful for Novice Programmers

    ERIC Educational Resources Information Center

    Hughes, Michael C.; Jadud, Matthew C.; Rodrigo, Ma. Mercedes T.

    2010-01-01

    In Java, "System.out.printf" and "String.format" consume a specialised kind of string commonly known as a format string. In our study of first-year students at the Ateneo de Manila University, we discovered that format strings present a substantial challenge for novice programmers. Focusing on their first laboratory we found…

  6. Charting the landscape of supercritical string theory.

    PubMed

    Hellerman, Simeon; Swanson, Ian

    2007-10-26

    Special solutions of string theory in supercritical dimensions can interpolate in time between theories with different numbers of spacetime dimensions and different amounts of world sheet supersymmetry. These solutions connect supercritical string theories to the more familiar string duality web in ten dimensions and provide a precise link between supersymmetric and purely bosonic string theories. Dimension quenching and c duality appear to be natural concepts in string theory, giving rise to large networks of interconnected theories.

  7. String solutions in spherically-symmetric f(R) gravity vacuum

    NASA Astrophysics Data System (ADS)

    Dil, Emre

    Dynamical evolution of the cosmic string in a spherically symmetric f(R) gravity vacuum is studied for a closed and straight string. We first set the background spacetime metric for a constant curvature scalar R = R0, and obtain the Killing fields for it. Using the standard gauge coordinates and constraints for both closed and straight strings, we present the equation of motions and find the solutions of them. We then analyze the dynamics of the string by studying the behavior of the string radius and periastron radius, with respect to both proper time and azimuthal angle, for various values of f(R) functions. Consequently, we conclude that the value of f(R) dramatically affects the closed string collapse time and the straight string angular deviation.

  8. Diffusion of massive particles around an Abelian-Higgs string

    NASA Astrophysics Data System (ADS)

    Saha, Abhisek; Sanyal, Soma

    2018-03-01

    We study the diffusion of massive particles in the space time of an Abelian Higgs string. The particles in the early universe plasma execute Brownian motion. This motion of the particles is modeled as a two dimensional random walk in the plane of the Abelian Higgs string. The particles move randomly in the space time of the string according to their geodesic equations. We observe that for certain values of their energy and angular momentum, an overdensity of particles is observed close to the string. We find that the string parameters determine the distribution of the particles. We make an estimate of the density fluctuation generated around the string as a function of the deficit angle. Though the thickness of the string is small, the length is large and the overdensity close to the string may have cosmological consequences in the early universe.

  9. Remarks on entanglement entropy in string theory

    NASA Astrophysics Data System (ADS)

    Balasubramanian, Vijay; Parrikar, Onkar

    2018-03-01

    Entanglement entropy for spatial subregions is difficult to define in string theory because of the extended nature of strings. Here we propose a definition for bosonic open strings using the framework of string field theory. The key difference (compared to ordinary quantum field theory) is that the subregion is chosen inside a Cauchy surface in the "space of open string configurations." We first present a simple calculation of this entanglement entropy in free light-cone string field theory, ignoring subtleties related to the factorization of the Hilbert space. We reproduce the answer expected from an effective field theory point of view, namely a sum over the one-loop entanglement entropies corresponding to all the particle-excitations of the string, and further show that the full string theory regulates ultraviolet divergences in the entanglement entropy. We then revisit the question of factorization of the Hilbert space by analyzing the covariant phase-space associated with a subregion in Witten's covariant string field theory. We show that the pure gauge (i.e., BRST exact) modes in the string field become dynamical at the entanglement cut. Thus, a proper definition of the entropy must involve an extended Hilbert space, with new stringy edge modes localized at the entanglement cut.

  10. Using glyphosate herbicide in converting aspen to conifers.

    Treesearch

    Donald A. Perala

    1985-01-01

    Glyphosate at 1.5 to 2.0 lbs. Per acre active ingredient will control aspen suckers, shrubs, and herbs when applied during August to early September. There appears to be some intersite variability in the efficacy of the herbicide.

  11. Cosmic string catalysis of skyrmion decay

    NASA Technical Reports Server (NTRS)

    Gregory, Ruth; Davis, Anne-Christine; Brandenberger, Robert

    1988-01-01

    The Callan-Witten picture is developed for monopole catalyzed skyrmion decay in order to analyze the corresponding cosmic string scenario. It is discovered that cosmic strings (both ordinary and superconducting) can catalyze proton decay, but that this catalysis only occurs on the scale of the core of the string. In order to do this we have to develop a vortex model for the superconducting string. An argument is also given for the difference in the enhancement factors for monopoles and strings.

  12. Hydraulics Graphics Package. Users Manual

    DTIC Science & Technology

    1985-11-01

    ENTER: VARIABLE/SEPARATOR/VALUE OR STRING GLBL, TETON DAM FAILURE ENTER: VARIABLE/SEPARATOR/VALUE OR STRING SLOC ,DISCHARGE HISTOGRAM ENTER: VARIABLE...ENTER: VARIABLE/SEPARATOR/VALUE OR STRING YLBL,FLOW IN 1000 CFS ENTER: VARIABLE/SEPARATORVA LUE OR STRING GLBL, TETON DAM FAILURE ENTER: VARIABLE...SEPARATOR/VALUE OR STRING SECNO, 0 ENTER: VARIABLE/SEPARATOR/VALUE OR STRING GO 1ee0. F go L 0 U I Goo. 200. TETON DAM FAILUPE N\\ rLOIJ Alr 4wi. fiNT. I .I

  13. Impaired letter-string processing in developmental dyslexia: what visual-to-phonology code mapping disorder?

    PubMed

    Valdois, Sylviane; Lassus-Sangosse, Delphine; Lobier, Muriel

    2012-05-01

    Poor parallel letter-string processing in developmental dyslexia was taken as evidence of poor visual attention (VA) span, that is, a limitation of visual attentional resources that affects multi-character processing. However, the use of letter stimuli in oral report tasks was challenged on its capacity to highlight a VA span disorder. In particular, report of poor letter/digit-string processing but preserved symbol-string processing was viewed as evidence of poor visual-to-phonology code mapping, in line with the phonological theory of developmental dyslexia. We assessed here the visual-to-phonological-code mapping disorder hypothesis. In Experiment 1, letter-string, digit-string and colour-string processing was assessed to disentangle a phonological versus visual familiarity account of the letter/digit versus symbol dissociation. Against a visual-to-phonological-code mapping disorder but in support of a familiarity account, results showed poor letter/digit-string processing but preserved colour-string processing in dyslexic children. In Experiment 2, two tasks of letter-string report were used, one of which was performed simultaneously to a high-taxing phonological task. Results show that dyslexic children are similarly impaired in letter-string report whether a concurrent phonological task is simultaneously performed or not. Taken together, these results provide strong evidence against a phonological account of poor letter-string processing in developmental dyslexia. Copyright © 2012 John Wiley & Sons, Ltd.

  14. Multiflavor string-net models

    NASA Astrophysics Data System (ADS)

    Lin, Chien-Hung

    2017-05-01

    We generalize the string-net construction to multiple flavors of strings, each of which is labeled by the elements of an Abelian group Gi. The same flavor of strings can branch, while different flavors of strings can cross one another and thus they form intersecting string nets. We systematically construct the exactly soluble lattice Hamiltonians and the ground-state wave functions for the intersecting string-net condensed phases. We analyze the braiding statistics of the low-energy quasiparticle excitations and find that our model can realize all the topological phases as the string-net model with group G =∏iGi . In this respect, our construction provides various ways of building lattice models which realize topological order G , corresponding to different partitions of G and thus different flavors of string nets. In fact, our construction concretely demonstrates the Künneth formula by constructing various lattice models with the same topological order. As an example, we construct the G =Z2×Z2×Z2 string-net model which realizes a non-Abelian topological phase by properly intersecting three copies of toric codes.

  15. Photovoltaic power generation system with photovoltaic cells as bypass diodes

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Lentine, Anthony L.; Nielson, Gregory N.; Tauke-Pedretti, Anna

    A photovoltaic power generation system that includes a solar panel is described herein. The solar panel includes a photovoltaic sub-module, which includes a group of microsystem enabled photovoltaic cells. The group includes a first string of photovoltaic cells, a second string of photovoltaic cells, and a differing photovoltaic cell. Photovoltaic cells in the first string are electrically connected in series, and photovoltaic cells in the second string are electrically connected in series. Further, the first string of photovoltaic cells, the second string of photovoltaic cells, and the differing photovoltaic cell are electrically connected in parallel. Moreover, the differing photovoltaic cellmore » is used as a bypass diode for the first string of photovoltaic cells and the second string of photovoltaic cells.« less

  16. DOE Office of Scientific and Technical Information (OSTI.GOV)

    Donnelly, William; Wong, Gabriel

    What is the meaning of entanglement in a theory of extended objects such as strings? To address this question we consider the spatial entanglement between two intervals in the Gross-Taylor model, the string theory dual to two-dimensional Yang-Mills theory at large N. The string diagrams that contribute to the entanglement entropy describe open strings with endpoints anchored to the entangling surface, as first argued by Susskind. We develop a canonical theory of these open strings, and describe how closed strings are divided into open strings at the level of the Hilbert space. Here, we derive the modular Hamiltonian for themore » Hartle-Hawking state and show that the corresponding reduced density matrix describes a thermal ensemble of open strings ending on an object at the entangling surface that we call an entanglement brane, or E-brane.« less

  17. Device for balancing parallel strings

    DOEpatents

    Mashikian, Matthew S.

    1985-01-01

    A battery plant is described which features magnetic circuit means in association with each of the battery strings in the battery plant for balancing the electrical current flow through the battery strings by equalizing the voltage across each of the battery strings. Each of the magnetic circuit means generally comprises means for sensing the electrical current flow through one of the battery strings, and a saturable reactor having a main winding connected electrically in series with the battery string, a bias winding connected to a source of alternating current and a control winding connected to a variable source of direct current controlled by the sensing means. Each of the battery strings is formed by a plurality of batteries connected electrically in series, and these battery strings are connected electrically in parallel across common bus conductors.

  18. Aspects of some dualities in string theory

    NASA Astrophysics Data System (ADS)

    Kim, Bom Soo

    AdS/CFT correspondence in string theory has changed landscape of the theoretical physics. Through this celebrated duality between gravity theory and field theory, one can investigate analytically strongly coupled gauge theories such as Quantum Chromodynamics (QCD) in terms of weakly coupled string theory such as supergravity theory and vice versa. In the first part of this thesis we used this duality to construct a new type of nonlocal field theory, called Puff Field Theory, in terms of D3 branes in type IIB string theory with a geometric twist. In addition to the strong-weak duality of AdS/CFT, there also exists a weak-weak duality, called Twistor String Theory. Twistor technique is successfully used to calculate the SYM scattering amplitude in an elegant fashion. Yet, the progress in the string theory side was hindered by a non-unitary conformal gravity. We extend the Twistor string theory by introducing mass terms, in the second part of the thesis. A chiral mass term is identified as a vacuum expectation value of a conformal supergravity field and is tied with the breaking of the conformal symmetry of gravity. As a prime candidate for a quantum theory of gravity, string theory revealed many promising successes such as counting the number of microstates in supersymmetric Black Holes thermodynamics and resolution of timelike and null singularities, to name a few. Yet, the fundamental string and M-theroy formulations are not yet available. Various string theories without gravity, such as Non-Commutative Open String (NCOS) and Open Membrane (OM) theories, are very nice playground to investigate the fundamental structure of string and M-theory without the complication of gravity. In the last part of the thesis, simpler Non-Relativistic String Theories are constructed and investigated. One important motivation for those theories is related to the connection between Non-Relativistic String Theories and Non-critical String Theories through the bosonization of betagamma CFT.

  19. The physics of unwound and wound strings on the electric guitar applied to the pitch intervals produced by tremolo/vibrato arm systems

    PubMed Central

    2017-01-01

    The physics of wound and unwound strings on the electric guitar are presented here, and the pitch intervals produced by the movements of a Fender Stratocaster tremolo unit are explained. Predicted changes in pitch sensitivity of different strings are given, and experimentally verified, for changes in saddle height, the distance of string free to move behind the nut and ratio of diameters/masses of the core and windings of wound strings. Also, it is shown that changes to the gauge of strings (assuming the string tension is sufficient for linear behaviour and in absence of changes to other construction details) don’t alter the pitch intervals produced by a given angle of tremolo arm use assuming the instrument is set up with the same sounding pitches and starting bridge angle. It is demonstrated that it not possible to equalise the relative sensitivity of unwound steel stings on a Fender Stratocaster type tremolo unit through string construction techniques. The ratio of core to winding mass in the string, on the other hand, was found to be a very powerful design parameter for choosing the sensitivity of the string to tremolo arm use and standard pitch bends. For instance, the pitch intervals produced by operation of tremolo arm for wound strings may be made to approximately match that for one of the unwound strings if they share very similar core gauges (assuming the winding masses are chosen to give approximately the same tension at their sounding pitches). Such a design, only available currently by custom order, also delivers the optimum equalisation in sensitivity of strings for standard string bends (due to these also being produced by altering the length of the string to generate changes in tension and therefore pitch). PMID:28934268

  20. Formation of Electron Strings in Narrow Band Polar Semiconductors

    NASA Astrophysics Data System (ADS)

    Kusmartsev, F. V.

    2000-01-01

    We show that linear electron strings may arise in polar semiconductors. A single string consists of M spinless fermions trapped by an extended polarization well of a cigar shape. Inside the string the particles are free although they interact with each other via Coulomb forces. The strings arise as a result of an electronic phase separation associated with an instability of small adiabatic polarons. We have found the length of the string which depends on dielectric constants of semiconductors. The appearance of these electron strings may have an impact on the effect of stripe formation observed in a variety of high- Tc experiments.

  1. Optimal management of batteries in electric systems

    DOEpatents

    Atcitty, Stanley; Butler, Paul C.; Corey, Garth P.; Symons, Philip C.

    2002-01-01

    An electric system including at least a pair of battery strings and an AC source minimizes the use and maximizes the efficiency of the AC source by using the AC source only to charge all battery strings at the same time. Then one or more battery strings is used to power the load while management, such as application of a finish charge, is provided to one battery string. After another charge cycle, the roles of the battery strings are reversed so that each battery string receives regular management.

  2. The waiting time problem in a model hominin population.

    PubMed

    Sanford, John; Brewer, Wesley; Smith, Franzine; Baumgardner, John

    2015-09-17

    Functional information is normally communicated using specific, context-dependent strings of symbolic characters. This is true within the human realm (texts and computer programs), and also within the biological realm (nucleic acids and proteins). In biology, strings of nucleotides encode much of the information within living cells. How do such information-bearing nucleotide strings arise and become established? This paper uses comprehensive numerical simulation to understand what types of nucleotide strings can realistically be established via the mutation/selection process, given a reasonable timeframe. The program Mendel's Accountant realistically simulates the mutation/selection process, and was modified so that a starting string of nucleotides could be specified, and a corresponding target string of nucleotides could be specified. We simulated a classic pre-human hominin population of at least 10,000 individuals, with a generation time of 20 years, and with very strong selection (50% selective elimination). Random point mutations were generated within the starting string. Whenever an instance of the target string arose, all individuals carrying the target string were assigned a specified reproductive advantage. When natural selection had successfully amplified an instance of the target string to the point of fixation, the experiment was halted, and the waiting time statistics were tabulated. Using this methodology we tested the effect of mutation rate, string length, fitness benefit, and population size on waiting time to fixation. Biologically realistic numerical simulations revealed that a population of this type required inordinately long waiting times to establish even the shortest nucleotide strings. To establish a string of two nucleotides required on average 84 million years. To establish a string of five nucleotides required on average 2 billion years. We found that waiting times were reduced by higher mutation rates, stronger fitness benefits, and larger population sizes. However, even using the most generous feasible parameters settings, the waiting time required to establish any specific nucleotide string within this type of population was consistently prohibitive. We show that the waiting time problem is a significant constraint on the macroevolution of the classic hominin population. Routine establishment of specific beneficial strings of two or more nucleotides becomes very problematic.

  3. The IMS Software Integration Platform

    DTIC Science & Technology

    1993-04-12

    products to incorporate all data shared by the IMS applications. Some entities (time-series, images, a algorithm -specific parameters) must be managed...dbwhoanii, dbcancel Transaction Management: dbcommit, dbrollback Key Counter Assignment: dbgetcounter String Handling: cstr ~to~pad, pad-to- cstr Error...increment *value; String Maniputation: int cstr topad (array, string, arraylength) char *array, *string; int arrayjlength; int pad tocstr (string

  4. Cosmic superstrings: Observable remnants of brane inflation

    NASA Astrophysics Data System (ADS)

    Wyman, Mark Charles

    Brane inflation provides a natural dynamical model for the physics which underlie the inflationary paradigm. Besides their inflationary predictions, brane models imply another observable consequence: cosmic strings. In this dissertation I outline the background of how cosmic strings arise in brane inflationary models and how the properties of the strings and the models are mutually tied (Chapter 2). I then use cosmological observations to put limits on the properties of any actually-existing cosmic string network (Chapter 3). Next, I study the question of how cosmic superstrings, as the cosmic strings arising from string theory are known, could be distinct from classical gauge- theory cosmic strings. In particular, I propose an analytical model for the cosmological evolution of a network of binding cosmic strings (Chapter 4); I also describe the distinctive gravitational lensing phenomena that are caused by binding strings (Chapter 5). Finally, I lay out the background for the numerical study of a gauge theory model for the dynamics of cosmic superstring binding (Chapter 6).

  5. String theory--the physics of string-bending and other electric guitar techniques.

    PubMed

    Grimes, David Robert

    2014-01-01

    Electric guitar playing is ubiquitous in practically all modern music genres. In the hands of an experienced player, electric guitars can sound as expressive and distinct as a human voice. Unlike other more quantised instruments where pitch is a discrete function, guitarists can incorporate micro-tonality and, as a result, vibrato and sting-bending are idiosyncratic hallmarks of a player. Similarly, a wide variety of techniques unique to the electric guitar have emerged. While the mechano-acoustics of stringed instruments and vibrating strings are well studied, there has been comparatively little work dedicated to the underlying physics of unique electric guitar techniques and strings, nor the mechanical factors influencing vibrato, string-bending, fretting force and whammy-bar dynamics. In this work, models for these processes are derived and the implications for guitar and string design discussed. The string-bending model is experimentally validated using a variety of strings and vibrato dynamics are simulated. The implications of these findings on the configuration and design of guitars is also discussed.

  6. PhD Thesis: String theory in the early universe

    NASA Astrophysics Data System (ADS)

    Gwyn, Rhiannon

    2009-11-01

    The intersection of string theory with cosmology is unavoidable in the early universe, and its exploration may shine light on both fields. In this thesis, three papers at this intersection are presented and reviewed, with the aim of providing a thorough and pedagogical guide to their results. First, we address the longstanding problem of finding a string theory realisation of the axion. Using warped compactifications in heterotic string theory, we show that the axion decay constant can be lowered to acceptable values by the warp factor. Next, we move to the subject of cosmic strings, whose network evolution could have important consequences for astrophysics and cosmology. In particular, there are quantitative differences between cosmic superstring networks and GUT cosmic string networks. We investigate the properties of cosmic superstring networks in warped backgrounds, giving the tension and properties of three-string junctions in these backgrounds. Finally, we examine the possibility that cosmic strings in heterotic string theory could be responsible for generating the galactic magnetic fields that seeded those observed today.

  7. DOE Office of Scientific and Technical Information (OSTI.GOV)

    Ciuca, Razvan; Hernández, Oscar F., E-mail: razvan.ciuca@mail.mcgill.ca, E-mail: oscarh@physics.mcgill.ca

    There exists various proposals to detect cosmic strings from Cosmic Microwave Background (CMB) or 21 cm temperature maps. Current proposals do not aim to find the location of strings on sky maps, all of these approaches can be thought of as a statistic on a sky map. We propose a Bayesian interpretation of cosmic string detection and within that framework, we derive a connection between estimates of cosmic string locations and cosmic string tension G μ. We use this Bayesian framework to develop a machine learning framework for detecting strings from sky maps and outline how to implement this frameworkmore » with neural networks. The neural network we trained was able to detect and locate cosmic strings on noiseless CMB temperature map down to a string tension of G μ=5 ×10{sup −9} and when analyzing a CMB temperature map that does not contain strings, the neural network gives a 0.95 probability that G μ≤2.3×10{sup −9}.« less

  8. String Theory - The Physics of String-Bending and Other Electric Guitar Techniques

    PubMed Central

    Grimes, David Robert

    2014-01-01

    Electric guitar playing is ubiquitous in practically all modern music genres. In the hands of an experienced player, electric guitars can sound as expressive and distinct as a human voice. Unlike other more quantised instruments where pitch is a discrete function, guitarists can incorporate micro-tonality and, as a result, vibrato and sting-bending are idiosyncratic hallmarks of a player. Similarly, a wide variety of techniques unique to the electric guitar have emerged. While the mechano-acoustics of stringed instruments and vibrating strings are well studied, there has been comparatively little work dedicated to the underlying physics of unique electric guitar techniques and strings, nor the mechanical factors influencing vibrato, string-bending, fretting force and whammy-bar dynamics. In this work, models for these processes are derived and the implications for guitar and string design discussed. The string-bending model is experimentally validated using a variety of strings and vibrato dynamics are simulated. The implications of these findings on the configuration and design of guitars is also discussed. PMID:25054880

  9. ASSESSING CONTAMINANT SENSITIVITY OF ENDANGERED AND THREATENED SPECIES: EFFLUENT TOXICITY TESTS

    EPA Science Inventory

    Toxicity tests using standard effluent test procedures were conducted (EPA 1994) with Ceriodaphnia dubia and fathead minnows and four endangered fish species: bonytail chub (Gila elegans), Colorado squawfish (Ptychocheilus lucias ), razorback sucker (Xyrauchen texanus) and Gila t...

  10. Minimal string theories and integrable hierarchies

    NASA Astrophysics Data System (ADS)

    Iyer, Ramakrishnan

    Well-defined, non-perturbative formulations of the physics of string theories in specific minimal or superminimal model backgrounds can be obtained by solving matrix models in the double scaling limit. They provide us with the first examples of completely solvable string theories. Despite being relatively simple compared to higher dimensional critical string theories, they furnish non-perturbative descriptions of interesting physical phenomena such as geometrical transitions between D-branes and fluxes, tachyon condensation and holography. The physics of these theories in the minimal model backgrounds is succinctly encoded in a non-linear differential equation known as the string equation, along with an associated hierarchy of integrable partial differential equations (PDEs). The bosonic string in (2,2m-1) conformal minimal model backgrounds and the type 0A string in (2,4 m) superconformal minimal model backgrounds have the Korteweg-de Vries system, while type 0B in (2,4m) backgrounds has the Zakharov-Shabat system. The integrable PDE hierarchy governs flows between backgrounds with different m. In this thesis, we explore this interesting connection between minimal string theories and integrable hierarchies further. We uncover the remarkable role that an infinite hierarchy of non-linear differential equations plays in organizing and connecting certain minimal string theories non-perturbatively. We are able to embed the type 0A and 0B (A,A) minimal string theories into this single framework. The string theories arise as special limits of a rich system of equations underpinned by an integrable system known as the dispersive water wave hierarchy. We find that there are several other string-like limits of the system, and conjecture that some of them are type IIA and IIB (A,D) minimal string backgrounds. We explain how these and several other string-like special points arise and are connected. In some cases, the framework endows the theories with a non-perturbative definition for the first time. Notably, we discover that the Painleve IV equation plays a key role in organizing the string theory physics, joining its siblings, Painleve I and II, whose roles have previously been identified in this minimal string context. We then present evidence that the conjectured type II theories have smooth non-perturbative solutions, connecting two perturbative asymptotic regimes, in a 't Hooft limit. Our technique also demonstrates evidence for new minimal string theories that are not apparent in a perturbative analysis.

  11. Dynamical AdS strings across horizons

    DOE PAGES

    Ishii, Takaaki; Murata, Keiju

    2016-03-01

    We examine the nonlinear classical dynamics of a fundamental string in anti-deSitter spacetime. The string is dual to the flux tube between an external quark-antiquark pair in $N = 4$ super Yang-Mills theory. We perturb the string by shaking the endpoints and compute its time evolution numerically. We find that with sufficiently strong perturbations the string continues extending and plunges into the Poincare´ horizon. In the evolution, effective horizons are also dynamically created on the string worldsheet. The quark and antiquark are thus causally disconnected, and the string transitions to two straight strings. The forces acting on the endpoints vanishmore » with a power law whose slope depends on the perturbations. Lastly, the condition for this transition to occur is that energy injection exceeds the static energy between the quark-antiquark pair.« less

  12. Cosmic strings and the microwave sky. I - Anisotropy from moving strings

    NASA Technical Reports Server (NTRS)

    Stebbins, Albert

    1988-01-01

    A method is developed for calculating the component of the microwave anisotropy around cosmic string loops due to their rapidly changing gravitational fields. The method is only valid for impact parameters from the string much smaller than the horizon size at the time the photon passes the string. The method makes it possible to calculate the temperature pattern around arbitrary string configurations numerically in terms of one-dimensional integrals. This method is applied to temperature jump across a string, confirming and extending previous work. It is also applied to cusps and kinks on strings, and to determining the temperature pattern far from a strong loop. The temperature pattern around a few loop configurations is explicitly calculated. Comparisons with the work of Brandenberger et al. (1986) indicates that they have overestimated the MBR anisotropy from gravitational radiation emitted from loops.

  13. The Development of a String Sight-Reading Pitch Skill Hierarchy

    ERIC Educational Resources Information Center

    Alexander, Michael L.; Henry, Michele L.

    2012-01-01

    This study was designed to determine a pitch skill hierarchy for string sight-reading, to determine the effects of key on string sight-reading achievement, and to determine the validity of a tonal pattern system as a measurement of melodic sight-reading skill for string players. High school string students (n = 94) obtained a mean score of 27.28…

  14. Actomyosin purse strings: renewable resources that make morphogenesis robust and resilient

    PubMed Central

    Rodriguez-Diaz, Alice; Toyama, Yusuke; Abravanel, Daniel L.; Wiemann, John M.; Wells, Adrienne R.; Tulu, U. Serdar; Edwards, Glenn S.; Kiehart, Daniel P.

    2008-01-01

    Dorsal closure in Drosophila is a model system for cell sheet morphogenesis and wound healing. During closure two sheets of lateral epidermis move dorsally to close over the amnioserosa and form a continuous epidermis. Forces from the amnioserosa and actomyosin-rich, supracellular purse strings at the leading edges of these lateral epidermal sheets drive closure. Purse strings generate the largest force for closure and occur during development and wound healing throughout phylogeny. We use laser microsurgery to remove some or all of the purse strings from developing embryos. Free edges produced by surgery undergo characteristic responses as follows. Intact cells in the free edges, which previously had no purse string, recoil away from the incision and rapidly assemble new, secondary purse strings. Next, recoil slows, then pauses at a turning point. Following a brief delay, closure resumes and is powered to completion by the secondary purse strings. We confirm that the assembly of the secondary purse strings requires RhoA. We show that α-actinin alternates with nonmuscle myosin II along purse strings and requires nonmuscle myosin II for its localization. Together our data demonstrate that purse strings are renewable resources that contribute to the robust and resilient nature of closure. PMID:19404432

  15. Physical cognition: birds learn the structural efficacy of nest material

    PubMed Central

    Bailey, Ida E.; Morgan, Kate V.; Bertin, Marion; Meddle, Simone L.; Healy, Susan D.

    2014-01-01

    It is generally assumed that birds’ choice of structurally suitable materials for nest building is genetically predetermined. Here, we tested that assumption by investigating whether experience affected male zebra finches’ (Taeniopygia guttata) choice of nest material. After a short period of building with relatively flexible string, birds preferred to build with stiffer string while those that had experienced a stiffer string were indifferent to string type. After building a complete nest with either string type, however, all birds increased their preference for stiff string. The stiffer string appeared to be the more effective building material as birds required fewer pieces of stiffer than flexible string to build a roofed nest. For birds that raised chicks successfully, there was no association between the material they used to build their nest and the type they subsequently preferred. Birds’ material preference reflected neither the preference of their father nor of their siblings but juvenile experience of either string type increased their preference for stiffer string. Our results represent two important advances: (i) birds choose nest material based on the structural properties of the material; (ii) nest material preference is not entirely genetically predetermined as both the type and amount of experience influences birds’ choices. PMID:24741011

  16. Physical cognition: birds learn the structural efficacy of nest material.

    PubMed

    Bailey, Ida E; Morgan, Kate V; Bertin, Marion; Meddle, Simone L; Healy, Susan D

    2014-06-07

    It is generally assumed that birds' choice of structurally suitable materials for nest building is genetically predetermined. Here, we tested that assumption by investigating whether experience affected male zebra finches' (Taeniopygia guttata) choice of nest material. After a short period of building with relatively flexible string, birds preferred to build with stiffer string while those that had experienced a stiffer string were indifferent to string type. After building a complete nest with either string type, however, all birds increased their preference for stiff string. The stiffer string appeared to be the more effective building material as birds required fewer pieces of stiffer than flexible string to build a roofed nest. For birds that raised chicks successfully, there was no association between the material they used to build their nest and the type they subsequently preferred. Birds' material preference reflected neither the preference of their father nor of their siblings but juvenile experience of either string type increased their preference for stiffer string. Our results represent two important advances: (i) birds choose nest material based on the structural properties of the material; (ii) nest material preference is not entirely genetically predetermined as both the type and amount of experience influences birds' choices.

  17. Text String Detection from Natural Scenes by Structure-based Partition and Grouping

    PubMed Central

    Yi, Chucai; Tian, YingLi

    2012-01-01

    Text information in natural scene images serves as important clues for many image-based applications such as scene understanding, content-based image retrieval, assistive navigation, and automatic geocoding. However, locating text from complex background with multiple colors is a challenging task. In this paper, we explore a new framework to detect text strings with arbitrary orientations in complex natural scene images. Our proposed framework of text string detection consists of two steps: 1) Image partition to find text character candidates based on local gradient features and color uniformity of character components. 2) Character candidate grouping to detect text strings based on joint structural features of text characters in each text string such as character size differences, distances between neighboring characters, and character alignment. By assuming that a text string has at least three characters, we propose two algorithms of text string detection: 1) adjacent character grouping method, and 2) text line grouping method. The adjacent character grouping method calculates the sibling groups of each character candidate as string segments and then merges the intersecting sibling groups into text string. The text line grouping method performs Hough transform to fit text line among the centroids of text candidates. Each fitted text line describes the orientation of a potential text string. The detected text string is presented by a rectangle region covering all characters whose centroids are cascaded in its text line. To improve efficiency and accuracy, our algorithms are carried out in multi-scales. The proposed methods outperform the state-of-the-art results on the public Robust Reading Dataset which contains text only in horizontal orientation. Furthermore, the effectiveness of our methods to detect text strings with arbitrary orientations is evaluated on the Oriented Scene Text Dataset collected by ourselves containing text strings in non-horizontal orientations. PMID:21411405

  18. Text string detection from natural scenes by structure-based partition and grouping.

    PubMed

    Yi, Chucai; Tian, YingLi

    2011-09-01

    Text information in natural scene images serves as important clues for many image-based applications such as scene understanding, content-based image retrieval, assistive navigation, and automatic geocoding. However, locating text from a complex background with multiple colors is a challenging task. In this paper, we explore a new framework to detect text strings with arbitrary orientations in complex natural scene images. Our proposed framework of text string detection consists of two steps: 1) image partition to find text character candidates based on local gradient features and color uniformity of character components and 2) character candidate grouping to detect text strings based on joint structural features of text characters in each text string such as character size differences, distances between neighboring characters, and character alignment. By assuming that a text string has at least three characters, we propose two algorithms of text string detection: 1) adjacent character grouping method and 2) text line grouping method. The adjacent character grouping method calculates the sibling groups of each character candidate as string segments and then merges the intersecting sibling groups into text string. The text line grouping method performs Hough transform to fit text line among the centroids of text candidates. Each fitted text line describes the orientation of a potential text string. The detected text string is presented by a rectangle region covering all characters whose centroids are cascaded in its text line. To improve efficiency and accuracy, our algorithms are carried out in multi-scales. The proposed methods outperform the state-of-the-art results on the public Robust Reading Dataset, which contains text only in horizontal orientation. Furthermore, the effectiveness of our methods to detect text strings with arbitrary orientations is evaluated on the Oriented Scene Text Dataset collected by ourselves containing text strings in nonhorizontal orientations.

  19. How to simulate global cosmic strings with large string tension

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Klaer, Vincent B.; Moore, Guy D., E-mail: vklaer@theorie.ikp.physik.tu-darmstadt.de, E-mail: guy.moore@physik.tu-darmstadt.de

    Global string networks may be relevant in axion production in the early Universe, as well as other cosmological scenarios. Such networks contain a large hierarchy of scales between the string core scale and the Hubble scale, ln( f {sub a} / H ) ∼ 70, which influences the network dynamics by giving the strings large tensions T ≅ π f {sub a} {sup 2} ln( f {sub a} / H ). We present a new numerical approach to simulate such global string networks, capturing the tension without an exponentially large lattice.

  20. Closed timelike curves produced by pairs of moving cosmic strings - Exact solutions

    NASA Technical Reports Server (NTRS)

    Gott, J. Richard, III

    1991-01-01

    Exact solutions of Einstein's field equations are presented for the general case of two moving straight cosmic strings that do not intersect. The solutions for parallel cosmic strings moving in opposite directions show closed timelike curves (CTCs) that circle the two strings as they pass, allowing observers to visit their own past. Similar results occur for nonparallel strings, and for masses in (2+1)-dimensional spacetime. For finite string loops the possibility that black-hole formation may prevent the formation of CTCs is discussed.

  1. Coulomb string tension, asymptotic string tension, and the gluon chain

    DOE PAGES

    Greensite, Jeff; Szczepaniak, Adam P.

    2015-02-01

    We compute, via numerical simulations, the non-perturbative Coulomb potential and position-space ghost propagator in pure SU(3) gauge theory in Coulomb gauge. We find that that the Coulomb potential scales nicely in accordance with asymptotic freedom, that the Coulomb potential is linear in the infrared, and that the Coulomb string tension is about four times larger than the asymptotic string tension. We explain how it is possible that the asymptotic string tension can be lower than the Coulomb string tension by a factor of four.

  2. Simulation of swimming strings immersed in a viscous fluid flow

    NASA Astrophysics Data System (ADS)

    Huang, Wei-Xi; Sung, Hyung Jin

    2006-11-01

    In nature, many phenomena involve interactions between flexible bodies and their surrounding viscous fluid, such as a swimming fish or a flapping flag. The intrinsic dynamics is complicate and not well understood. A flexible string can be regarded as a one-dimensional flag model. Many similarities can be found between the flapping string and swimming fish, although different wake speed results in a drag force for the flapping string and a propulsion force for the swimming fish. In the present study, we propose a mathematical formulation for swimming strings immersed in a viscous fluid flow. Fluid motion is governed by the Navier-Stokes equations and a momentum forcing is added in order to bring the fluid to move at the same velocity with the immersed surface. A flexible inextensible string model is described by another set of equations with an additional momentum forcing which is a result of the fluid viscosity and the pressure difference across the string. The momentum forcing is calculated by a feedback loop. Simulations of several numerical examples are carried out, including a hanging string which starts moving under gravity without ambient fluid, a swinging string immersed in a quiescent viscous fluid, a string swimming within a uniform surrounding flow, and flow over two side-by-side strings. The numerical results agree well with the theoretical analysis and previous experimental observations. Further simulation of a swimming fish is under consideration.

  3. Noncommutative Field Theories and (super)string Field Theories

    NASA Astrophysics Data System (ADS)

    Aref'eva, I. Ya.; Belov, D. M.; Giryavets, A. A.; Koshelev, A. S.; Medvedev, P. B.

    2002-11-01

    In this lecture notes we explain and discuss some ideas concerning noncommutative geometry in general, as well as noncommutative field theories and string field theories. We consider noncommutative quantum field theories emphasizing an issue of their renormalizability and the UV/IR mixing. Sen's conjectures on open string tachyon condensation and their application to the D-brane physics have led to wide investigations of the covariant string field theory proposed by Witten about 15 years ago. We review main ingredients of cubic (super)string field theories using various formulations: functional, operator, conformal and the half string formalisms. The main technical tools that are used to study conjectured D-brane decay into closed string vacuum through the tachyon condensation are presented. We describe also methods which are used to study the cubic open string field theory around the tachyon vacuum: construction of the sliver state, "comma" and matrix representations of vertices.

  4. Gravitational lensing effects of vacuum strings - Exact solutions

    NASA Technical Reports Server (NTRS)

    Gott, J. R., III

    1985-01-01

    Exact interior and exterior solutions to Einstein's field equations are derived for vacuum strings. The exterior solution for a uniform density vacuum string corresponds to a conical space while the interior solution is that of a spherical cap. For Mu equals 0-1/4 the external metric is ds-squared = -dt-squared + dr-squared + (1-4 Mu)-squared r-squared dphi-squared + dz-squared, where Mu is the mass per unit length in the string in Planck masses per Planck length. A maximum mass per unit length for a string is 6.73 x 10 to the 27th g/cm. It is shown that strings cause temperature fluctuations in the cosmic microwave background and produce equal brightness double QSO images separated by up to several minutes of arc. Formulae for lensing probabilities, image splittings, and time delays are derived for strings in a realistic cosmological setting. String searches using ST, the VLA, and the COBE satellite are discussed.

  5. Fitting cosmic microwave background data with cosmic strings and inflation.

    PubMed

    Bevis, Neil; Hindmarsh, Mark; Kunz, Martin; Urrestilla, Jon

    2008-01-18

    We perform a multiparameter likelihood analysis to compare measurements of the cosmic microwave background (CMB) power spectra with predictions from models involving cosmic strings. Adding strings to the standard case of a primordial spectrum with power-law tilt ns, we find a 2sigma detection of strings: f10=0.11+/-0.05, where f10 is the fractional contribution made by strings in the temperature power spectrum (at l=10). CMB data give moderate preference to the model ns=1 with cosmic strings over the standard zero-strings model with variable tilt. When additional non-CMB data are incorporated, the two models become on a par. With variable ns and these extra data, we find that f10<0.11, which corresponds to Gmicro<0.7x10(-6) (where micro is the string tension and G is the gravitational constant).

  6. Modal analysis of a nonuniform string with end mass and variable tension

    NASA Technical Reports Server (NTRS)

    Rheinfurth, M. H.; Galaboff, Z. J.

    1983-01-01

    Modal synthesis techniques for dynamic systems containing strings describe the lateral displacements of these strings by properly chosen shape functions. An iterative algorithm is provided to calculate the natural modes of a nonuniform string and variable tension for some typical boundary conditions including one end mass. Numerical examples are given for a string in a constant and a gravity gradient force field.

  7. Self-energy and self-force in the space-time of a thick cosmic string

    NASA Astrophysics Data System (ADS)

    Khusnutdinov, N. R.; Bezerra, V. B.

    2001-10-01

    We calculate the self-energy and self-force for an electrically charged particle at rest in the background of Gott-Hiscock cosmic string space-time. We find the general expression for the self-energy which is expressed in terms of the S matrix of the scattering problem. The self-energy continuously falls down outward from the string's center with the maximum at the origin of the string. The self-force is repulsive for an arbitrary position of the particle. It tends to zero in the string's center and also far from the string and it has a maximum value at the string's surface. The plots of the numerical calculations of the self-energy and self-force are shown.

  8. The implications of the COBE diffuse microwave radiation results for cosmic strings

    NASA Technical Reports Server (NTRS)

    Bennett, David P.; Stebbins, Albert; Bouchet, Francois R.

    1992-01-01

    We compare the anisotropies in the cosmic microwave background radiation measured by the COBE experiment to those predicted by cosmic string theories. We use an analytic model for the Delta T/T power spectrum that is based on our previous numerical simulations of strings, under the assumption that cosmic strings are the sole source of the measured anisotropy. This implies a value for the string mass per unit length of 1.5 +/- 0.5 x 10 exp -6 C-squared/G. This is within the range of values required for cosmic strings to successfully seed the formation of large-scale structures in the universe. These results clearly encourage further studies of Delta T/T and large-scale structure in the cosmic string model.

  9. Identification of market trends with string and D2-brane maps

    NASA Astrophysics Data System (ADS)

    Bartoš, Erik; Pinčák, Richard

    2017-08-01

    The multidimensional string objects are introduced as a new alternative for an application of string models for time series forecasting in trading on financial markets. The objects are represented by open string with 2-endpoints and D2-brane, which are continuous enhancement of 1-endpoint open string model. We show how new object properties can change the statistics of the predictors, which makes them the candidates for modeling a wide range of time series systems. String angular momentum is proposed as another tool to analyze the stability of currency rates except the historical volatility. To show the reliability of our approach with application of string models for time series forecasting we present the results of real demo simulations for four currency exchange pairs.

  10. Mechanism of Tennis Racket Spin Performance

    NASA Astrophysics Data System (ADS)

    Kawazoe, Yoshihiko; Okimoto, Kenji; Okimoto, Keiko

    Players often say that some strings provide a better grip and more spin than others, but ball spin did not depend on string type, gauge, or tension in pervious laboratory experiments. There was no research work on spin to uncover what is really happening during an actual tennis impact because of the difficulty of performing the appropriate experiments. The present paper clarified the mechanism of top spin and its improvement by lubrication of strings through the use of high-speed video analysis. It also provided a more detailed explanation of spin behavior by comparing a racket with lubricated strings with the famous “spaghetti” strung racket, which was banned in 1978 by the International Tennis Federation because it used plastic spaghetti tubing over the strings to reduce friction, resulting in excessive ball spin. As the main strings stretch and slide sideways more, the ball is given additional spin due to the restoring force parallel to the string face when the main strings spring back and the ball is released from the strings. Herein, we also showed that the additional spin results in a reduction of shock vibrations of the wrist joint during impact.

  11. Contaminants of legacy and emerging concern in largescale suckers (Catostomus macrocheilus) and the foodweb in the lower Columbia River, Oregon and Washington, USA.

    PubMed

    Nilsen, Elena; Zaugg, Steven; Alvarez, David; Morace, Jennifer; Waite, Ian; Counihan, Timothy; Hardiman, Jill; Torres, Leticia; Patiño, Reynaldo; Mesa, Matthew; Grove, Robert

    2014-06-15

    We investigated occurrence, transport pathways, and effects of polybrominated diphenyl ether (PBDE) flame retardants and other endocrine disrupting chemicals (EDCs) in aquatic media and the foodweb in the lower Columbia River. In 2009 and 2010, foodweb sampling at three sites along a gradient of contaminant exposure near Skamania (Washington), Columbia City (Oregon) and Longview (Washington) included water (via passive samplers), bed sediment, invertebrate biomass residing in sediment, a resident fish species (largescale suckers [Catostomus macrocheilus]), and eggs from osprey (Pandion haliaetus). This paper primarily reports fish tissue concentrations. In 2009, composites of fish brain, fillet, liver, stomach, and gonad tissues revealed that overall contaminant concentrations were highest in livers, followed by brain, stomach, gonad, and fillet. Concentrations of halogenated compounds in tissue samples from all three sites ranged from <1 to 400nanograms per gram of wet tissue. Several chemical classes, including PBDEs, organochlorine pesticides, and polychlorinated biphenyls (PCBs), were detected at all sites and in nearly all fish tissues sampled. In 2010, only fish livers were sampled and inter-site concentration differences were not as pronounced as in 2009. Chemical concentrations in sediments, fish tissues, and osprey eggs increased moving downstream from Skamania to the urbanized sites near Columbia City and Longview. Numerous organochlorine (OC) pesticides, both banned and currently used, and PBDEs, were present at each site in multiple media and concentrations exceeded environmental quality benchmarks in some cases. Frequently detected OC compounds included hexachlorobenzene, pentachloroanisole, dichlorodiphenyltrichloroethane (DDT) and its degradates, chlorpyrifos, and oxyfluorofen. Biomagnification of BDE47, 100, 153, and 154 occurred in largescale suckers and osprey eggs. Results support the hypothesis that contaminants in the environment lead to bioaccumulation and potential negative effects in multiple levels of the foodweb. Published by Elsevier B.V.

  12. Stellantchasmus falcatus (Digenea: Heterophyidae) in Cambodia: Discovery of Metacercariae in Mullets and Recovery of Adult Flukes in an Experimental Hamster

    PubMed Central

    Chai, Jong-Yil; Sohn, Woon-Mok; Na, Byoung-Kuk; Jeoung, Hoo-Gn; Sinuon, Muth; Socheat, Duong

    2016-01-01

    Stellantchasmus falcatus (Digenea: Heterophyidae) is first reported from Cambodia through recovery of the metacercariae from mullet fish and adult flukes from an experimentally infected hamster. We purchased 7 mullets, Chelon macrolepis, in a local market of Phnom Penh, Cambodia, and each of them was examined by the artificial digestion method on May 2010. The metacercariae of S. falcatus were detected in all mullets (100%) examined, and their average density was 177 per fish. They were elliptical, 220×168 μm in average size. They were orally infected to an hamster to obtain adult flukes. Adults recovered at day 10 post infection were observed with a light microscope and a scanning electron microscope (SEM). They were small, 450×237 μm in average size, had a small oral sucker (41×50 μm), subglobular pharynx (29×21 μm), slender esophagus (57 μm), long and thick-walled expulsor (119×32 μm), spherical ovary (58×69 μm), and 2 ovoid testes (right: 117×74 μm; left: 114×63 μm). Eggs were small, yellow, and 23×12 μm in average size. In SEM observations, tegumental spines were densely distributed on the whole tegument, and single small type I sensory papillae were distributed around the lip of oral sucker. The small ventral sucker was dextrally located and had 8 type I sensory papillae on the left margin. It has been first confirmed in the present study that the mullet, C. macrolepis, is playing the role of a second intermediate host of S. falcatus in Cambodia. PMID:27658608

  13. New circumscription of freshwater fish parasites Monobothrium diesing, 1863 and Promonobothrium mackiewicz, 1968 (Cestoda: Caryophyllidea) using morphological and molecular evidence.

    PubMed

    Scholz, Tomáš; Oros, Mikuláš; Choudhury, Anindo; Brabec, Jan; Waeschenbach, Andrea

    2015-02-01

    Two genera of caryophyllidean cestodes, Monobothrium Diesing, 1863 and Promonobothrium Mackiewicz, 1968 , from cypriniform fishes (cyprinids and suckers) in the Holarctic Region, are revised using morphological and molecular evidence. Monobothrium, which includes morphologically distinct species that occur in European cyprinids (Cyprinidae) and North American suckers (Catostomidae), is separated into 3 genera. Monobothrium becomes monotypic and is represented by its type species, Monobothrium wageneri Nybelin, 1922 , which occurs in European tench, Tinca tinca (Cyprinidae). Monobothrium auriculatum Kulakovskaya, 1961 from Leuciscus danilevskii (Cyprinidae) in the Ukraine, is tentatively transferred to Caryophyllaeus Gmelin, 1790 as Caryophyllaeus auriculatus ( Kulakovskaya, 1961 ) n. comb. because it possesses morphological characteristics considered to be typical of Caryophyllaeus but which are absent in Monobothrium (shape of the scolex, presence of a seminal receptacle, short neck, and absence of a large, muscular papilla surrounding the large gonopores). The 5 remaining Monobothrium species from suckers in North America are transferred to Promonobothrium based on shared morphological features and molecular data. Species of Promonobothrium differ from M. wageneri by having an external seminal vesicle (absent in M. wageneri), the absence of postovarian vitelline follicles in North American species (present in M. wageneri), and a scolex that is digitiform papillate, loculopapillate, or loculotruncate, i.e., equipped with weak loculi and a terminal introvert (vs. claviform, bluntly ended, with 6 weak, shallow, longitudinal grooves in M. wageneri). Phylogenetic analysis of nuclear ribosomal RNA genes ssr- and lsrDNA placed Promonobothrium minytremi within a clade of 3 of the 5 North American Monobothrium species. The position of M. wageneri, although distinct from the remaining Monobothrium species, is unresolved within the Caryophyllidea. This study further supports the pattern that the biogeography of caryophyllidean taxa is generally constrained by host and continental association.

  14. Redescription of Corallobothrium solidum (Cestoda: Proteocephalidea) and erection of a new genus, Essexiella, for tapeworms from channel catfishes (Ictaluridae).

    PubMed

    Scholz, Tomáš; de Chambrier, Alain; Mariaux, Jean; Kuchta, Roman

    2011-12-01

    The proteocephalidean tapeworm, Corallobothrium solidum, type species of the genus, is redescribed on the basis of the examination of its type specimens and extensive material recently collected from Malapterurus electricus (type host). Some morphological characteristics of taxonomic importance are reported for the first time, such as the presence of semispherical (U-shaped) sphincters on the external (outer) margin of the suckers, a vaginal sphincter, a well-developed seminal receptacle, and a unique morphology of the eggs. Corallobothrium solidum differs from the 2 remaining species of the genus, both parasitic in channel catfishes (Ictaluridae), in its scolex shape, morphology of its suckers, presence of longitudinal and transverse grooves on the body surface, dense network of excretory canals in the apical part of the scolex, morphology of the eggs, and uterine development. The non-monophyletic nature of Corallobothrium is further supported by molecular data (partial sequences of the 28S rRNA gene) because C. solidum and the 2 remaining species from ictalurids do not form a monophyletic assemblage. Therefore, Essexiella n. gen. is proposed to accommodate Essexiella fimbriatum new comb. (type and only species; syn. Corallobothrium fimbriatum) from channel catfish. Essexiella n. gen. differs from Corallobothrium, Megathylacoides, and Megathylacus by the absence of a sphincter in the suckers, from Corallotaenia by the shape of the scolex and the number and shape of proglottids, and from Paraproteocephalus by the structure of the uterus. The diagnosis of Corallobothrium, which becomes monotypic and restricted to electric catfishes in Africa, is emended. The remaining species of Corallobothrium, Corallobothrium parafimbriatum, is tentatively transferred to Corallotaenia as Corallotaenia parafimbriata n. comb., based on molecular data, small size of the strobila, and shape of the scolex.

  15. Echinostoma macrorchis (Digenea: Echinostomatidae): Metacercariae in Cipangopaludina chinensis malleata Snails and Adults from Experimental Rats in Korea.

    PubMed

    Sohn, Woon-Mok; Na, Byoung-Kuk

    2017-10-01

    A survey was performed to know the infection status of echinostome metacercariae in Cipangopaludina chinensis malleata snails from Korea. Total 75 snails collected in 5 localities, i.e., Imsil-gun, Jeollabuk-do, Hwasun-gun and Shinan-gun (Aphae and Jido), Jeollanam-do, and Jinju-si, Gyeongsangnam-do, were examined for metacercariae by the artificial digestion method. Infection rates of metacercariae were 80.0%, 66.7%, 100%, 60.0%, and 73.3%, and their densities were 39, 32, 183, 19, and 30 per snail infected, respectively. The metacercariae were round, 105-118×105-118 μm in size, with a thin cyst wall, collar spines on the head collar, and excretory granules in 2 canals of excretory tube. Adult flukes were elongated, ventrally curved, and 5,167×939 μm in average size. Head collar distinct, bearing 45 collar spines with 5 end groups on each side. Oral sucker subterminal, pharynx well developed, and esophagus somewhat short. Cirrus sac well developed, with a saccular seminal vesicle, and ventral sucker very large. Ovary elliptical and on the median line of the body. Testes tandem and slightly lobed. Eggs operculated, elliptical, and 90-103×55-60 μm in size. By scanning electron microscopy, the head collar was prominent with 45 collar spines resembling horns of younger stags. Scale-like tegumental spines were densely distributed on the body surface between the head collar and ventral sucker. Conclusively, it has been first confirmed that the life cycle of E. macrorchis is indigenously maintained in Korea, and C. chinensis malleata snails are popularly infected with the metacercariae of this echinostome.

  16. The Carcinogenic Liver Fluke, Clonorchis sinensis: New Assembly, Reannotation and Analysis of the Genome and Characterization of Tissue Transcriptomes

    PubMed Central

    Wang, Xiaoyun; Liu, Hailiang; Chen, Yangyi; Guo, Lei; Luo, Fang; Sun, Jiufeng; Mao, Qiang; Liang, Pei; Xie, Zhizhi; Zhou, Chenhui; Tian, Yanli; Lv, Xiaoli; Huang, Lisi; Zhou, Juanjuan; Hu, Yue; Li, Ran; Zhang, Fan; Lei, Huali; Li, Wenfang; Hu, Xuchu; Liang, Chi; Xu, Jin; Li, Xuerong; Yu, Xinbing

    2013-01-01

    Clonorchis sinensis (C. sinensis), an important food-borne parasite that inhabits the intrahepatic bile duct and causes clonorchiasis, is of interest to both the public health field and the scientific research community. To learn more about the migration, parasitism and pathogenesis of C. sinensis at the molecular level, the present study developed an upgraded genomic assembly and annotation by sequencing paired-end and mate-paired libraries. We also performed transcriptome sequence analyses on multiple C. sinensis tissues (sucker, muscle, ovary and testis). Genes encoding molecules involved in responses to stimuli and muscle-related development were abundantly expressed in the oral sucker. Compared with other species, genes encoding molecules that facilitate the recognition and transport of cholesterol were observed in high copy numbers in the genome and were highly expressed in the oral sucker. Genes encoding transporters for fatty acids, glucose, amino acids and oxygen were also highly expressed, along with other molecules involved in metabolizing these substrates. All genes involved in energy metabolism pathways, including the β-oxidation of fatty acids, the citrate cycle, oxidative phosphorylation, and fumarate reduction, were expressed in the adults. Finally, we also provide valuable insights into the mechanism underlying the process of pathogenesis by characterizing the secretome of C. sinensis. The characterization and elaborate analysis of the upgraded genome and the tissue transcriptomes not only form a detailed and fundamental C. sinensis resource but also provide novel insights into the physiology and pathogenesis of C. sinensis. We anticipate that this work will aid the development of innovative strategies for the prevention and control of clonorchiasis. PMID:23382950

  17. A revision of the haploporinae nicoll, 1914 (digenea: haploporidae) from mullets (mugilidae): Dicrogaster Looss, 1902 and Forticulcita overstreet, 1982.

    PubMed

    Blasco-Costa, Isabel; Montero, Francisco E; Balbuena, Juan Antonio; Raga, Juan Antonio; Kostadinova, Aneta

    2009-03-01

    The status of the nominal species of Dicrogaster is re-assessed by means of a comparative morphological study based on newly collected material from the western Mediterranean and a critical evaluation of the published data. D. perpusilla Looss, 1902 (type-species) and D. contracta Looss, 1902 are redescribed on the basis of new material from Liza spp. D. fragilis Fernández Bargiela, 1987 is considered a junior synonym of D. fastigata Thatcher & Sparks, 1958 and D. maryutensis Al-Bassel, 1990 is considered to be nomen nudum. A key to the recognised species of Dicrogaster is presented. A new species of Forticulcita Overstreet, 1982 is described from Mugil cephalus L. in the Spanish Mediterranean and differentiated from the type-species, F. glabra Overstreet, 1982, and the recently described F. mugilis Hassanine, 2007 by its significantly smaller body size and most of its metrical data. The new species, F. gibsoni n. sp., is characterised by a narrow 'neck' region anterior to the genital pore and a long striated 'tail' region in the hindbody which is devoid of organs, such that the bulk of the uterus is restricted to the anterior half of the hindbody and the testis is located more anteriorly. F. gibsoni differs from F. glabra in having a longer forebody, smaller sucker width ratio and larger eggs, and from F. mugilis in having a smaller sucker width ratio, a hermaphroditic sac much longer than the ventral sucker and smaller eggs. The three Mediterranean forms, D. perpusilla, D. contracta and F. gibsoni are further distinguished by multivariate analyses. Keys to the species of both genera are presented.

  18. Identifying preservation and restoration priority areas for desert fishes in an increasingly invaded world.

    PubMed

    Pool, Thomas K; Strecker, Angela L; Olden, Julian D

    2013-03-01

    A commonly overlooked aspect of conservation planning assessments is that wildlife managers are increasingly focused on habitats that contain non-native species. We examine this management challenge in the Gila River basin (150,730 km(2)), and present a new planning strategy for fish conservation. By applying a hierarchical prioritization algorithm to >850,000 fish records in 27,181 sub-watersheds we first identified high priority areas (PAs) termed "preservation PAs" with high native fish richness and low non-native richness; these represent traditional conservation targets. Second, we identified "restoration PAs" with high native fish richness that also contained high numbers of non-native species; these represent less traditional conservation targets. The top 10 % of preservation and restoration PAs contained common native species (e.g., Catostomus clarkii, desert sucker; Catostomus insignis, Sonora sucker) in addition to native species with limited distributions (i.e., Xyrauchen texanus, razorback sucker; Oncorhynchus gilae apache, Apache trout). The top preservation and restoration PAs overlapped by 42 %, indicating areas with high native fish richness range from minimally to highly invaded. Areas exclusively identified as restoration PAs also encompassed a greater percentage of native species ranges than would be expected by the random addition of an equivalent basin area. Restoration PAs identified an additional 19.0 and 26.6 % of the total ranges of two federally endangered species-Meda fulgida (spikedace) and Gila intermedia (Gila chub), respectively, compared to top preservation PAs alone-despite adding only 5.8 % of basin area. We contend that in addition to preservation PAs, restoration PAs are well suited for complementary management activities benefiting native fishes.

  19. Inhibition of erythrocytes δ-aminolevulinic acid dehydratase (ALAD) activity in fish from waters affected by lead smelters

    USGS Publications Warehouse

    Schmitt, Christopher J.; Caldwell, Colleen A.; Olsen, Bill; Serdar, Dave; Coffey, Mike

    2002-01-01

    We assessed the effects on fish of lead (Pb) released to streamsby smelters located in Trail, BC (Canada), E. Helena, MT, Herculaneum, MO, and Glover, MO. Fish were collected by electrofishing from sites located downstream of smelters and from reference sites. Blood from each fish was analyzed for δ-aminolevulinic acid dehydratase (ALAD) activity and hemoglobin (Hb), and samples of blood, liver, or carcass were analyzed for Pb, zinc (Zn), or both. Fish collected downstreamof all four smelters sites had elevated Pb concentrations, decreased ALAD activity, or both relative to their respectivereference sites. At E. Helena, fish from the downstream site also had lower Hb concentrations than fish from upstream. Differences among taxa were also apparent. Consistent with previous studies, ALAD activity in catostomids (Pisces: Catostomidae-northern hog sucker,Hypentelium nigricans;river carpsucker, Carpiodes carpio; largescale sucker, Catostomus macrocheilus; and mountain sucker, C. platyrhynchus) seemed more sensitive to Pb-induced ALADinhibition than the salmonids (Pisces: Salmonidae-rainbow trout,Oncorhynchus mykiss; brook trout,Salvelinus fontinalis) or common carp (Cyprinus carpio). Some of these differences may have resulted from differential accumulation of Zn, which was not measured at all sites. We detected noALAD activity in channel catfish (Ictaluruspunctatus) from either site on the Mississippi River at Herculaneum, MO. Our findings confirmed that Pb is releasedto aquatic ecosystems by smelters and accumulated by fish, andwe documented potentially adverse effects of Pb in fish. We recommend that Zn be measured along with Pb when ALAD activityis used as a biomarker and the collection of at least 10 fish ofa species at each site to facilitate statistical analysis.

  20. Echinostoma macrorchis (Digenea: Echinostomatidae): Metacercariae in Cipangopaludina chinensis malleata Snails and Adults from Experimental Rats in Korea

    PubMed Central

    Sohn, Woon-Mok; Na, Byoung-Kuk

    2017-01-01

    A survey was performed to know the infection status of echinostome metacercariae in Cipangopaludina chinensis malleata snails from Korea. Total 75 snails collected in 5 localities, i.e., Imsil-gun, Jeollabuk-do, Hwasun-gun and Shinan-gun (Aphae and Jido), Jeollanam-do, and Jinju-si, Gyeongsangnam-do, were examined for metacercariae by the artificial digestion method. Infection rates of metacercariae were 80.0%, 66.7%, 100%, 60.0%, and 73.3%, and their densities were 39, 32, 183, 19, and 30 per snail infected, respectively. The metacercariae were round, 105–118×105–118 μm in size, with a thin cyst wall, collar spines on the head collar, and excretory granules in 2 canals of excretory tube. Adult flukes were elongated, ventrally curved, and 5,167×939 μm in average size. Head collar distinct, bearing 45 collar spines with 5 end groups on each side. Oral sucker subterminal, pharynx well developed, and esophagus somewhat short. Cirrus sac well developed, with a saccular seminal vesicle, and ventral sucker very large. Ovary elliptical and on the median line of the body. Testes tandem and slightly lobed. Eggs operculated, elliptical, and 90–103×55–60 μm in size. By scanning electron microscopy, the head collar was prominent with 45 collar spines resembling horns of younger stags. Scale-like tegumental spines were densely distributed on the body surface between the head collar and ventral sucker. Conclusively, it has been first confirmed that the life cycle of E. macrorchis is indigenously maintained in Korea, and C. chinensis malleata snails are popularly infected with the metacercariae of this echinostome. PMID:29103269

  1. Contaminants of legacy and emerging concern in largescale suckers (Catostomus macrocheilus) and the foodweb in the lower Columbia River, Oregon and Washington, USA

    USGS Publications Warehouse

    Nilsen, Elena B.; Zaugg, Steven D.; Alvarez, David A.; Morace, Jennifer L.; Waite, Ian R.; Counihan, Timothy D.; Hardiman, Jill M.; Torres, Leticia; Patino, Reynaldo; Mesa, Matthew G.; Grove, Robert

    2014-01-01

    We investigated occurrence, transport pathways, and effects of polybrominated diphenyl ether (PBDE) flame retardants and other endocrine disrupting chemicals (EDCs) in aquatic media and the foodweb in the lower Columbia River. In 2009 and 2010, foodweb sampling at three sites along a gradient of contaminant exposure near Skamania (Washington), Columbia City (Oregon) and Longview (Washington) included water (via passive samplers), bed sediment, invertebrate biomass residing in sediment, a resident fish species (largescale suckers [Catostomus macrocheilus]), and eggs from osprey (Pandion haliaetus). This paper primarily reports fish tissue concentrations. In 2009, composites of fish brain, fillet, liver, stomach, and gonad tissues revealed that overall contaminant concentrations were highest in livers, followed by brain, stomach, gonad, and fillet. Concentrations of halogenated compounds in tissue samples from all three sites ranged from < 1 to 400 nanograms per gram of wet tissue. Several chemical classes, including PBDEs, organochlorine pesticides, and polychlorinated biphenyls (PCBs), were detected at all sites and in nearly all fish tissues sampled. In 2010, only fish livers were sampled and inter-site concentration differences were not as pronounced as in 2009. Chemical concentrations in sediments, fish tissues, and osprey eggs increased moving downstream from Skamania to the urbanized sites near Columbia City and Longview. Numerous organochlorine (OC) pesticides, both banned and currently used, and PBDEs, were present at each site in multiple media and concentrations exceeded environmental quality benchmarks in some cases. Frequently detected OC compounds included hexachlorobenzene, pentachloroanisole, dichlorodiphenyltrichloroethane (DDT) and its degradates, chlorpyrifos, and oxyfluorofen. Biomagnification of BDE47, 100, 153, and 154 occurred in largescale suckers and osprey eggs. Results support the hypothesis that contaminants in the environment lead to bioaccumulation and potential negative effects in multiple levels of the foodweb.

  2. Replacement of fish oil with soybean oil in diets for juvenile Chinese sucker (Myxocyprinus asiaticus): effects on liver lipid peroxidation and biochemical composition.

    PubMed

    Yu, Deng-Hang; Chang, Jia-Zhi; Dong, Gui-Fang; Liu, Jun

    2017-10-01

    This study was designed to evaluate the effect of the replacement of fish oil (FO) by soybean oil (SO) on growth performance, liver lipid peroxidation, and biochemical composition in juvenile Chinese sucker, Myxocyprinus asiaticus. Fish (13.7 ± 0.2 g) in triplicate were fed five experimental diets in which 0% (FO as control), 40% (SO40), 60% (SO60), 80% (SO40), and 100% (SO100) FO were replaced by SO. The body weight gain of fish fed SO40, SO60, or SO80 diet was similar to FO group, but diets that have 100% soybean oil as dietary lipid significantly reduced fish growth (P < 0.05). Although the level of SO resulted in increasing crude lipid content of the liver, the level of SO did not significantly alter the hepatosomatic index (HSI). Indicators of peroxidation, such as vitamin E (V E ) and thiobarbituric acid-reactive substance (TBARS) contents, were changed as increasing dietary SO. It was shown that the inclusion of SO in the diets increased V E concentrations, but reduced TBARS in the liver and total cholesterol (T-CHO) in the plasma. Linoleic acid (LA) and linolenic acid (LNA) significantly increased in fish liver fed diets that contained SO, but eicosapentaenoic acid (EPA), docosahexaenoic acid (DHA), and the ratio n-3/n-6 were significantly reduced by the inclusion of dietary SO (P < 0.05). Our results indicated that the inclusion of SO increased the hepatic V E content and reduced lipid peroxidation in fish. However, diet containing 100% SO as dietary lipid could reduce growth performance. Thus, we recommended that 40-80% SO can be used as dietary lipid to replace FO for juvenile Chinese sucker.

  3. Cardiorespiratory toxicity of environmentally relevant zinc oxide nanoparticles in the freshwater fish Catostomus commersonii.

    PubMed

    Bessemer, Robin Anne; Butler, Kathryn Marie Alison; Tunnah, Louise; Callaghan, Neal Ingraham; Rundle, Amanda; Currie, Suzanne; Dieni, Christopher Anthony; MacCormack, Tyson James

    2015-01-01

    The inhalation of zinc oxide engineered nanomaterials (ENMs) has been linked to cardiorespiratory dysfunction in mammalian models but the effects of aquatic ENM exposure on fish have not been fully investigated. Nano-zinc oxide (nZnO) is widely used in consumer products such as sunscreens and can make its way into aquatic ecosystems from domestic and commercial wastewater. This study examined the impact of an environmentally relevant nZnO formulation on cardiorespiratory function and energy metabolism in the white sucker (Catostomus commersonii), a freshwater teleost fish. Evidence of oxidative and cellular stress was present in gill tissue, including increases in malondialdehyde levels, heat shock protein (HSP) expression, and caspase 3/7 activity. Gill Na(+)/K(+)-ATPase activity was also higher by approximately three-fold in nZnO-treated fish, likely in response to increased epithelial permeability or structural remodeling. Despite evidence of toxicity in gill, plasma cortisol and lactate levels did not change in animals exposed to 1.0 mg L(-1) nZnO. White suckers also exhibited a 35% decrease in heart rate during nZnO exposure, with no significant changes in resting oxygen consumption or tissue energy stores. Our results suggest that tissue damage or cellular stress resulting from nZnO exposure activates gill neuroepithelial cells, triggering a whole-animal hypoxic response. An increase in parasympathetic nervous signaling will decrease heart rate and may reduce energy demand, even in the face of an environmental toxicant. We have shown that acute exposure to nZnO is toxic to white suckers and that ENMs have the potential to negatively impact cardiorespiratory function in adult fish.

  4. Two Studies of Pitch in String Instrument Vibrato: Perception and Pitch Matching Responses of University and High School String Players

    ERIC Educational Resources Information Center

    Geringer, John M.; MacLeod, Rebecca B.; Ellis, Julia C.

    2014-01-01

    We investigated pitch perception of string vibrato tones among string players in two separate studies. In both studies we used tones of acoustic instruments (violin and cello) as stimuli. In the first, we asked 192 high school and university string players to listen to a series of tonal pairs: one tone of each pair was performed with vibrato and…

  5. Geometric phase for a static two-level atom in cosmic string spacetime

    NASA Astrophysics Data System (ADS)

    Cai, Huabing; Ren, Zhongzhou

    2018-05-01

    We investigate the geometric phase of a static two-level atom immersed in a bath of fluctuating vacuum electromagnetic field in the background of a cosmic string. Our results indicate that due to the existence of the string, the geometric phase depends crucially on the position and the polarizability of the atom relative to the string. This can be ascribed to the fact that the presence of the string profoundly modify the distribution of electric field in Minkowski spacetime. So in principle, we can detect the cosmic string by experiments involving geometric phase.

  6. Current balancing for battery strings

    DOEpatents

    Galloway, James H.

    1985-01-01

    A battery plant is described which features magnetic circuit means for balancing the electrical current flow through a pluraliircuitbattery strings which are connected electrically in parallel. The magnetic circuit means is associated with the battery strings such that the conductors carrying the electrical current flow through each of the battery strings pass through the magnetic circuit means in directions which cause the electromagnetic fields of at least one predetermined pair of the conductors to oppose each other. In an alternative embodiment, a low voltage converter is associated with each of the battery strings for balancing the electrical current flow through the battery strings.

  7. Constraints on cosmic strings due to black holes formed from collapsed cosmic string loops

    NASA Technical Reports Server (NTRS)

    Caldwell, R. R.; Gates, Evalyn

    1993-01-01

    The cosmological features of primordial black holes formed from collapsed cosmic string loops are studied. Observational restrictions on a population of primordial black holes are used to restrict f, the fraction of cosmic string loops which collapse to form black holes, and mu, the cosmic string mass-per-unit length. Using a realistic model of cosmic strings, we find the strongest restriction on the parameters f and mu is due to the energy density in 100MeV photons radiated by the black holes. We also find that inert black hole remnants cannot serve as the dark matter. If earlier, crude estimates of f are reliable, our results severely restrict mu, and therefore limit the viability of the cosmic string large-scale structure scenario.

  8. Stochastic gravitational wave background from light cosmic strings

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    DePies, Matthew R.; Hogan, Craig J.

    2007-06-15

    Spectra of the stochastic gravitational wave backgrounds from cosmic strings are calculated and compared with present and future experimental limits. Motivated by theoretical expectations of light cosmic strings in superstring cosmology, improvements in experimental sensitivity, and recent demonstrations of large, stable loop formation from a primordial network, this study explores a new range of string parameters with masses lighter than previously investigated. A standard 'one-scale' model for string loop formation is assumed. Background spectra are calculated numerically for dimensionless string tensions G{mu}/c{sup 2} between 10{sup -7} and 10{sup -18}, and initial loop sizes as a fraction of the Hubble radiusmore » {alpha} from 0.1 to 10{sup -6}. The spectra show a low frequency power-law tail, a broad spectral peak due to loops decaying at the present epoch (including frequencies higher than their fundamental mode, and radiation associated with cusps), and a flat (constant energy density) spectrum at high frequencies due to radiation from loops that decayed during the radiation-dominated era. The string spectrum is distinctive and unlike any other known source. The peak of the spectrum for light strings appears at high frequencies, significantly affecting predicted signals. The spectra of the cosmic string backgrounds are compared with current millisecond pulsar limits and Laser Interferometer Space Antenna (LISA) sensitivity curves. For models with large stable loops ({alpha}=0.1), current pulsar-timing limits exclude G{mu}/c{sup 2}>10{sup -9}, a much tighter limit on string tension than achievable with other techniques, and within the range of current models based on brane inflation. LISA may detect a background from strings as light as G{mu}/c{sup 2}{approx_equal}10{sup -16}, corresponding to field theory strings formed at roughly 10{sup 11} GeV.« less

  9. CMB ISW-lensing bispectrum from cosmic strings

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Yamauchi, Daisuke; Sendouda, Yuuiti; Takahashi, Keitaro, E-mail: yamauchi@resceu.s.u-tokyo.ac.jp, E-mail: sendouda@cc.hirosaki-u.ac.jp, E-mail: keitaro@sci.kumamoto-u.ac.jp

    2014-02-01

    We study the effect of weak lensing by cosmic (super-)strings on the higher-order statistics of the cosmic microwave background (CMB). A cosmic string segment is expected to cause weak lensing as well as an integrated Sachs-Wolfe (ISW) effect, the so-called Gott-Kaiser-Stebbins (GKS) effect, to the CMB temperature fluctuation, which are thus naturally cross-correlated. We point out that, in the presence of such a correlation, yet another kind of the post-recombination CMB temperature bispectra, the ISW-lensing bispectra, will arise in the form of products of the auto- and cross-power spectra. We first present an analytic method to calculate the autocorrelation ofmore » the temperature fluctuations induced by the strings, and the cross-correlation between the temperature fluctuation and the lensing potential both due to the string network. In our formulation, the evolution of the string network is assumed to be characterized by the simple analytic model, the velocity-dependent one scale model, and the intercommutation probability is properly incorporated in order to characterize the possible superstringy nature. Furthermore, the obtained power spectra are dominated by the Poisson-distributed string segments, whose correlations are assumed to satisfy the simple relations. We then estimate the signal-to-noise ratios of the string-induced ISW-lensing bispectra and discuss the detectability of such CMB signals from the cosmic string network. It is found that in the case of the smaller string tension, Gμ << 10{sup -7}, the ISW-lensing bispectrum induced by a cosmic string network can constrain the string-model parameters even more tightly than the purely GKS-induced bispectrum in the ongoing and future CMB observations on small scales.« less

  10. CMB ISW-lensing bispectrum from cosmic strings

    NASA Astrophysics Data System (ADS)

    Yamauchi, Daisuke; Sendouda, Yuuiti; Takahashi, Keitaro

    2014-02-01

    We study the effect of weak lensing by cosmic (super-)strings on the higher-order statistics of the cosmic microwave background (CMB). A cosmic string segment is expected to cause weak lensing as well as an integrated Sachs-Wolfe (ISW) effect, the so-called Gott-Kaiser-Stebbins (GKS) effect, to the CMB temperature fluctuation, which are thus naturally cross-correlated. We point out that, in the presence of such a correlation, yet another kind of the post-recombination CMB temperature bispectra, the ISW-lensing bispectra, will arise in the form of products of the auto- and cross-power spectra. We first present an analytic method to calculate the autocorrelation of the temperature fluctuations induced by the strings, and the cross-correlation between the temperature fluctuation and the lensing potential both due to the string network. In our formulation, the evolution of the string network is assumed to be characterized by the simple analytic model, the velocity-dependent one scale model, and the intercommutation probability is properly incorporated in order to characterize the possible superstringy nature. Furthermore, the obtained power spectra are dominated by the Poisson-distributed string segments, whose correlations are assumed to satisfy the simple relations. We then estimate the signal-to-noise ratios of the string-induced ISW-lensing bispectra and discuss the detectability of such CMB signals from the cosmic string network. It is found that in the case of the smaller string tension, Gμ << 10-7, the ISW-lensing bispectrum induced by a cosmic string network can constrain the string-model parameters even more tightly than the purely GKS-induced bispectrum in the ongoing and future CMB observations on small scales.

  11. Rapid vegetative propagation method for carob

    USDA-ARS?s Scientific Manuscript database

    Many fruit species are propagated by vegetative methods such as budding, grafting, cutting, suckering, layering etc. to avoid heterozygosity. Carob trees (Ceratonia siliqua L.) are of highly economical value and it is among the most difficult-to-propagate fruit species. In this study, air-layering p...

  12. 7 CFR 29.3518 - Foreign matter.

    Code of Federal Regulations, 2014 CFR

    2014-01-01

    ... 7 Agriculture 2 2014-01-01 2014-01-01 false Foreign matter. 29.3518 Section 29.3518 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Type 95) § 29.3518 Foreign matter. Any extraneous substance or material such as stalks, suckers, straw...

  13. 7 CFR 29.3518 - Foreign matter.

    Code of Federal Regulations, 2011 CFR

    2011-01-01

    ... 7 Agriculture 2 2011-01-01 2011-01-01 false Foreign matter. 29.3518 Section 29.3518 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Type 95) § 29.3518 Foreign matter. Any extraneous substance or material such as stalks, suckers, straw...

  14. 7 CFR 29.3518 - Foreign matter.

    Code of Federal Regulations, 2013 CFR

    2013-01-01

    ... 7 Agriculture 2 2013-01-01 2013-01-01 false Foreign matter. 29.3518 Section 29.3518 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Type 95) § 29.3518 Foreign matter. Any extraneous substance or material such as stalks, suckers, straw...

  15. 7 CFR 29.3518 - Foreign matter.

    Code of Federal Regulations, 2012 CFR

    2012-01-01

    ... 7 Agriculture 2 2012-01-01 2012-01-01 false Foreign matter. 29.3518 Section 29.3518 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Type 95) § 29.3518 Foreign matter. Any extraneous substance or material such as stalks, suckers, straw...

  16. 7 CFR 29.3518 - Foreign matter.

    Code of Federal Regulations, 2010 CFR

    2010-01-01

    ... 7 Agriculture 2 2010-01-01 2010-01-01 false Foreign matter. 29.3518 Section 29.3518 Agriculture Regulations of the Department of Agriculture AGRICULTURAL MARKETING SERVICE (Standards, Inspections, Marketing... Type 95) § 29.3518 Foreign matter. Any extraneous substance or material such as stalks, suckers, straw...

  17. Lateral line pore diameters correlate with the development of gas bubble trauma signs in several Columbia River fishes

    USGS Publications Warehouse

    Morris, R.G.; Beeman, J.W.; VanderKooi, S.P.; Maule, A.G.

    2003-01-01

    Gas bubble trauma (GBT) caused by gas supersaturation of river water continues to be a problem in the Columbia River Basin. A common indicator of GBT is the percent of the lateral line occluded with gas bubbles; however, this effect has never been examined in relation to lateral line morphology. The effects of 115, 125 and 130% total dissolved gas levels were evaluated on five fish species common to the upper Columbia River. Trunk lateral line pore diameters differed significantly (P<0.0001) among species (longnose sucker>largescale sucker>northern pikeminnow≥chinook salmon≥redside shiner). At all supersaturation levels evaluated, percent of lateral line occlusion exhibited an inverse correlation to pore size but was not generally related to total dissolved gas level or time of exposure. This study suggests that the differences in lateral line pore diameters between species should be considered when using lateral line occlusion as an indicator of gas bubble trauma.

  18. Morphological and histological identification of Paramphistomum cervi (Trematoda: Paramiphistoma) in the rumen of infected sheep

    PubMed Central

    Chaoudhary, Vijayata; Hasnani, J. J.; Khyalia, Mukesh K.; Pandey, Sunanda; Chauhan, Vandip D.; Pandya, Suchit S.; Patel, P. V.

    2015-01-01

    Aim: This study was undertaken to identify Paramphistomum cervi on the basis of its morphology and histology to be the common cause of paramphistomosis in infected sheep and its differentiation from other similar Paramphistomes in Gujarat. Materials and Methods: Adult rumen flukes were recovered from the rumen of naturally infected sheep slaughtered in various abattoirs in Gujarat. Some adult flukes were flattened and stained in Borax carmine, and some were sectioned in the median sagittal plane and histological slides of the flukes were prepared for detailed morphological and histological studies. Result: Microscopic pictures of the parasite used in identification define the similarity in the morphology and histology of the anterior sucker, pharynx, esophagus, genital atrium, posterior sucker (acetabulum) and testes to the P. cervi. Conclusion: It can be concluded that the most common species found in sheep infected with Paramphistomosis is P. cervi on the basis of its histo-morphological appearance in Gujarat. PMID:27047009

  19. Determination of selenium in fish from designated critical habitat in the Gunnison River, Colorado, March through October, 2012

    USGS Publications Warehouse

    May, Thomas W.; Walther, Michael J.

    2013-01-01

    This report presents results for the summer 2012 sam-pling of muscle plugs from common carp (Cyprinus carpio), bonytail chub (Gila elegans), Colorado pikeminnow (Ptycho-cheilus lucius), and razorback suckers (Xyrauchen texanus) inhabiting critical habitat in the Gunnison River in western Colorado. Total selenium in fish muscle plugs was determinedby instrumental neutron activation analysis. Total selenium concentrations (range and mean ± standard deviation) in micrograms per gram dry weight were 6.0 to 10.7, 8.8 ± 1.3 for common carp; 2.9 to 8.7, 5.6 ± 2.4 for Colorado pikemin-now; and 1.4 to 7.3, 3.4 ± 2.7 for razorback sucker. The selenium concentration for one bonytail chub sample was 0.8 micrograms per gram dry weight. Selenium concentrations in muscle plugs from 1 Colorado pikeminnow and 12 common carp exceeded the 8 micrograms per gram dry weight toxicity guideline for selenium in fish muscle tissue.

  20. Assessing genetic diversity of wild and hatchery samples of the Chinese sucker (Myxocyprinus asiaticus) by the mitochondrial DNA control region.

    PubMed

    Wu, Jiayun; Wu, Bo; Hou, Feixia; Chen, Yongbai; Li, Chong; Song, Zhaobin

    2016-01-01

    To restore the natural populations of Chinese sucker (Myxocyprinus asiaticus), a hatchery release program has been underway for nearly 10 years. Using DNA sequences of the mitochondrial control region, we assessed the genetic diversity and genetic structure among samples collected from three sites of the wild population as well as from three hatcheries. The haplotype diversity of the wild samples (h = 0.899-0.975) was significantly higher than that of the hatchery ones (h = 0.296-0.666), but the nucleotide diversity was almost identical between them (π = 0.0170-0.0280). Relatively high gene flow was detected between the hatchery and wild samples. Analysis of effective population size indicated that M. asiaticus living in the Yangtze River has been expanding following a bottleneck in the recent past. Our results suggest the hatchery release programs for M. asiaticus have not reduced the genetic diversity, but have influenced the genetic structure of the species in the upper Yangtze River.

  1. Discovery of Maritrema jebuensis n. sp. (Digenea: Microphallidae) from the Asian Shore Crab, Hemigrapsus sanguineus, in Korea

    PubMed Central

    Chung, Ok-Sik; Lee, Hye-Jung; Sohn, Woon-Mok; Lee, Seung-Ha; Park, Il-Yong; Oh, Sang-Ah; Chai, Jong-Yil

    2010-01-01

    Maritrema spp. (Digenea: Microphallidae) are parasites of birds, but have not been found in the Republic of Korea. In this study, metacercariae of Maritrema sp. were discovered in the Asian shore crab, Hemigrapsus sanguineus, caught in the mud-flats of Jebu-do, Hwasung-gun, Gyeonggi-do, and the adult flukes were confirmed by experimental infection into mice. Based on the symmetric ribbon-like vitellarium, adult flukes of Maritrema sp. were identified, but did not belong to previously described species in terms of the following morphologic characteristics: ceca reaching to the lateral wall at the anterior border of the ovary; ventral sucker larger than oral sucker; a prominent metraterm; and vitellarium forming a complete ring. Hence, we named this microphallid M. jebuensis n. sp. after the island where the second intermediate hosts were collected. From this study, it has been shown that Maritrema sp. is distributed in Korea and transmitted by the Asian shore crab, H. sanguineus. PMID:21234238

  2. Discovery of Maritrema jebuensis n. sp. (Digenea: Microphallidae) from the Asian Shore Crab, Hemigrapsus sanguineus, in Korea.

    PubMed

    Chung, Ok-Sik; Lee, Hye-Jung; Sohn, Woon-Mok; Lee, Seung-Ha; Park, Il-Yong; Oh, Sang-Ah; Chai, Jong-Yil; Seo, Min

    2010-12-01

    Maritrema spp. (Digenea: Microphallidae) are parasites of birds, but have not been found in the Republic of Korea. In this study, metacercariae of Maritrema sp. were discovered in the Asian shore crab, Hemigrapsus sanguineus, caught in the mud-flats of Jebu-do, Hwasung-gun, Gyeonggi-do, and the adult flukes were confirmed by experimental infection into mice. Based on the symmetric ribbon-like vitellarium, adult flukes of Maritrema sp. were identified, but did not belong to previously described species in terms of the following morphologic characteristics: ceca reaching to the lateral wall at the anterior border of the ovary; ventral sucker larger than oral sucker; a prominent metraterm; and vitellarium forming a complete ring. Hence, we named this microphallid M. jebuensis n. sp. after the island where the second intermediate hosts were collected. From this study, it has been shown that Maritrema sp. is distributed in Korea and transmitted by the Asian shore crab, H. sanguineus.

  3. Ghost vertices for the bosonic string using the group-theoretic approach to string theory

    NASA Astrophysics Data System (ADS)

    Freeman, M. D.; West, P.

    1988-04-01

    The N-string tree-level scattering vertices for the bosonic string are extended to include anticommuting (ghost) oscillators. These vertices behave correctly under the action of the BRST charge Q and reproduce the known results for the scattering of physical states. This work is an application of the group-theoretic approach to string theory. Permanent address: Mathematics Department, King's College, Strand, London WC2R 2LS, UK.

  4. Cosmic strings and galaxy formation

    NASA Technical Reports Server (NTRS)

    Bertschinger, Edmund

    1989-01-01

    The cosmogonical model proposed by Zel'dovich and Vilenkin (1981), in which superconducting cosmic strings act as seeds for the origin of structure in the universe, is discussed, summarizing the results of recent theoretical investigations. Consideration is given to the formation of cosmic strings, the microscopic structure of strings, gravitational effects, cosmic string evolution, and the formation of galaxies and large-scale structure. Simulation results are presented in graphs, and several outstanding issues are listed and briefly characterized.

  5. String Things.

    ERIC Educational Resources Information Center

    Mesa Public Schools, AZ.

    Designed for music educators instructing grades 4 through 8 in string instruments, this Mesa (Arizona) public schools guide presents information on the string curriculum, orchestras, and practicing. The goals and objectives for string instruments delineate grade levels and how student skills will be verified. Following 17 curriculum goal tests,…

  6. Embellished String Prints. Cover Story.

    ERIC Educational Resources Information Center

    Smith, Mary Ruth

    1999-01-01

    Focuses on a printmaking activity in which students create embellished string prints using the relief process of string glued to chip board. Explains that string prints can easily be embellished with oil pastels. Provides a description of the procedure and a list of materials and methods. (CMK)

  7. DOE Office of Scientific and Technical Information (OSTI.GOV)

    Marolf, Donald; Palmer, Belkis Cabrera; Physics Department, Syracuse University, Syracuse, New York 13244

    A thermodynamic argument is presented suggesting that near-extremal spinning D1-D5-P black strings become unstable when their angular momentum exceeds J{sub crit}=3Q{sub 1}Q{sub 5}/2{radical}(2). In contrast, the dimensionally reduced black holes are thermodynamically stable. The proposed instability involves a phase in which the spin angular momentum above J{sub crit} is transferred to gyration of the string in space, i.e., to orbital angular momentum of parts of the string about the mean location in space. Thus the string becomes a rotating helical coil. We note that an instability of this form would yield a counter-example to the Gubser-Mitra conjecture, which proposes amore » particular link between dynamic black string instabilities and the thermodynamics of black strings. There may also be other instabilities associated with radiation modes of various fields. Our arguments also apply to the D-brane bound states associated with these black strings in weakly coupled string theory.« less

  8. Semiclassical (qft) and Quantum (string) Rotating Black Holes and Their Evaporation:. New Results

    NASA Astrophysics Data System (ADS)

    Bouchareb, A.; Ramón Medrano, M.; Sánchez, N. G.

    Combination of both quantum field theory (QFT) and string theory in curved backgrounds in a consistent framework, the string analogue model, allows us to provide a full picture of the Kerr-Newman black hole and its evaporation going beyond the current picture. We compute the quantum emission cross-section of strings by a Kerr-Newman black hole (KNbh). It shows the black hole emission at the Hawking temperature Tsem in the early stage of evaporation and the new string emission featuring a Hagedorn transition into a string state of temperature Ts at the last stages. New bounds on J and Q emerge in the quantum string regime (besides the known ones of the classical/semiclassical QFT regime). The last state of evaporation of a semiclassical Kerr-Newman black hole with mass M > mPl, angular momentum J and charge Q is a string state of temperature Ts, string mass Ms, J = 0 and Q = 0, decaying as usual quantum strings do into all kinds of particles. (Naturally, in this framework, there is no loss of information, (there is no paradox at all).) We compute the string entropy Ss(m, j) from the microscopic string density of states of mass m and spin mode j, ρ(m, j). (Besides the Hagedorn transition at Ts) we find for high j (extremal string states j → m2α‧c), a new phase transition at a temperature Tsj = √ {j/hbar }Ts, higher than Ts. By precisely identifying the semiclassical and quantum (string) gravity regimes, we find a new formula for the Kerr black hole entropy Ssem(M, J), as a function of the usual Bekenstein-Hawking entropy S sem(0). For M ≫ mPl and J < GM2/c, S sem(0) is the leading term, but for high angular momentum, (nearly extremal case J = GM2/c), a gravitational phase transition operates and the whole entropy Ssem is drastically different from the Bekenstein-Hawking entropy S sem(0). This new extremal black hole transition occurs at a temperature Tsem J = (J/ℏ)Tsem, higher than the Hawking temperature Tsem.

  9. Light Z' in heterotic string standardlike models

    NASA Astrophysics Data System (ADS)

    Athanasopoulos, P.; Faraggi, A. E.; Mehta, V. M.

    2014-05-01

    The discovery of the Higgs boson at the LHC supports the hypothesis that the Standard Model provides an effective parametrization of all subatomic experimental data up to the Planck scale. String theory, which provides a viable perturbative approach to quantum gravity, requires for its consistency the existence of additional gauge symmetries beyond the Standard Model. The construction of heterotic string models with a viable light Z' is, however, highly constrained. We outline the construction of standardlike heterotic string models that allow for an additional Abelian gauge symmetry that may remain unbroken down to low scales. We present a string inspired model, consistent with the string constraints.

  10. Scaling properties of cosmic (super)string networks

    NASA Astrophysics Data System (ADS)

    Martins, C. J. A. P.

    2014-10-01

    I use a combination of state-of-the-art numerical simulations and analytic modelling to discuss the scaling properties of cosmic defect networks, including superstrings. Particular attention is given to the role of extra degrees of freedom in the evolution of these networks. Compared to the 'plain vanilla' case of Goto-Nambu strings, three such extensions play important but distinct roles in the network dynamics: the presence of charges/currents on the string worldsheet, the existence of junctions, and the possibility of a hierarchy of string tensions. I also comment on insights gained from studying simpler defect networks, including Goto-Nambu strings themselves, domain walls and semilocal strings.

  11. Evolution of cosmic string networks

    NASA Technical Reports Server (NTRS)

    Albrecht, Andreas; Turok, Neil

    1989-01-01

    A discussion of the evolution and observable consequences of a network of cosmic strings is given. A simple model for the evolution of the string network is presented, and related to the statistical mechanics of string networks. The model predicts the long string density throughout the history of the universe from a single parameter, which researchers calculate in radiation era simulations. The statistical mechanics arguments indicate a particular thermal form for the spectrum of loops chopped off the network. Detailed numerical simulations of string networks in expanding backgrounds are performed to test the model. Consequences for large scale structure, the microwave and gravity wave backgrounds, nucleosynthesis and gravitational lensing are calculated.

  12. Gödel universes in string theory

    NASA Astrophysics Data System (ADS)

    Barrow, John D.; Dabrowski, Mariusz P.

    1998-11-01

    We show that homogeneous Gödel spacetimes need not contain closed timelike curves in low-energy-effective string theories. We find exact solutions for the Gödel metric in string theory for the full O(α') action including both dilaton and axion fields. The results are valid for bosonic, heterotic and super-strings. To first order in the inverse string tension α', these solutions display a simple relation between the angular velocity of the Gödel universe, Ω, and the inverse string tension of the form α'=1/Ω2 in the absence of the axion field. The generalization of this relationship is also found when the axion field is present.

  13. A note on closed-string interactions a la witten

    NASA Astrophysics Data System (ADS)

    Romans, L. J.

    1987-08-01

    We consider the problem of formulating a field theory of interacting closed strings analogous to Witten's open-string field theory. Two natural candidates have been suggested for an off-shell three-string interaction vertex: one scheme involves a cyclic geometric overlap in spacetime, while the other is obtained by ``stuttering'' the Fock-space realization of the open-string vertex. We demonstrate that these two approaches are in fact equivalent, utilizing the operator formalism as developed to describe Witten's theory. Implications of this result for the construction of closed-string theories are briefly discussed. Address after August 1, 1987: Department of Physics, University of Southern California, Los Angeles, CA 90089, USA.

  14. Reconstruction of piano hammer force from string velocity.

    PubMed

    Chaigne, Antoine

    2016-11-01

    A method is presented for reconstructing piano hammer forces through appropriate filtering of the measured string velocity. The filter design is based on the analysis of the pulses generated by the hammer blow and propagating along the string. In the five lowest octaves, the hammer force is reconstructed by considering two waves only: the incoming wave from the hammer and its first reflection at the front end. For the higher notes, four- or eight-wave schemes must be considered. The theory is validated on simulated string velocities by comparing imposed and reconstructed forces. The simulations are based on a nonlinear damped stiff string model previously developed by Chabassier, Chaigne, and Joly [J. Acoust. Soc. Am. 134(1), 648-665 (2013)]. The influence of absorption, dispersion, and amplitude of the string waves on the quality of the reconstruction is discussed. Finally, the method is applied to real piano strings. The measured string velocity is compared to the simulated velocity excited by the reconstructed force, showing a high degree of accuracy. A number of simulations are compared to simulated strings excited by a force derived from measurements of mass and acceleration of the hammer head. One application to an historic piano is also presented.

  15. Improving Upon String Methods for Transition State Discovery.

    PubMed

    Chaffey-Millar, Hugh; Nikodem, Astrid; Matveev, Alexei V; Krüger, Sven; Rösch, Notker

    2012-02-14

    Transition state discovery via application of string methods has been researched on two fronts. The first front involves development of a new string method, named the Searching String method, while the second one aims at estimating transition states from a discretized reaction path. The Searching String method has been benchmarked against a number of previously existing string methods and the Nudged Elastic Band method. The developed methods have led to a reduction in the number of gradient calls required to optimize a transition state, as compared to existing methods. The Searching String method reported here places new beads on a reaction pathway at the midpoint between existing beads, such that the resolution of the path discretization in the region containing the transition state grows exponentially with the number of beads. This approach leads to favorable convergence behavior and generates more accurate estimates of transition states from which convergence to the final transition states occurs more readily. Several techniques for generating improved estimates of transition states from a converged string or nudged elastic band have been developed and benchmarked on 13 chemical test cases. Optimization approaches for string methods, and pitfalls therein, are discussed.

  16. Social Impacts Module (SIM) Transition

    DTIC Science & Technology

    2012-09-28

    User String The authorized user’s name to access the PAVE database. Applies only to Microsoft SQL Server; leave blank, otherwise. passwd String The...otherwise. passwd String The password if an authorized user’s name is required; otherwise, leave blank driver String The class name for the driver to

  17. Novel string field theory with also negative energy constituents/objects gives Veneziano amplitude

    NASA Astrophysics Data System (ADS)

    Nielsen, H. B.; Ninomiya, M.

    2018-02-01

    We have proposed a new type of string field theory. The main point of the present article is to cure some technical troubles: missing two out three terms in Veneziano amplitude. Our novel string field theory, describes a theory with many strings in terms of "objects", which are not exactly, but close to Charles Thorn's string bits. The new point is that the objects in terms of which the universe states are constructed, and which have an essentially 26-momentum variable called J μ , can have the energy J 0 be also negative as well as positive. We get a long way in deriving in this model the Veneziano model and obtain all the three terms needed for a four point amplitude. This result strongly indicates that our novel string field theory is indeed string theory.

  18. Axion string dynamics I: 2+1D

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Fleury, Leesa M.; Moore, Guy D.

    2016-05-03

    If the axion exists and if the initial axion field value is uncorrelated at causally disconnected points, then it should be possible to predict the efficiency of cosmological axion production, relating the axionic dark matter density to the axion mass. The main obstacle to making this prediction is correctly treating the axion string cores. We develop a new algorithm for treating the axionic string cores correctly in 2+1 dimensions. When the axionic string cores are given their full physical string tension, axion production is about twice as efficient as in previous simulations. We argue that the string network in 2+1more » dimensions should behave very differently than in 3+1 dimensions, so this result cannot be simply carried over to the physical case. We outline how to extend our method to 3+1D axion string dynamics.« less

  19. A numerical study of the string function using a primitive equation ocean model

    NASA Astrophysics Data System (ADS)

    Tyler, R. H.; Käse, R.

    We use results from a primitive-equation ocean numerical model (SCRUM) to test a theoretical 'string function' formulation put forward by Tyler and Käse in another article in this issue. The string function acts as a stream function for the large-scale potential energy flow under the combined beta and topographic effects. The model results verify that large-scale anomalies propagate along the string function contours with a speed correctly given by the cross-string gradient. For anomalies having a scale similar to the Rossby radius, material rates of change in the layer mass following the string velocity are balanced by material rates of change in relative vorticity following the flow velocity. It is shown that large-amplitude anomalies can be generated when wind stress is resonant with the string function configuration.

  20. Scattering of Cosmic Strings by Black Holes:. Loop Formation

    NASA Astrophysics Data System (ADS)

    Dubath, Florian; Sakellariadou, Mairi; Viallet, Claude Michel

    We study the deformation of a long cosmic string by a nearby rotating black hole. We examine whether the deformation of a cosmic string, induced by the gravitational field of a Kerr black hole, may lead to the formation of a string loop. The segment of the string which enters the ergo-sphere of a rotating black hole gets deformed and, if it is sufficiently twisted, it can self-intersect, chopping off a loop. We find that the formation of a loop, via such a mechanism, is a rare event. It will only arise in a small region of the collision phase space, which depends on the string velocity, the impact parameter and the black hole angular momentum. We conclude that, generically, a long cosmic string is simply scattered, or captured, by a nearby rotating black hole.

  1. Factorization of chiral string amplitudes

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Huang, Yu-tin; Siegel, Warren; Yuan, Ellis Ye

    We re-examine a closed-string model defined by altering the boundary conditions for one handedness of two-dimensional propagators in otherwise-standard string theory. We evaluate the amplitudes using Kawai-Lewellen-Tye factorization into open-string amplitudes. The only modification to standard string theory is effectively that the spacetime Minkowski metric changes overall sign in one open-string factor. This cancels all but a finite number of states: as found in earlier approaches, with enough supersymmetry (e.g., type II) the tree amplitudes reproduce those of the massless truncation of ordinary string theory. However, we now find for the other cases that additional fields, formerly thought to bemore » auxiliary, describe new spin-2 states at the two adjacent mass levels (tachyonic and tardyonic). The tachyon is always a ghost, but can be avoided in the heterotic case.« less

  2. Factorization of chiral string amplitudes

    DOE PAGES

    Huang, Yu-tin; Siegel, Warren; Yuan, Ellis Ye

    2016-09-16

    We re-examine a closed-string model defined by altering the boundary conditions for one handedness of two-dimensional propagators in otherwise-standard string theory. We evaluate the amplitudes using Kawai-Lewellen-Tye factorization into open-string amplitudes. The only modification to standard string theory is effectively that the spacetime Minkowski metric changes overall sign in one open-string factor. This cancels all but a finite number of states: as found in earlier approaches, with enough supersymmetry (e.g., type II) the tree amplitudes reproduce those of the massless truncation of ordinary string theory. However, we now find for the other cases that additional fields, formerly thought to bemore » auxiliary, describe new spin-2 states at the two adjacent mass levels (tachyonic and tardyonic). The tachyon is always a ghost, but can be avoided in the heterotic case.« less

  3. Pauses enhance chunk recognition in song element strings by zebra finches.

    PubMed

    Spierings, Michelle; de Weger, Anouk; Ten Cate, Carel

    2015-07-01

    When learning a language, it is crucial to know which syllables of a continuous sound string belong together as words. Human infants achieve this by attending to pauses between words or to the co-occurrence of syllables. It is not only humans that can segment a continuous string. Songbirds learning their song tend to copy 'chunks' from one or more tutors' songs and combine these into their own song. In the tutor songs, these chunks are often separated by pauses and a high co-occurrence of elements, suggesting that these features affect chunking and song learning. We examined experimentally whether the presence of pauses and element co-occurrence affect the ability of adult zebra finches to discriminate strings of song elements. Using a go/no-go design, two groups of birds were trained to discriminate between two strings. In one group (Pause-group), pauses were inserted between co-occurring element triplets in the strings, and in the other group (No-pause group), both strings were continuous. After making a correct discrimination, an individual proceeded to a reversal training using string segments. Segments were element triplets consistent in co-occurrence, triplets that were partly consistent in composition and triplets consisting of elements that did not co-occur in the strings. The Pause-group was faster in discriminating between the two strings. This group also responded differently to consistent triplets in the reversal training, compared to inconsistent triplets. The No-pause group did not differentiate among the triplet types. These results indicate that pauses in strings of song elements aid song discrimination and memorization of co-occurring element groups.

  4. From the currency rate quotations onto strings and brane world scenarios

    NASA Astrophysics Data System (ADS)

    Horváth, D.; Pincak, R.

    2012-11-01

    In the paper, we study the projections of the real exchange rate dynamics onto the string-like topology. Our approach is inspired by the contemporary movements in the string theory. The string map of data is defined here by the boundary conditions, characteristic length, real valued and the method of redistribution of information. As a practical matter, this map represents the detrending and data standardization procedure. We introduced maps onto 1-end-point and 2-end-point open strings that satisfy the Dirichlet and Neumann boundary conditions. The questions of the choice of extra-dimensions, symmetries, duality and ways to the partial compactification are discussed. Subsequently, we pass to higher dimensional and more complex objects. The 2D-Brane was suggested which incorporated bid-ask spreads. Polarization by the spread was considered which admitted analyzing arbitrage opportunities on the market where transaction costs are taken into account. The model of the rotating string which naturally yields calculation of angular momentum is suitable for tracking of several currency pairs. The systematic way which allows one suggest more structured maps suitable for a simultaneous study of several currency pairs was analyzed by means of the Gâteaux generalized differential calculus. The effect of the string and brane maps on test data was studied by comparing their mean statistical characteristics. The study revealed notable differences between topologies. We review the dependence on the characteristic string length, mean fluctuations and properties of the intra-string statistics. The study explores the coupling of the string amplitude and volatility. The possible utilizations of the string theory approach in financial markets are slight.

  5. Haptic Distal Spatial Perception Mediated by Strings: Haptic "Looming"

    ERIC Educational Resources Information Center

    Cabe, Patrick A.

    2011-01-01

    Five experiments tested a haptic analog of optical looming, demonstrating string-mediated haptic distal spatial perception. Horizontally collinear hooks supported a weighted string held taut by a blindfolded participant's finger midway between the hooks. At the finger, the angle between string segments increased as the finger approached…

  6. NC-140 peach rootstock testing in 13 U.S. states

    USDA-ARS?s Scientific Manuscript database

    Eighteen Prunus rootstocks budded with ‘Redhaven’ peach were planted at 16 locations in North America in 2009. Fourth-year performance from 14 locations show that significant differences among rootstocks and sites were found for survival, root suckers, growth, bloom date, fruit maturity date, fruit ...

  7. Tilted string cosmologies

    NASA Astrophysics Data System (ADS)

    Clancy, Dominic; Feinstein, Alexander; Lidsey, James E.; Tavakol, Reza

    1999-04-01

    Global symmetries of the string effective action are employed to generate tilted, homogeneous Bianchi type VIh string cosmologies from a previously known stiff perfect fluid solution to Einstein gravity. The dilaton field is not constant on the surfaces of homogeneity. The future asymptotic state of the models is interpreted as a plane wave and is itself an exact solution to the string equations of motion to all orders in the inverse string tension. An inhomogeneous generalization of the Bianchi type III model is also found.

  8. Propagating stress-pulses and wiggling transition revealed in string dynamics

    NASA Astrophysics Data System (ADS)

    Yao, Zhenwei

    2018-02-01

    Understanding string dynamics yields insights into the intricate dynamic behaviors of various filamentary thin structures in nature and industry covering multiple length scales. In this work, we investigate the planar dynamics of a flexible string where one end is free and the other end is subject to transverse and longitudinal motions. Under transverse harmonic motion, we reveal the propagating pulse structure in the stress profile over the string, and analyze its role in bringing the system into a chaotic state. For a string where one end is under longitudinal uniform acceleration, we identify the wiggling transition, derive the analytical wiggling solution from the string equations, and present the phase diagram.

  9. Noncommutative-geometry model for closed bosonic strings

    NASA Technical Reports Server (NTRS)

    Sen, Siddhartha; Holman, R.

    1987-01-01

    It is shown how Witten's (1986) noncommutative geometry may be extended to describe the closed bosonic string. For closed strings, an explicit representation is provided of the integral operator needed to construct an action and of an associative product on string fields. The proper choice of the action of the integral operator and the associative product in order to give rise to a reasonable theory is explained, and the consequences of such a choice are discussed. It is shown that the ghost numbers of the operator and associative product can be chosen arbitrarily for both open and closed strings, and that this construct can be used as an action for interacting closed bosonic strings.

  10. Surface operators from M -strings

    NASA Astrophysics Data System (ADS)

    Mori, Hironori; Sugimoto, Yuji

    2017-01-01

    It has been found that surface operators have a significant role in Alday-Gaiotto-Tachikawa (AGT) relation. This duality is an outstanding consequence of M -theory, but it is actually encoded into the brane web for which the topological string can work. From this viewpoint, the surface defect in AGT relation is geometrically engineered as a toric brane realization. Also, there is a class of the brane configuration in M -theory called M -strings which can be translated into the language of the topological string. In this work, we propose a new M -string configuration which can realize AGT relation in the presence of the surface defect by utilizing the geometric transition in the refined topological string.

  11. Voltage-Matched, Monolithic, Multi-Band-Gap Devices

    DOEpatents

    Wanlass, M. W.; Mascarenhas, A.

    2006-08-22

    Monolithic, tandem, photonic cells include at least a first semiconductor layer and a second semiconductor layer, wherein each semiconductor layer includes an n-type region, a p-type region, and a given band-gap energy. Formed within each semiconductor layer is a string of electrically connected photonic sub-cells. By carefully selecting the numbers of photonic sub-cells in the first and second layer photonic sub-cell string(s), and by carefully selecting the manner in which the sub-cells in a first and second layer photonic sub-cell string(s) are electrically connected, each of the first and second layer sub-cell strings may be made to achieve one or more substantially identical electrical characteristics.

  12. Non-perturbative String Theory from Water Waves

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Iyer, Ramakrishnan; Johnson, Clifford V.; /Southern California U.

    2012-06-14

    We use a combination of a 't Hooft limit and numerical methods to find non-perturbative solutions of exactly solvable string theories, showing that perturbative solutions in different asymptotic regimes are connected by smooth interpolating functions. Our earlier perturbative work showed that a large class of minimal string theories arise as special limits of a Painleve IV hierarchy of string equations that can be derived by a similarity reduction of the dispersive water wave hierarchy of differential equations. The hierarchy of string equations contains new perturbative solutions, some of which were conjectured to be the type IIA and IIB string theoriesmore » coupled to (4, 4k ? 2) superconformal minimal models of type (A, D). Our present paper shows that these new theories have smooth non-perturbative extensions. We also find evidence for putative new string theories that were not apparent in the perturbative analysis.« less

  13. The Neural Substrates for Letter String Readings in The Normal and Reverse Directions: An fMRI Study

    NASA Astrophysics Data System (ADS)

    Ge, Sheng; Saito, Takashi; Wu, Jing-Long; Ogasawara, Jun-Ichi; Yamauchi, Shuichi; Matsunaga, Naofumi; Iramina, Keiji

    In order to investigate the difference in cortical activations between reading letter strings in the normal direction and the reverse direction, an fMRI study was conducted. In this study, the cortical activations elicited by Japanese letter string reading and Chinese letter string reading were investigated. The subjects performed the normal direction reading task (read letter strings from left to right), and the reverse direction reading task (read letter strings from right to left). According to the experimental results, the activated brain regions during the normal and the reverse direction reading tasks were compared. It was found that visuospatial transformation was involved in the reverse direction reading task, while this function was not significant during the normal direction reading task. Furthermore, we found that there was no significant difference in cortical activation between Japanese and Chinese letter string readings.

  14. Constraints on Cosmic Strings from the LIGO-Virgo Gravitational-Wave Detectors

    NASA Astrophysics Data System (ADS)

    Aasi, J.; Abadie, J.; Abbott, B. P.; Abbott, R.; Abbott, T.; Abernathy, M. R.; Accadia, T.; Acernese, F.; Adams, C.; Adams, T.; Adhikari, R. X.; Affeldt, C.; Agathos, M.; Aggarwal, N.; Aguiar, O. D.; Ajith, P.; Allen, B.; Allocca, A.; Amador Ceron, E.; Amariutei, D.; Anderson, R. A.; Anderson, S. B.; Anderson, W. G.; Arai, K.; Araya, M. C.; Arceneaux, C.; Areeda, J.; Ast, S.; Aston, S. M.; Astone, P.; Aufmuth, P.; Aulbert, C.; Austin, L.; Aylott, B. E.; Babak, S.; Baker, P. T.; Ballardin, G.; Ballmer, S. W.; Barayoga, J. C.; Barker, D.; Barnum, S. H.; Barone, F.; Barr, B.; Barsotti, L.; Barsuglia, M.; Barton, M. A.; Bartos, I.; Bassiri, R.; Basti, A.; Batch, J.; Bauchrowitz, J.; Bauer, Th. S.; Bebronne, M.; Behnke, B.; Bejger, M.; Beker, M. G.; Bell, A. S.; Bell, C.; Belopolski, I.; Bergmann, G.; Berliner, J. M.; Bersanetti, D.; Bertolini, A.; Bessis, D.; Betzwieser, J.; Beyersdorf, P. T.; Bhadbhade, T.; Bilenko, I. A.; Billingsley, G.; Birch, J.; Bitossi, M.; Bizouard, M. A.; Black, E.; Blackburn, J. K.; Blackburn, L.; Blair, D.; Blom, M.; Bock, O.; Bodiya, T. P.; Boer, M.; Bogan, C.; Bond, C.; Bondu, F.; Bonelli, L.; Bonnand, R.; Bork, R.; Born, M.; Boschi, V.; Bose, S.; Bosi, L.; Bowers, J.; Bradaschia, C.; Brady, P. R.; Braginsky, V. B.; Branchesi, M.; Brannen, C. A.; Brau, J. E.; Breyer, J.; Briant, T.; Bridges, D. O.; Brillet, A.; Brinkmann, M.; Brisson, V.; Britzger, M.; Brooks, A. F.; Brown, D. A.; Brown, D. D.; Brückner, F.; Bulik, T.; Bulten, H. J.; Buonanno, A.; Buskulic, D.; Buy, C.; Byer, R. L.; Cadonati, L.; Cagnoli, G.; Calderón Bustillo, J.; Calloni, E.; Camp, J. B.; Campsie, P.; Cannon, K. C.; Canuel, B.; Cao, J.; Capano, C. D.; Carbognani, F.; Carbone, L.; Caride, S.; Castiglia, A.; Caudill, S.; Cavaglià, M.; Cavalier, F.; Cavalieri, R.; Cella, G.; Cepeda, C.; Cesarini, E.; Chakraborty, R.; Chalermsongsak, T.; Chao, S.; Charlton, P.; Chassande-Mottin, E.; Chen, X.; Chen, Y.; Chincarini, A.; Chiummo, A.; Cho, H. S.; Chow, J.; Christensen, N.; Chu, Q.; Chua, S. S. Y.; Chung, S.; Ciani, G.; Clara, F.; Clark, D. E.; Clark, J. A.; Cleva, F.; Coccia, E.; Cohadon, P.-F.; Colla, A.; Colombini, M.; Constancio, M.; Conte, A.; Conte, R.; Cook, D.; Corbitt, T. R.; Cordier, M.; Cornish, N.; Corsi, A.; Costa, C. A.; Coughlin, M. W.; Coulon, J.-P.; Countryman, S.; Couvares, P.; Coward, D. M.; Cowart, M.; Coyne, D. C.; Craig, K.; Creighton, J. D. E.; Creighton, T. D.; Crowder, S. G.; Cumming, A.; Cunningham, L.; Cuoco, E.; Dahl, K.; Canton, T. Dal; Damjanic, M.; Danilishin, S. L.; D'Antonio, S.; Danzmann, K.; Dattilo, V.; Daudert, B.; Daveloza, H.; Davier, M.; Davies, G. S.; Daw, E. J.; Day, R.; Dayanga, T.; De Rosa, R.; Debreczeni, G.; Degallaix, J.; Del Pozzo, W.; Deleeuw, E.; Deléglise, S.; Denker, T.; Dent, T.; Dereli, H.; Dergachev, V.; DeRosa, R.; DeSalvo, R.; Dhurandhar, S.; Di Fiore, L.; Di Lieto, A.; Di Palma, I.; Di Virgilio, A.; Díaz, M.; Dietz, A.; Dmitry, K.; Donovan, F.; Dooley, K. L.; Doravari, S.; Drago, M.; Drever, R. W. P.; Driggers, J. C.; Du, Z.; Dumas, J.-C.; Dwyer, S.; Eberle, T.; Edwards, M.; Effler, A.; Ehrens, P.; Eichholz, J.; Eikenberry, S. S.; Endrőczi, G.; Essick, R.; Etzel, T.; Evans, K.; Evans, M.; Evans, T.; Factourovich, M.; Fafone, V.; Fairhurst, S.; Fang, Q.; Farinon, S.; Farr, B.; Farr, W.; Favata, M.; Fazi, D.; Fehrmann, H.; Feldbaum, D.; Ferrante, I.; Ferrini, F.; Fidecaro, F.; Finn, L. S.; Fiori, I.; Fisher, R.; Flaminio, R.; Foley, E.; Foley, S.; Forsi, E.; Fotopoulos, N.; Fournier, J.-D.; Franco, S.; Frasca, S.; Frasconi, F.; Frede, M.; Frei, M.; Frei, Z.; Freise, A.; Frey, R.; Fricke, T. T.; Fritschel, P.; Frolov, V. V.; Fujimoto, M.-K.; Fulda, P.; Fyffe, M.; Gair, J.; Gammaitoni, L.; Garcia, J.; Garufi, F.; Gehrels, N.; Gemme, G.; Genin, E.; Gennai, A.; Gergely, L.; Ghosh, S.; Giaime, J. A.; Giampanis, S.; Giardina, K. D.; Giazotto, A.; Gil-Casanova, S.; Gill, C.; Gleason, J.; Goetz, E.; Goetz, R.; Gondan, L.; González, G.; Gordon, N.; Gorodetsky, M. L.; Gossan, S.; Goßler, S.; Gouaty, R.; Graef, C.; Graff, P. B.; Granata, M.; Grant, A.; Gras, S.; Gray, C.; Greenhalgh, R. J. S.; Gretarsson, A. M.; Griffo, C.; Groot, P.; Grote, H.; Grover, K.; Grunewald, S.; Guidi, G. M.; Guido, C.; Gushwa, K. E.; Gustafson, E. K.; Gustafson, R.; Hall, B.; Hall, E.; Hammer, D.; Hammond, G.; Hanke, M.; Hanks, J.; Hanna, C.; Hanson, J.; Harms, J.; Harry, G. M.; Harry, I. W.; Harstad, E. D.; Hartman, M. T.; Haughian, K.; Hayama, K.; Heefner, J.; Heidmann, A.; Heintze, M.; Heitmann, H.; Hello, P.; Hemming, G.; Hendry, M.; Heng, I. S.; Heptonstall, A. W.; Heurs, M.; Hild, S.; Hoak, D.; Hodge, K. A.; Holt, K.; Holtrop, M.; Hong, T.; Hooper, S.; Horrom, T.; Hosken, D. J.; Hough, J.; Howell, E. J.; Hu, Y.; Hua, Z.; Huang, V.; Huerta, E. A.; Hughey, B.; Husa, S.; Huttner, S. H.; Huynh, M.; Huynh-Dinh, T.; Iafrate, J.; Ingram, D. R.; Inta, R.; Isogai, T.; Ivanov, A.; Iyer, B. R.; Izumi, K.; Jacobson, M.; James, E.; Jang, H.; Jang, Y. J.; Jaranowski, P.; Jiménez-Forteza, F.; Johnson, W. W.; Jones, D.; Jones, D. I.; Jones, R.; Jonker, R. J. G.; Ju, L.; Haris, K.; Kalmus, P.; Kalogera, V.; Kandhasamy, S.; Kang, G.; Kanner, J. B.; Kasprzack, M.; Kasturi, R.; Katsavounidis, E.; Katzman, W.; Kaufer, H.; Kaufman, K.; Kawabe, K.; Kawamura, S.; Kawazoe, F.; Kéfélian, F.; Keitel, D.; Kelley, D. B.; Kells, W.; Keppel, D. G.; Khalaidovski, A.; Khalili, F. Y.; Khazanov, E. A.; Kim, B. K.; Kim, C.; Kim, K.; Kim, N.; Kim, W.; Kim, Y.-M.; King, E. J.; King, P. J.; Kinzel, D. L.; Kissel, J. S.; Klimenko, S.; Kline, J.; Koehlenbeck, S.; Kokeyama, K.; Kondrashov, V.; Koranda, S.; Korth, W. Z.; Kowalska, I.; Kozak, D.; Kremin, A.; Kringel, V.; Królak, A.; Kucharczyk, C.; Kudla, S.; Kuehn, G.; Kumar, A.; Kumar, P.; Kumar, R.; Kurdyumov, R.; Kwee, P.; Landry, M.; Lantz, B.; Larson, S.; Lasky, P. D.; Lawrie, C.; Lazzarini, A.; Le Roux, A.; Leaci, P.; Lebigot, E. O.; Lee, C.-H.; Lee, H. K.; Lee, H. M.; Lee, J.; Lee, J.; Leonardi, M.; Leong, J. R.; Leroy, N.; Letendre, N.; Levine, B.; Lewis, J. B.; Lhuillier, V.; Li, T. G. F.; Lin, A. C.; Littenberg, T. B.; Litvine, V.; Liu, F.; Liu, H.; Liu, Y.; Liu, Z.; Lloyd, D.; Lockerbie, N. A.; Lockett, V.; Lodhia, D.; Loew, K.; Logue, J.; Lombardi, A. L.; Lorenzini, M.; Loriette, V.; Lormand, M.; Losurdo, G.; Lough, J.; Luan, J.; Lubinski, M. J.; Lück, H.; Lundgren, A. P.; Macarthur, J.; Macdonald, E.; Machenschalk, B.; MacInnis, M.; Macleod, D. M.; Magana-Sandoval, F.; Mageswaran, M.; Mailand, K.; Majorana, E.; Maksimovic, I.; Malvezzi, V.; Man, N.; Manca, G. M.; Mandel, I.; Mandic, V.; Mangano, V.; Mantovani, M.; Marchesoni, F.; Marion, F.; Márka, S.; Márka, Z.; Markosyan, A.; Maros, E.; Marque, J.; Martelli, F.; Martin, I. W.; Martin, R. M.; Martinelli, L.; Martynov, D.; Marx, J. N.; Mason, K.; Masserot, A.; Massinger, T. J.; Matichard, F.; Matone, L.; Matzner, R. A.; Mavalvala, N.; May, G.; Mazumder, N.; Mazzolo, G.; McCarthy, R.; McClelland, D. E.; McGuire, S. C.; McIntyre, G.; McIver, J.; Meacher, D.; Meadors, G. D.; Mehmet, M.; Meidam, J.; Meier, T.; Melatos, A.; Mendell, G.; Mercer, R. A.; Meshkov, S.; Messenger, C.; Meyer, M. S.; Miao, H.; Michel, C.; Mikhailov, E. E.; Milano, L.; Miller, J.; Minenkov, Y.; Mingarelli, C. M. F.; Mitra, S.; Mitrofanov, V. P.; Mitselmakher, G.; Mittleman, R.; Moe, B.; Mohan, M.; Mohapatra, S. R. P.; Mokler, F.; Moraru, D.; Moreno, G.; Morgado, N.; Mori, T.; Morriss, S. R.; Mossavi, K.; Mours, B.; Mow-Lowry, C. M.; Mueller, C. L.; Mueller, G.; Mukherjee, S.; Mullavey, A.; Munch, J.; Murphy, D.; Murray, P. G.; Mytidis, A.; Nagy, M. F.; Nanda Kumar, D.; Nardecchia, I.; Nash, T.; Naticchioni, L.; Nayak, R.; Necula, V.; Nelemans, G.; Neri, I.; Neri, M.; Newton, G.; Nguyen, T.; Nishida, E.; Nishizawa, A.; Nitz, A.; Nocera, F.; Nolting, D.; Normandin, M. E.; Nuttall, L. K.; Ochsner, E.; O'Dell, J.; Oelker, E.; Ogin, G. H.; Oh, J. J.; Oh, S. H.; Ohme, F.; Oppermann, P.; O'Reilly, B.; Ortega Larcher, W.; O'Shaughnessy, R.; Osthelder, C.; Ott, C. D.; Ottaway, D. J.; Ottens, R. S.; Ou, J.; Overmier, H.; Owen, B. J.; Padilla, C.; Pai, A.; Palomba, C.; Pan, Y.; Pankow, C.; Paoletti, F.; Paoletti, R.; Papa, M. A.; Paris, H.; Pasqualetti, A.; Passaquieti, R.; Passuello, D.; Pedraza, M.; Peiris, P.; Penn, S.; Perreca, A.; Phelps, M.; Pichot, M.; Pickenpack, M.; Piergiovanni, F.; Pierro, V.; Pinard, L.; Pindor, B.; Pinto, I. M.; Pitkin, M.; Poeld, J.; Poggiani, R.; Poole, V.; Poux, C.; Predoi, V.; Prestegard, T.; Price, L. R.; Prijatelj, M.; Principe, M.; Privitera, S.; Prix, R.; Prodi, G. A.; Prokhorov, L.; Puncken, O.; Punturo, M.; Puppo, P.; Quetschke, V.; Quintero, E.; Quitzow-James, R.; Raab, F. J.; Rabeling, D. S.; Rácz, I.; Radkins, H.; Raffai, P.; Raja, S.; Rajalakshmi, G.; Rakhmanov, M.; Ramet, C.; Rapagnani, P.; Raymond, V.; Re, V.; Reed, C. M.; Reed, T.; Regimbau, T.; Reid, S.; Reitze, D. H.; Ricci, F.; Riesen, R.; Riles, K.; Robertson, N. A.; Robinet, F.; Rocchi, A.; Roddy, S.; Rodriguez, C.; Rodruck, M.; Roever, C.; Rolland, L.; Rollins, J. G.; Romano, R.; Romanov, G.; Romie, J. H.; Rosińska, D.; Rowan, S.; Rüdiger, A.; Ruggi, P.; Ryan, K.; Salemi, F.; Sammut, L.; Sandberg, V.; Sanders, J.; Sannibale, V.; Santiago-Prieto, I.; Saracco, E.; Sassolas, B.; Sathyaprakash, B. S.; Saulson, P. R.; Savage, R.; Schilling, R.; Schnabel, R.; Schofield, R. M. S.; Schreiber, E.; Schuette, D.; Schulz, B.; Schutz, B. F.; Schwinberg, P.; Scott, J.; Scott, S. M.; Seifert, F.; Sellers, D.; Sengupta, A. S.; Sentenac, D.; Sergeev, A.; Shaddock, D.; Shah, S.; Shahriar, M. S.; Shaltev, M.; Shapiro, B.; Shawhan, P.; Shoemaker, D. H.; Sidery, T. L.; Siellez, K.; Siemens, X.; Sigg, D.; Simakov, D.; Singer, A.; Singer, L.; Sintes, A. M.; Skelton, G. R.; Slagmolen, B. J. J.; Slutsky, J.; Smith, J. R.; Smith, M. R.; Smith, R. J. E.; Smith-Lefebvre, N. D.; Soden, K.; Son, E. J.; Sorazu, B.; Souradeep, T.; Sperandio, L.; Staley, A.; Steinert, E.; Steinlechner, J.; Steinlechner, S.; Steplewski, S.; Stevens, D.; Stochino, A.; Stone, R.; Strain, K. A.; Straniero, N.; Strigin, S.; Stroeer, A. S.; Sturani, R.; Stuver, A. L.; Summerscales, T. Z.; Susmithan, S.; Sutton, P. J.; Swinkels, B.; Szeifert, G.; Tacca, M.; Talukder, D.; Tang, L.; Tanner, D. B.; Tarabrin, S. P.; Taylor, R.; ter Braack, A. P. M.; Thirugnanasambandam, M. P.; Thomas, M.; Thomas, P.; Thorne, K. A.; Thorne, K. S.; Thrane, E.; Tiwari, V.; Tokmakov, K. V.; Tomlinson, C.; Toncelli, A.; Tonelli, M.; Torre, O.; Torres, C. V.; Torrie, C. I.; Travasso, F.; Traylor, G.; Tse, M.; Ugolini, D.; Unnikrishnan, C. S.; Vahlbruch, H.; Vajente, G.; Vallisneri, M.; van den Brand, J. F. J.; Van Den Broeck, C.; van der Putten, S.; van der Sluys, M. V.; van Heijningen, J.; van Veggel, A. A.; Vass, S.; Vasúth, M.; Vaulin, R.; Vecchio, A.; Vedovato, G.; Veitch, J.; Veitch, P. J.; Venkateswara, K.; Verkindt, D.; Verma, S.; Vetrano, F.; Viceré, A.; Vincent-Finley, R.; Vinet, J.-Y.; Vitale, S.; Vlcek, B.; Vo, T.; Vocca, H.; Vorvick, C.; Vousden, W. D.; Vrinceanu, D.; Vyachanin, S. P.; Wade, A.; Wade, L.; Wade, M.; Waldman, S. J.; Walker, M.; Wallace, L.; Wan, Y.; Wang, J.; Wang, M.; Wang, X.; Wanner, A.; Ward, R. L.; Was, M.; Weaver, B.; Wei, L.-W.; Weinert, M.; Weinstein, A. J.; Weiss, R.; Welborn, T.; Wen, L.; Wessels, P.; West, M.; Westphal, T.; Wette, K.; Whelan, J. T.; Whitcomb, S. E.; White, D. J.; Whiting, B. F.; Wibowo, S.; Wiesner, K.; Wilkinson, C.; Williams, L.; Williams, R.; Williams, T.; Willis, J. L.; Willke, B.; Wimmer, M.; Winkelmann, L.; Winkler, W.; Wipf, C. C.; Wittel, H.; Woan, G.; Worden, J.; Yablon, J.; Yakushin, I.; Yamamoto, H.; Yancey, C. C.; Yang, H.; Yeaton-Massey, D.; Yoshida, S.; Yum, H.; Yvert, M.; ZadroŻny, A.; Zanolin, M.; Zendri, J.-P.; Zhang, F.; Zhang, L.; Zhao, C.; Zhu, H.; Zhu, X. J.; Zotov, N.; Zucker, M. E.; Zweizig, J.; LIGO Scientific Collaboration; Virgo Collaboration

    2014-04-01

    Cosmic strings can give rise to a large variety of interesting astrophysical phenomena. Among them, powerful bursts of gravitational waves (GWs) produced by cusps are a promising observational signature. In this Letter we present a search for GWs from cosmic string cusps in data collected by the LIGO and Virgo gravitational wave detectors between 2005 and 2010, with over 625 days of live time. We find no evidence of GW signals from cosmic strings. From this result, we derive new constraints on cosmic string parameters, which complement and improve existing limits from previous searches for a stochastic background of GWs from cosmic microwave background measurements and pulsar timing data. In particular, if the size of loops is given by the gravitational backreaction scale, we place upper limits on the string tension Gμ below 10-8 in some regions of the cosmic string parameter space.

  15. Constraints on Cosmic Strings from the LIGO-Virgo Gravitational-Wave Detectors

    NASA Technical Reports Server (NTRS)

    Aasi, J.; Abadie, J.; Abbott, B.P.; Abbott, R.; Abbott, T.; Abernathy, M.R.; Accadia, T.; Adams, C.; Adams, T.; Adhikari, R.X.; hide

    2014-01-01

    Cosmic strings can give rise to a large variety of interesting astrophysical phenomena. Among them, powerful bursts of gravitational waves (GWs) produced by cusps are a promising observational signature. In this Letter we present a search for GWs from cosmic string cusps in data collected by the LIGO and Virgo gravitational wave detectors between 2005 and 2010, with over 625 days of live time. We find no evidence of GW signals from cosmic strings. From this result, we derive new constraints on cosmic string parameters, which complement and improve existing limits from previous searches for a stochastic background of GWs from cosmic microwave background measurements and pulsar timing data. In particular, if the size of loops is given by the gravitational backreaction scale, we place upper limits on the string tension (Newton's Constant x mass per unit length) below 10(exp -8) in some regions of the cosmic string parameter space.

  16. Constraints on cosmic strings from the LIGO-Virgo gravitational-wave detectors.

    PubMed

    Aasi, J; Abadie, J; Abbott, B P; Abbott, R; Abbott, T; Abernathy, M R; Accadia, T; Acernese, F; Adams, C; Adams, T; Adhikari, R X; Affeldt, C; Agathos, M; Aggarwal, N; Aguiar, O D; Ajith, P; Allen, B; Allocca, A; Amador Ceron, E; Amariutei, D; Anderson, R A; Anderson, S B; Anderson, W G; Arai, K; Araya, M C; Arceneaux, C; Areeda, J; Ast, S; Aston, S M; Astone, P; Aufmuth, P; Aulbert, C; Austin, L; Aylott, B E; Babak, S; Baker, P T; Ballardin, G; Ballmer, S W; Barayoga, J C; Barker, D; Barnum, S H; Barone, F; Barr, B; Barsotti, L; Barsuglia, M; Barton, M A; Bartos, I; Bassiri, R; Basti, A; Batch, J; Bauchrowitz, J; Bauer, Th S; Bebronne, M; Behnke, B; Bejger, M; Beker, M G; Bell, A S; Bell, C; Belopolski, I; Bergmann, G; Berliner, J M; Bersanetti, D; Bertolini, A; Bessis, D; Betzwieser, J; Beyersdorf, P T; Bhadbhade, T; Bilenko, I A; Billingsley, G; Birch, J; Bitossi, M; Bizouard, M A; Black, E; Blackburn, J K; Blackburn, L; Blair, D; Blom, M; Bock, O; Bodiya, T P; Boer, M; Bogan, C; Bond, C; Bondu, F; Bonelli, L; Bonnand, R; Bork, R; Born, M; Boschi, V; Bose, S; Bosi, L; Bowers, J; Bradaschia, C; Brady, P R; Braginsky, V B; Branchesi, M; Brannen, C A; Brau, J E; Breyer, J; Briant, T; Bridges, D O; Brillet, A; Brinkmann, M; Brisson, V; Britzger, M; Brooks, A F; Brown, D A; Brown, D D; Brückner, F; Bulik, T; Bulten, H J; Buonanno, A; Buskulic, D; Buy, C; Byer, R L; Cadonati, L; Cagnoli, G; Calderón Bustillo, J; Calloni, E; Camp, J B; Campsie, P; Cannon, K C; Canuel, B; Cao, J; Capano, C D; Carbognani, F; Carbone, L; Caride, S; Castiglia, A; Caudill, S; Cavaglià, M; Cavalier, F; Cavalieri, R; Cella, G; Cepeda, C; Cesarini, E; Chakraborty, R; Chalermsongsak, T; Chao, S; Charlton, P; Chassande-Mottin, E; Chen, X; Chen, Y; Chincarini, A; Chiummo, A; Cho, H S; Chow, J; Christensen, N; Chu, Q; Chua, S S Y; Chung, S; Ciani, G; Clara, F; Clark, D E; Clark, J A; Cleva, F; Coccia, E; Cohadon, P-F; Colla, A; Colombini, M; Constancio, M; Conte, A; Conte, R; Cook, D; Corbitt, T R; Cordier, M; Cornish, N; Corsi, A; Costa, C A; Coughlin, M W; Coulon, J-P; Countryman, S; Couvares, P; Coward, D M; Cowart, M; Coyne, D C; Craig, K; Creighton, J D E; Creighton, T D; Crowder, S G; Cumming, A; Cunningham, L; Cuoco, E; Dahl, K; Dal Canton, T; Damjanic, M; Danilishin, S L; D'Antonio, S; Danzmann, K; Dattilo, V; Daudert, B; Daveloza, H; Davier, M; Davies, G S; Daw, E J; Day, R; Dayanga, T; De Rosa, R; Debreczeni, G; Degallaix, J; Del Pozzo, W; Deleeuw, E; Deléglise, S; Denker, T; Dent, T; Dereli, H; Dergachev, V; DeRosa, R; DeSalvo, R; Dhurandhar, S; Di Fiore, L; Di Lieto, A; Di Palma, I; Di Virgilio, A; Díaz, M; Dietz, A; Dmitry, K; Donovan, F; Dooley, K L; Doravari, S; Drago, M; Drever, R W P; Driggers, J C; Du, Z; Dumas, J-C; Dwyer, S; Eberle, T; Edwards, M; Effler, A; Ehrens, P; Eichholz, J; Eikenberry, S S; Endrőczi, G; Essick, R; Etzel, T; Evans, K; Evans, M; Evans, T; Factourovich, M; Fafone, V; Fairhurst, S; Fang, Q; Farinon, S; Farr, B; Farr, W; Favata, M; Fazi, D; Fehrmann, H; Feldbaum, D; Ferrante, I; Ferrini, F; Fidecaro, F; Finn, L S; Fiori, I; Fisher, R; Flaminio, R; Foley, E; Foley, S; Forsi, E; Fotopoulos, N; Fournier, J-D; Franco, S; Frasca, S; Frasconi, F; Frede, M; Frei, M; Frei, Z; Freise, A; Frey, R; Fricke, T T; Fritschel, P; Frolov, V V; Fujimoto, M-K; Fulda, P; Fyffe, M; Gair, J; Gammaitoni, L; Garcia, J; Garufi, F; Gehrels, N; Gemme, G; Genin, E; Gennai, A; Gergely, L; Ghosh, S; Giaime, J A; Giampanis, S; Giardina, K D; Giazotto, A; Gil-Casanova, S; Gill, C; Gleason, J; Goetz, E; Goetz, R; Gondan, L; González, G; Gordon, N; Gorodetsky, M L; Gossan, S; Goßler, S; Gouaty, R; Graef, C; Graff, P B; Granata, M; Grant, A; Gras, S; Gray, C; Greenhalgh, R J S; Gretarsson, A M; Griffo, C; Groot, P; Grote, H; Grover, K; Grunewald, S; Guidi, G M; Guido, C; Gushwa, K E; Gustafson, E K; Gustafson, R; Hall, B; Hall, E; Hammer, D; Hammond, G; Hanke, M; Hanks, J; Hanna, C; Hanson, J; Harms, J; Harry, G M; Harry, I W; Harstad, E D; Hartman, M T; Haughian, K; Hayama, K; Heefner, J; Heidmann, A; Heintze, M; Heitmann, H; Hello, P; Hemming, G; Hendry, M; Heng, I S; Heptonstall, A W; Heurs, M; Hild, S; Hoak, D; Hodge, K A; Holt, K; Holtrop, M; Hong, T; Hooper, S; Horrom, T; Hosken, D J; Hough, J; Howell, E J; Hu, Y; Hua, Z; Huang, V; Huerta, E A; Hughey, B; Husa, S; Huttner, S H; Huynh, M; Huynh-Dinh, T; Iafrate, J; Ingram, D R; Inta, R; Isogai, T; Ivanov, A; Iyer, B R; Izumi, K; Jacobson, M; James, E; Jang, H; Jang, Y J; Jaranowski, P; Jiménez-Forteza, F; Johnson, W W; Jones, D; Jones, D I; Jones, R; Jonker, R J G; Ju, L; K, Haris; Kalmus, P; Kalogera, V; Kandhasamy, S; Kang, G; Kanner, J B; Kasprzack, M; Kasturi, R; Katsavounidis, E; Katzman, W; Kaufer, H; Kaufman, K; Kawabe, K; Kawamura, S; Kawazoe, F; Kéfélian, F; Keitel, D; Kelley, D B; Kells, W; Keppel, D G; Khalaidovski, A; Khalili, F Y; Khazanov, E A; Kim, B K; Kim, C; Kim, K; Kim, N; Kim, W; Kim, Y-M; King, E J; King, P J; Kinzel, D L; Kissel, J S; Klimenko, S; Kline, J; Koehlenbeck, S; Kokeyama, K; Kondrashov, V; Koranda, S; Korth, W Z; Kowalska, I; Kozak, D; Kremin, A; Kringel, V; Królak, A; Kucharczyk, C; Kudla, S; Kuehn, G; Kumar, A; Kumar, P; Kumar, R; Kurdyumov, R; Kwee, P; Landry, M; Lantz, B; Larson, S; Lasky, P D; Lawrie, C; Lazzarini, A; Le Roux, A; Leaci, P; Lebigot, E O; Lee, C-H; Lee, H K; Lee, H M; Lee, J; Lee, J; Leonardi, M; Leong, J R; Leroy, N; Letendre, N; Levine, B; Lewis, J B; Lhuillier, V; Li, T G F; Lin, A C; Littenberg, T B; Litvine, V; Liu, F; Liu, H; Liu, Y; Liu, Z; Lloyd, D; Lockerbie, N A; Lockett, V; Lodhia, D; Loew, K; Logue, J; Lombardi, A L; Lorenzini, M; Loriette, V; Lormand, M; Losurdo, G; Lough, J; Luan, J; Lubinski, M J; Lück, H; Lundgren, A P; Macarthur, J; Macdonald, E; Machenschalk, B; MacInnis, M; Macleod, D M; Magana-Sandoval, F; Mageswaran, M; Mailand, K; Majorana, E; Maksimovic, I; Malvezzi, V; Man, N; Manca, G M; Mandel, I; Mandic, V; Mangano, V; Mantovani, M; Marchesoni, F; Marion, F; Márka, S; Márka, Z; Markosyan, A; Maros, E; Marque, J; Martelli, F; Martin, I W; Martin, R M; Martinelli, L; Martynov, D; Marx, J N; Mason, K; Masserot, A; Massinger, T J; Matichard, F; Matone, L; Matzner, R A; Mavalvala, N; May, G; Mazumder, N; Mazzolo, G; McCarthy, R; McClelland, D E; McGuire, S C; McIntyre, G; McIver, J; Meacher, D; Meadors, G D; Mehmet, M; Meidam, J; Meier, T; Melatos, A; Mendell, G; Mercer, R A; Meshkov, S; Messenger, C; Meyer, M S; Miao, H; Michel, C; Mikhailov, E E; Milano, L; Miller, J; Minenkov, Y; Mingarelli, C M F; Mitra, S; Mitrofanov, V P; Mitselmakher, G; Mittleman, R; Moe, B; Mohan, M; Mohapatra, S R P; Mokler, F; Moraru, D; Moreno, G; Morgado, N; Mori, T; Morriss, S R; Mossavi, K; Mours, B; Mow-Lowry, C M; Mueller, C L; Mueller, G; Mukherjee, S; Mullavey, A; Munch, J; Murphy, D; Murray, P G; Mytidis, A; Nagy, M F; Nanda Kumar, D; Nardecchia, I; Nash, T; Naticchioni, L; Nayak, R; Necula, V; Nelemans, G; Neri, I; Neri, M; Newton, G; Nguyen, T; Nishida, E; Nishizawa, A; Nitz, A; Nocera, F; Nolting, D; Normandin, M E; Nuttall, L K; Ochsner, E; O'Dell, J; Oelker, E; Ogin, G H; Oh, J J; Oh, S H; Ohme, F; Oppermann, P; O'Reilly, B; Ortega Larcher, W; O'Shaughnessy, R; Osthelder, C; Ott, C D; Ottaway, D J; Ottens, R S; Ou, J; Overmier, H; Owen, B J; Padilla, C; Pai, A; Palomba, C; Pan, Y; Pankow, C; Paoletti, F; Paoletti, R; Papa, M A; Paris, H; Pasqualetti, A; Passaquieti, R; Passuello, D; Pedraza, M; Peiris, P; Penn, S; Perreca, A; Phelps, M; Pichot, M; Pickenpack, M; Piergiovanni, F; Pierro, V; Pinard, L; Pindor, B; Pinto, I M; Pitkin, M; Poeld, J; Poggiani, R; Poole, V; Poux, C; Predoi, V; Prestegard, T; Price, L R; Prijatelj, M; Principe, M; Privitera, S; Prix, R; Prodi, G A; Prokhorov, L; Puncken, O; Punturo, M; Puppo, P; Quetschke, V; Quintero, E; Quitzow-James, R; Raab, F J; Rabeling, D S; Rácz, I; Radkins, H; Raffai, P; Raja, S; Rajalakshmi, G; Rakhmanov, M; Ramet, C; Rapagnani, P; Raymond, V; Re, V; Reed, C M; Reed, T; Regimbau, T; Reid, S; Reitze, D H; Ricci, F; Riesen, R; Riles, K; Robertson, N A; Robinet, F; Rocchi, A; Roddy, S; Rodriguez, C; Rodruck, M; Roever, C; Rolland, L; Rollins, J G; Romano, R; Romanov, G; Romie, J H; Rosińska, D; Rowan, S; Rüdiger, A; Ruggi, P; Ryan, K; Salemi, F; Sammut, L; Sandberg, V; Sanders, J; Sannibale, V; Santiago-Prieto, I; Saracco, E; Sassolas, B; Sathyaprakash, B S; Saulson, P R; Savage, R; Schilling, R; Schnabel, R; Schofield, R M S; Schreiber, E; Schuette, D; Schulz, B; Schutz, B F; Schwinberg, P; Scott, J; Scott, S M; Seifert, F; Sellers, D; Sengupta, A S; Sentenac, D; Sergeev, A; Shaddock, D; Shah, S; Shahriar, M S; Shaltev, M; Shapiro, B; Shawhan, P; Shoemaker, D H; Sidery, T L; Siellez, K; Siemens, X; Sigg, D; Simakov, D; Singer, A; Singer, L; Sintes, A M; Skelton, G R; Slagmolen, B J J; Slutsky, J; Smith, J R; Smith, M R; Smith, R J E; Smith-Lefebvre, N D; Soden, K; Son, E J; Sorazu, B; Souradeep, T; Sperandio, L; Staley, A; Steinert, E; Steinlechner, J; Steinlechner, S; Steplewski, S; Stevens, D; Stochino, A; Stone, R; Strain, K A; Straniero, N; Strigin, S; Stroeer, A S; Sturani, R; Stuver, A L; Summerscales, T Z; Susmithan, S; Sutton, P J; Swinkels, B; Szeifert, G; Tacca, M; Talukder, D; Tang, L; Tanner, D B; Tarabrin, S P; Taylor, R; ter Braack, A P M; Thirugnanasambandam, M P; Thomas, M; Thomas, P; Thorne, K A; Thorne, K S; Thrane, E; Tiwari, V; Tokmakov, K V; Tomlinson, C; Toncelli, A; Tonelli, M; Torre, O; Torres, C V; Torrie, C I; Travasso, F; Traylor, G; Tse, M; Ugolini, D; Unnikrishnan, C S; Vahlbruch, H; Vajente, G; Vallisneri, M; van den Brand, J F J; Van Den Broeck, C; van der Putten, S; van der Sluys, M V; van Heijningen, J; van Veggel, A A; Vass, S; Vasúth, M; Vaulin, R; Vecchio, A; Vedovato, G; Veitch, J; Veitch, P J; Venkateswara, K; Verkindt, D; Verma, S; Vetrano, F; Viceré, A; Vincent-Finley, R; Vinet, J-Y; Vitale, S; Vlcek, B; Vo, T; Vocca, H; Vorvick, C; Vousden, W D; Vrinceanu, D; Vyachanin, S P; Wade, A; Wade, L; Wade, M; Waldman, S J; Walker, M; Wallace, L; Wan, Y; Wang, J; Wang, M; Wang, X; Wanner, A; Ward, R L; Was, M; Weaver, B; Wei, L-W; Weinert, M; Weinstein, A J; Weiss, R; Welborn, T; Wen, L; Wessels, P; West, M; Westphal, T; Wette, K; Whelan, J T; Whitcomb, S E; White, D J; Whiting, B F; Wibowo, S; Wiesner, K; Wilkinson, C; Williams, L; Williams, R; Williams, T; Willis, J L; Willke, B; Wimmer, M; Winkelmann, L; Winkler, W; Wipf, C C; Wittel, H; Woan, G; Worden, J; Yablon, J; Yakushin, I; Yamamoto, H; Yancey, C C; Yang, H; Yeaton-Massey, D; Yoshida, S; Yum, H; Yvert, M; Zadrożny, A; Zanolin, M; Zendri, J-P; Zhang, F; Zhang, L; Zhao, C; Zhu, H; Zhu, X J; Zotov, N; Zucker, M E; Zweizig, J

    2014-04-04

    Cosmic strings can give rise to a large variety of interesting astrophysical phenomena. Among them, powerful bursts of gravitational waves (GWs) produced by cusps are a promising observational signature. In this Letter we present a search for GWs from cosmic string cusps in data collected by the LIGO and Virgo gravitational wave detectors between 2005 and 2010, with over 625 days of live time. We find no evidence of GW signals from cosmic strings. From this result, we derive new constraints on cosmic string parameters, which complement and improve existing limits from previous searches for a stochastic background of GWs from cosmic microwave background measurements and pulsar timing data. In particular, if the size of loops is given by the gravitational backreaction scale, we place upper limits on the string tension Gμ below 10(-8) in some regions of the cosmic string parameter space.

  17. Dualities in String Cosmology

    NASA Astrophysics Data System (ADS)

    Meissner, K. A.

    We describe in this chapter a set of duality symmetries present in the string-inspired theory of gravity coupled to the dilaton. These dualities are the cornerstones of String Cosmology, which provides alternatives to the usual inflation scenario. The crucial role of Prof. Gabriele Veneziano in the discovery and the development of string dualities is described and emphasized.

  18. Effects of overlapping strings in pp collisions

    DOE PAGES

    Bierlich, Christian; Gustafson, Gösta; Lönnblad, Leif; ...

    2015-03-26

    In models for hadron collisions based on string hadronization, the strings are usually treated as independent, allowing no interaction between the confined colour fields. In studies of nucleus collisions it has been suggested that strings close in space can fuse to form "colour ropes." Such ropes are expected to give more strange particles and baryons, which also has been suggested as a signal for plasma formation. Overlapping strings can also be expected in pp collisions, where usually no phase transition is expected. In particular at the high LHC energies the expected density of strings is quite high. To investigate possiblemore » effects of rope formation, we present a model in which strings are allowed to combine into higher multiplets, giving rise to increased production of baryons and strangeness, or recombine into singlet structures and vanish. Also a crude model for strings recombining into junction structures is considered, again giving rise to increased baryon production. The models are implemented in the DIPSY MC event generator, using PYTHIA8 for hadronization, and comparison to pp minimum bias data, reveals improvement in the description of identified particle spectra.« less

  19. Hydraulic drill string breakdown and bleed off unit

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Zeringue, F.J. Jr.

    1987-02-17

    An apparatus is described for use within an oil well rig for decoupling a tubing string into pipe segments comprising, in combination: rotary tong means for applying an unthreading torque to a first, upper pipe segment within the tubing string; torque resisting means for securing a second, lower pipe segment within the tubing string against the unthreading torque; containing means, intermediate the rotary tong means and the torque resisting means, enclosing a threaded joint of the tubing string, adapted for containing pressurized gases, liquids, and particulates, released from the threaded joint upon the decoupling; fluid communicating means for allowing fluidmore » communication between the containing means and a receiving point adapted for receiving the pressurized gases, liquids, and particulates; means for moving the rotary tong means, the torque resisting means and the containing means between a closed, engaging position with the tubing string and an open position; and means for horizontally moving the rotary tong means, the torque resisting means and the containing means between a position adjacent the tubing string and a position away from the tubing string.« less

  20. Perceiving the affordance of string tension for power strokes in badminton: expertise allows effective use of all string tensions.

    PubMed

    Zhu, Qin

    2013-01-01

    Affordances mean opportunities for action. These affordances are important for sports performance and relevant to the abilities developed by skilled athletes. In racquet sports such as badminton, different players prefer widely different string tension because it is believed to provide opportunities for effective strokes. The current study examined whether badminton players can perceive the affordance of string tension for power strokes and whether the perception of affordance itself changed as a function of skill level. The results showed that string tension constrained the striking performance of both novice and recreational players, but not of expert players. When perceptual capability was assessed, perceptual mode did not affect perception of the optimal string tension. Skilled players successfully perceived the affordance of string tension, but only experts were concerned about saving energy. Our findings demonstrated that perception of the affordance of string tension in badminton was determined by action abilities. Furthermore, experts could adjust the action to maintain a superior level of performance based on the perception of affordance.

  1. The three-dimensional simulation analysis of dynamic response on perforated strings

    NASA Astrophysics Data System (ADS)

    Li, M. F.; Liu, H. F.; Dou, Y. H.; Cao, L. H.; Liu, Y. X.

    2018-06-01

    It analyzes the dynamic response and stresses of perforating tubular string to detonating impact load in oil-gas well in ANSYS, obtains the response of vibration displacement, velocity and acceleration of perforating tubularstring caused by detonating impact load, finds the influence of the length and wall thickness of perforating tubular string to working stresses. The result shows that:when the detonating impact load exerts the perforating tubular string with compressive and tensile axial force alternatively;the vibration displacement, velocity and acceleration of perfora-ting tubular string change periodically at same cycle;the closer to the perforating gun, the larger the amplitude of vi-bration velocity and acceleration;the closer to the packer the smaller the vibration displacement, the larger the work-ing equivalent stress of perforating tubular string;the longer or the thicker the perforating tubular string, the smaller the working equivalent stress and the higher the strength safety. Therefore, it uses the damping tube between packer and perforating gun as well as thick walled tubing to increase the strength safety of perforating tubular string.

  2. Worldsheet factorization for twistor-strings

    NASA Astrophysics Data System (ADS)

    Adamo, Tim

    2014-04-01

    We study the multiparticle factorization properties of two worldsheet theories which — at tree-level — describe the scattering of massless particles in four dimensions: the Berkovits-Witten twistor-string for = 4 super-Yang-Mills coupled to = 4 conformal supergravity, and the Skinner twistor-string for = 8 supergravity. By considering these string-like theories, we can study factorization at the level of the worldsheet before any Wick contractions or integrals have been performed; this is much simpler than considering the factorization properties of the amplitudes themselves. In Skinner's twistor-string this entails the addition of worldsheet gravity as well as a formalism that represents all external states in a manifestly symmetric way, which we develop explicitly at genus zero. We confirm that the scattering amplitudes of Skinner's theory, as well as the gauge theory amplitudes for the planar sector of the Berkovits-Witten theory, factorize appropriately at genus zero. In the non-planar sector, we find behavior indicative of conformal gravity in the Berkovits-Witten twistor-string. We contrast factorization in twistor-strings with the story in ordinary string theory, and also make some remarks on higher genus factorization and disconnected prescriptions.

  3. The role of visual spatial attention in adult developmental dyslexia.

    PubMed

    Collis, Nathan L; Kohnen, Saskia; Kinoshita, Sachiko

    2013-01-01

    The present study investigated the nature of visual spatial attention deficits in adults with developmental dyslexia, using a partial report task with five-letter, digit, and symbol strings. Participants responded by a manual key press to one of nine alternatives, which included other characters in the string, allowing an assessment of position errors as well as intrusion errors. The results showed that the dyslexic adults performed significantly worse than age-matched controls with letter and digit strings but not with symbol strings. Both groups produced W-shaped serial position functions with letter and digit strings. The dyslexics' deficits with letter string stimuli were limited to position errors, specifically at the string-interior positions 2 and 4. These errors correlated with letter transposition reading errors (e.g., reading slat as "salt"), but not with the Rapid Automatized Naming (RAN) task. Overall, these results suggest that the dyslexic adults have a visual spatial attention deficit; however, the deficit does not reflect a reduced span in visual-spatial attention, but a deficit in processing a string of letters in parallel, probably due to difficulty in the coding of letter position.

  4. Self-similar motion of a Nambu-Goto string

    NASA Astrophysics Data System (ADS)

    Igata, Takahisa; Houri, Tsuyoshi; Harada, Tomohiro

    2016-09-01

    We study the self-similar motion of a string in a self-similar spacetime by introducing the concept of a self-similar string, which is defined as the world sheet to which a homothetic vector field is tangent. It is shown that in Nambu-Goto theory, the equations of motion for a self-similar string reduce to those for a particle. Moreover, under certain conditions such as the hypersurface orthogonality of the homothetic vector field, the equations of motion for a self-similar string simplify to the geodesic equations on a (pseudo)Riemannian space. As a concrete example, we investigate a self-similar Nambu-Goto string in a spatially flat Friedmann-Lemaître-Robertson-Walker expanding universe with self-similarity and obtain solutions of open and closed strings, which have various nontrivial configurations depending on the rate of the cosmic expansion. For instance, we obtain a circular solution that evolves linearly in the cosmic time while keeping its configuration by the balance between the effects of the cosmic expansion and string tension. We also show the instability for linear radial perturbation of the circular solutions.

  5. Wavelet-Bayesian inference of cosmic strings embedded in the cosmic microwave background

    NASA Astrophysics Data System (ADS)

    McEwen, J. D.; Feeney, S. M.; Peiris, H. V.; Wiaux, Y.; Ringeval, C.; Bouchet, F. R.

    2017-12-01

    Cosmic strings are a well-motivated extension to the standard cosmological model and could induce a subdominant component in the anisotropies of the cosmic microwave background (CMB), in addition to the standard inflationary component. The detection of strings, while observationally challenging, would provide a direct probe of physics at very high-energy scales. We develop a framework for cosmic string inference from observations of the CMB made over the celestial sphere, performing a Bayesian analysis in wavelet space where the string-induced CMB component has distinct statistical properties to the standard inflationary component. Our wavelet-Bayesian framework provides a principled approach to compute the posterior distribution of the string tension Gμ and the Bayesian evidence ratio comparing the string model to the standard inflationary model. Furthermore, we present a technique to recover an estimate of any string-induced CMB map embedded in observational data. Using Planck-like simulations, we demonstrate the application of our framework and evaluate its performance. The method is sensitive to Gμ ∼ 5 × 10-7 for Nambu-Goto string simulations that include an integrated Sachs-Wolfe contribution only and do not include any recombination effects, before any parameters of the analysis are optimized. The sensitivity of the method compares favourably with other techniques applied to the same simulations.

  6. A string theory which isn't about strings

    NASA Astrophysics Data System (ADS)

    Lee, Kanghoon; Rey, Soo-Jong; Rosabal, J. A.

    2017-11-01

    Quantization of closed string proceeds with a suitable choice of worldsheet vacuum. A priori, the vacuum may be chosen independently for left-moving and right-moving sectors. We construct ab initio quantized bosonic string theory with left-right asymmetric worldsheet vacuum and explore its consequences and implications. We critically examine the validity of new vacuum and carry out first-quantization using standard operator formalism. Remarkably, the string spectrum consists only of a finite number of degrees of freedom: string gravity (massless spin-two, Kalb-Ramond and dilaton fields) and two massive spin-two Fierz-Pauli fields. The massive spin-two fields have negative norm, opposite mass-squared, and provides a Lee-Wick type extension of string gravity. We compute two physical observables: tree-level scattering amplitudes and one-loop cosmological constant. Scattering amplitude of four dilatons is shown to be a rational function of kinematic invariants, and in D = 26 factorizes into contributions of massless spin-two and a pair of massive spin-two fields. The string one loop partition function is shown to perfectly agree with one loop Feynman diagram of string gravity and two massive spin-two fields. In particular, it does not exhibit modular invariance. We critically compare our construction with recent studies and contrast differences.

  7. Critical non-Abelian vortex in four dimensions and little string theory

    NASA Astrophysics Data System (ADS)

    Shifman, M.; Yung, A.

    2017-08-01

    As was shown recently, non-Abelian vortex strings supported in four-dimensional N =2 supersymmetric QCD with the U(2) gauge group and Nf=4 quark multiplets (flavors) become critical superstrings. In addition to the translational moduli, non-Abelian strings under consideration carry six orientational and size moduli. Together, they form a ten-dimensional target space required for a superstring to be critical. The target space of the string sigma model is a product of the flat four-dimensional space and a Calabi-Yau noncompact threefold, namely, the conifold. We study closed string states which emerge in four dimensions and identify them with hadrons of four-dimensional N =2 QCD. One massless state was found previously; it emerges as a massless hypermultiplet associated with the deformation of the complex structure of the conifold. In this paper, we find a number of massive states. To this end, we exploit the approach used in LST little string theory, namely, the equivalence between the critical string on the conifold and noncritical c =1 string with the Liouville field and a compact scalar at the self-dual radius. The states we find carry "baryonic" charge (its definition differs from standard). We interpret them as "monopole necklaces" formed (at strong coupling) by the closed string with confined monopoles attached.

  8. Patterned-string tasks: relation between fine motor skills and visual-spatial abilities in parrots.

    PubMed

    Krasheninnikova, Anastasia

    2013-01-01

    String-pulling and patterned-string tasks are often used to analyse perceptual and cognitive abilities in animals. In addition, the paradigm can be used to test the interrelation between visual-spatial and motor performance. Two Australian parrot species, the galah (Eolophus roseicapilla) and the cockatiel (Nymphicus hollandicus), forage on the ground, but only the galah uses its feet to manipulate food. I used a set of string pulling and patterned-string tasks to test whether usage of the feet during foraging is a prerequisite for solving the vertical string pulling problem. Indeed, the two species used techniques that clearly differed in the extent of beak-foot coordination but did not differ in terms of their success in solving the string pulling task. However, when the visual-spatial skills of the subjects were tested, the galahs outperformed the cockatiels. This supports the hypothesis that the fine motor skills needed for advanced beak-foot coordination may be interrelated with certain visual-spatial abilities needed for solving patterned-string tasks. This pattern was also found within each of the two species on the individual level: higher motor abilities positively correlated with performance in patterned-string tasks. This is the first evidence of an interrelation between visual-spatial and motor abilities in non-mammalian animals.

  9. Spontaneous knotting of an agitated string.

    PubMed

    Raymer, Dorian M; Smith, Douglas E

    2007-10-16

    It is well known that a jostled string tends to become knotted; yet the factors governing the "spontaneous" formation of various knots are unclear. We performed experiments in which a string was tumbled inside a box and found that complex knots often form within seconds. We used mathematical knot theory to analyze the knots. Above a critical string length, the probability P of knotting at first increased sharply with length but then saturated below 100%. This behavior differs from that of mathematical self-avoiding random walks, where P has been proven to approach 100%. Finite agitation time and jamming of the string due to its stiffness result in lower probability, but P approaches 100% with long, flexible strings. We analyzed the knots by calculating their Jones polynomials via computer analysis of digital photos of the string. Remarkably, almost all were identified as prime knots: 120 different types, having minimum crossing numbers up to 11, were observed in 3,415 trials. All prime knots with up to seven crossings were observed. The relative probability of forming a knot decreased exponentially with minimum crossing number and Möbius energy, mathematical measures of knot complexity. Based on the observation that long, stiff strings tend to form a coiled structure when confined, we propose a simple model to describe the knot formation based on random "braid moves" of the string end. Our model can qualitatively account for the observed distribution of knots and dependence on agitation time and string length.

  10. A method for the automated processing and analysis of images of ULVWF-platelet strings.

    PubMed

    Reeve, Scott R; Abbitt, Katherine B; Cruise, Thomas D; Hose, D Rodney; Lawford, Patricia V

    2013-01-01

    We present a method for identifying and analysing unusually large von Willebrand factor (ULVWF)-platelet strings in noisy low-quality images. The method requires relatively inexpensive, non-specialist equipment and allows multiple users to be employed in the capture of images. Images are subsequently enhanced and analysed, using custom-written software to perform the processing tasks. The formation and properties of ULVWF-platelet strings released in in vitro flow-based assays have recently become a popular research area. Endothelial cells are incorporated into a flow chamber, chemically stimulated to induce ULVWF release and perfused with isolated platelets which are able to bind to the ULVWF to form strings. The numbers and lengths of the strings released are related to characteristics of the flow. ULVWF-platelet strings are routinely identified by eye from video recordings captured during experiments and analysed manually using basic NIH image software to determine the number of strings and their lengths. This is a laborious, time-consuming task and a single experiment, often consisting of data from four to six dishes of endothelial cells, can take 2 or more days to analyse. The method described here allows analysis of the strings to provide data such as the number and length of strings, number of platelets per string and the distance between each platelet to be found. The software reduces analysis time, and more importantly removes user subjectivity, producing highly reproducible results with an error of less than 2% when compared with detailed manual analysis.

  11. Characterization of binary string statistics for syntactic landmine detection

    NASA Astrophysics Data System (ADS)

    Nasif, Ahmed O.; Mark, Brian L.; Hintz, Kenneth J.

    2011-06-01

    Syntactic landmine detection has been proposed to detect and classify non-metallic landmines using ground penetrating radar (GPR). In this approach, the GPR return is processed to extract characteristic binary strings for landmine and clutter discrimination. In our previous work, we discussed the preprocessing methodology by which the amplitude information of the GPR A-scan signal can be effectively converted into binary strings, which identify the impedance discontinuities in the signal. In this work, we study the statistical properties of the binary string space. In particular, we develop a Markov chain model to characterize the observed bit sequence of the binary strings. The state is defined as the number of consecutive zeros between two ones in the binarized A-scans. Since the strings are highly sparse (the number of zeros is much greater than the number of ones), defining the state this way leads to fewer number of states compared to the case where each bit is defined as a state. The number of total states is further reduced by quantizing the number of consecutive zeros. In order to identify the correct order of the Markov model, the mean square difference (MSD) between the transition matrices of mine strings and non-mine strings is calculated up to order four using training data. The results show that order one or two maximizes this MSD. The specification of the transition probabilities of the chain can be used to compute the likelihood of any given string. Such a model can be used to identify characteristic landmine strings during the training phase. These developments on modeling and characterizing the string statistics can potentially be part of a real-time landmine detection algorithm that identifies landmine and clutter in an adaptive fashion.

  12. CONAMINANT EXPOSURE, BIOCHEMICAL, AND HISTOPATHOLOGICAL BIOMARKERS IN WHITE SUCKERS FROM CONTAMINATED AND REFERENCE SITES IN THE SHEBOYGAN RIVER, WISCONSIN

    EPA Science Inventory

    Fish populations of the lower Sheboygan River, located in east-central Wisconsin, are considered impaired under the Great Lakes Water Qualtiy Agreement between the United States and Canada. Pollutants in the Sheboygan River system include: polychlorinated biphenyls (PCBs), polycy...

  13. CONTAMINANT EXPOSURE, BIOCHEMICAL, AND HISTOPATHOLOGICAL BIOMARKERS IN WHITE SUCKERS FROM CONTAMINATED AND REFERENCE SITES IN THE SHEBOYGAN RIVER, WISCONSIN

    EPA Science Inventory

    Fish populations of the lower Sheboygan River, located in east-central Wisconsin, are considered impaired under the Great Lakes Water Quality Agreement between the United States and Canada.Pollutants in the Sheboygan River system include: polychlorinated biphenyls (PCBs), polycyc...

  14. Managing Sandalwood for conservation in North Queensland, Australia

    Treesearch

    Grahame B. Applegate; Allan G. W. Davis; Peter Annable

    1990-01-01

    Santalum lanceolatum, the commercial species of sandalwood harvested in Queensland, was worth $4.2 million in export earnings in 1988. The ecology of the species in natural forests is summarized, and information on seedling regeneration and coppice and root suckering strategies is provided. Stand characteristics and size class distribution in two...

  15. ESTIMATION OF EXPOSURE CRITERIA VALUES FOR BILIARY POLYCYCLIC AROMATIC HYDROCARBON METABOLITE CONCENTRATIONS IN WHITE SUCKERS (CATOSTOMUS COMMERSONI)

    EPA Science Inventory

    A methodology was developed for deriving quantitative exposure criteria useful for comparing a site or a watershed to a reference condition and for defining the occurrence of extreme exposures. The prototype method used indicators of exposures to oil contamination and combustion ...

  16. ESTIMATION OF EXPOSURE CRITERIA VALUES FOR BILIARY POLYCYCLIC AROMATIC HYDROCARBON METABOLITE CONCENTRATION IN WHITE SUCKERS (CATASTOMUS COMMERSONI)

    EPA Science Inventory

    A methodology was developed for deriving quantitative exposure criteria useful for comparing a site or a watershed to a reference condition and for defining the occurrence of extreme exposures. The prototype method used indicators of exposures to oil contamination and combustion ...

  17. Regeneration and productivity of aspen grown on repeated short rotations.

    Treesearch

    Donald A. Perala

    1979-01-01

    Regeneration and productivity of aspen cannot be sustained on rotations shorter than about 8 years. Productivity losses on short rotations are physiological and morphological rather than nutritional in nature. Stump and root collar sprouts, which are rare in mature stands, were more numerous than root suckers.

  18. Manipulations to regenerate aspen ecosystems

    Treesearch

    Wayne D. Shepperd

    2001-01-01

    Vegetative regeneration of aspen can be initiated through manipulations that provide hormonal stimulation, proper growth environment, and sucker protection - the three elements of the aspen regeneration triangle. The correct course of action depends upon a careful evaluation of the size, vigor, age, and successional status of the existing clone. Soils and site...

  19. 75 FR 38441 - Endangered and Threatened Wildlife and Plants; Revised Critical Habitat for Santa Ana Sucker

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-07-02

    ... Ayres Suites Corona West, 1900 W Frontage Road, Corona, CA 92882. We will post all comments on http....m. and from 6 p.m. to 8 p.m. at Ayres Suites Corona West, 1900 W. Frontage Road, Corona, CA 92882...

  20. Fish population modeling approaches for assessing direct effects and recovery following mitigation of a pulp mill effluent in Jackfish Bay

    EPA Science Inventory

    We present an approach to link chemically-induced alterations in molecular and biochemical endpoints to adverse outcomes in whole organisms and populations. A predictive population model was developed to translate changes in fecundity measures of white sucker (Catostomus commers...

  1. Vegetative Propagation of Aspen by Greenwood Cuttings

    Treesearch

    Robert E. Farmer

    1963-01-01

    Greenwood stem cuttings taken from root suckers of Populus tremuloides and P. grandidentata were rooted in coarse sand under both intermittent mist and polyethylene sheeting in a greenhouse and under mist in an outdoor propagation bed. Prerooting treatment of cuttings with aqueous solutions of lBA (20-100 p.p.m. ) stimulated...

  2. CONNECTICUT RIVER FISH TISSUE CONTAMINANT STUDY (2000): ECOLOGICAL AND HUMAN HEALTH RISK SCREENING

    EPA Science Inventory

    The study targeted commonly caught recreational fish, as well as other fish that are important in the river food chain. Smallmouth bass, white suckers and yellow perch were collected during 2000 from the mainstem of the Connecticut River and composite samples were analyzed for t...

  3. Grafting Technique to Eliminate Rootstock Suckering of Grafted Tomatoes

    USDA-ARS?s Scientific Manuscript database

    Vegetable grafting has been proposed as a technique for avoiding disease problems in tomatoes in open field production. In this study we investigated the current use of grafting in an open field scenario and found a serious problem with the grafting techniques. In the Fall of 2007, commercially pr...

  4. Drill string enclosure

    DOEpatents

    Jorgensen, D.K.; Kuhns, D.J.; Wiersholm, O.; Miller, T.A.

    1993-03-02

    The drill string enclosure consists of six component parts, including; a top bracket, an upper acrylic cylinder, an acrylic drill casing guide, a lower acrylic cylinder, a bottom bracket, and three flexible ducts. The upper acrylic cylinder is optional based upon the drill string length. The drill string enclosure allows for an efficient drill and sight operation at a hazardous waste site.

  5. Drill string enclosure

    DOEpatents

    Jorgensen, Douglas K.; Kuhns, Douglass J.; Wiersholm, Otto; Miller, Timothy A.

    1993-01-01

    The drill string enclosure consists of six component parts, including; a top bracket, an upper acrylic cylinder, an acrylic drill casing guide, a lower acrylic cylinder, a bottom bracket, and three flexible ducts. The upper acrylic cylinder is optional based upon the drill string length. The drill string enclosure allows for an efficient drill and sight operation at a hazardous waste site.

  6. The Illusive Sound of a Bundengan String

    ERIC Educational Resources Information Center

    Parikesit, Gea O. F.; Kusumaningtyas, Indraswari

    2017-01-01

    The acoustics of a vibrating string is frequently used as a simple example of how physics can be applied in the field of art. In this paper we describe a simple experiment and analysis using a clipped string. This experiment can generate scientific curiosity among students because the sound generated by the string seem surprising to our senses.…

  7. Hammered Strings

    NASA Astrophysics Data System (ADS)

    Rossing, Thomas D.

    In the next three chapters we consider the science of hammered string instruments. In this chapter, we present a brief discussion of vibrating strings excited by a hard or soft hammer. Chapter 20 discusses the most important hammered string instrument, the piano - probably the most versatile and popular of all musical instruments. Chapter 21 discusses hammered dulcimers, especially the American folk dulcimer.

  8. Reflecting on the Rationales for String Study in Schools

    ERIC Educational Resources Information Center

    Brenner, Brenda

    2010-01-01

    This essay will address the question of the value of string education by first examining arguments offered on behalf of string education in schools, and noting their somewhat mixed value. Then a set of arguments will be presented that may have greater promise. The focal point will be the establishment of excellence in string teaching and playing.…

  9. Got 'Em on a String: The Skills, Knowledge and Attributes of Group String Teachers in Queensland

    ERIC Educational Resources Information Center

    Ashton, Graham R.; Klopper, Christopher J.

    2018-01-01

    There appear to be considerable differences in the outcomes of group string teaching programs in Queensland. Some teachers appear to be able to generate, manage, and administrate highly efficacious programs; others seem to experience difficulty transferring the knowledge and skills required for students to become successful string players. As a…

  10. Upgrades to the Closed Bomb Facility and Measurement of Propellant Burning Rate

    DTIC Science & Technology

    2010-01-01

    attenuation ratio myScope.WriteString (":CHAN1:RANGe " + CStr (Me.tbVRange.value)) ‘Sets the vertical voltage range myScope.WriteString (":CHAN1...OFFSet " + CStr (Me.tboffset.value)) ‘Sets the voltage offset myScope.WriteString ":CHAN1:PROB:STYP SING" ‘Sets the signal type...myScope.WriteString (":TRIG:EDGE:SOURce CHAN" + CStr (Int(val(GetRegistry(CollType, cCHANNEL))))) ‘Sets the source channel myScope.WriteString

  11. Bell's Inequalities, Superquantum Correlations, and String Theory

    DOE PAGES

    Chang, Lay Nam; Lewis, Zachary; Minic, Djordje; ...

    2011-01-01

    We offermore » an interpretation of superquantum correlations in terms of a “doubly” quantum theory. We argue that string theory, viewed as a quantum theory with two deformation parameters, the string tension α ' , and the string coupling constant g s , is such a superquantum theory that transgresses the usual quantum violations of Bell's inequalities. We also discuss the ℏ → ∞ limit of quantum mechanics in this context. As a superquantum theory, string theory should display distinct experimentally observable supercorrelations of entangled stringy states.« less

  12. The Hubble Web: The Dark Matter Problem and Cosmic Strings

    NASA Astrophysics Data System (ADS)

    Alexander, Stephon

    2009-07-01

    I propose a reinterpretation of cosmic dark matter in which a rigid network of cosmic strings formed at the end of inflation. The cosmic strings fulfill three functions: At recombination they provide an accretion mechanism for virializing baryonic and warm dark matter into disks. These cosmic strings survive as configurations which thread spiral and elliptical galaxies leading to the observed flatness of rotation curves and the Tully-Fisher relation. We find a relationship between the rotational velocity of the galaxy and the string tension and discuss the testability of this model.

  13. Surface controlled blade stabilizer

    DOEpatents

    Russell, Larry R.

    1983-01-01

    Drill string stabilizer apparatus, controllable to expand and retract entirely from the surface by control of drill string pressure, wherein increase of drill string pressure from the surface closes a valve to create a piston means which is moved down by drill string pressure to expand the stabilizer blades, said valve being opened and the piston moving upward upon reduction of drill string pressure to retract the stabilizer blades. Upward and downward movements of the piston and an actuator sleeve therebelow are controlled by a barrel cam acting between the housing and the actuator sleeve.

  14. Null cosmological singularities and free strings

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Narayan, K.

    2010-03-15

    We continue exploring free strings in the background of null Kasner-like cosmological singularities, following K. Narayan, arXiv:0904.4532. We study the free string Schrodinger wave functional along the lines of K. Narayan, arXiv:0807.1517. We find the wave functional to be nonsingular in the vicinity of singularities whose Kasner exponents satisfy certain relations. We compare this with the description in other variables. We then study certain regulated versions of these singularities where the singular region is replaced by a substringy but nonsingular region and study the string spectra in these backgrounds. The string modes can again be solved for exactly, giving somemore » insight into how string oscillator states get excited near the singularity.« less

  15. CMB temperature trispectrum of cosmic strings

    NASA Astrophysics Data System (ADS)

    Hindmarsh, Mark; Ringeval, Christophe; Suyama, Teruaki

    2010-03-01

    We provide an analytical expression for the trispectrum of the cosmic microwave background (CMB) temperature anisotropies induced by cosmic strings. Our result is derived for the small angular scales under the assumption that the temperature anisotropy is induced by the Gott-Kaiser-Stebbins effect. The trispectrum is predicted to decay with a noninteger power-law exponent ℓ-ρ with 6<ρ<7, depending on the string microstructure, and thus on the string model. For Nambu-Goto strings, this exponent is related to the string mean square velocity and the loop distribution function. We then explore two classes of wave number configuration in Fourier space, the kite and trapezium quadrilaterals. The trispectrum can be of any sign and appears to be strongly enhanced for all squeezed quadrilaterals.

  16. Galileon string measure and other modified measure extended objects

    NASA Astrophysics Data System (ADS)

    Vulfs, T. O.; Guendelman, E. I.

    2017-12-01

    We show that it is possible to formulate string theory as a “Galileon string theory”. The Galileon field χ enters in the definition of the integration measure in the action. Following the methods of the modified measure string theory, we find that the final equations are again those of the sigma-model. Moreover, the string tension appears again as an additional dynamical degree of freedom. At the same time, the theory satisfies all requirements of the Galileon higher derivative theory at the action level while the equations of motion are still of the second-order. A Galileon symmetry is displayed explicitly in the conformal string worldsheet frame. Also, we define the Galileon gauge transformations. Generalizations to branes with other modified measures are discussed.

  17. Large-D gravity and low-D strings.

    PubMed

    Emparan, Roberto; Grumiller, Daniel; Tanabe, Kentaro

    2013-06-21

    We show that in the limit of a large number of dimensions a wide class of nonextremal neutral black holes has a universal near-horizon limit. The limiting geometry is the two-dimensional black hole of string theory with a two-dimensional target space. Its conformal symmetry explains the properties of massless scalars found recently in the large-D limit. For black branes with string charges, the near-horizon geometry is that of the three-dimensional black strings of Horne and Horowitz. The analogies between the α' expansion in string theory and the large-D expansion in gravity suggest a possible effective string description of the large-D limit of black holes. We comment on applications to several subjects, in particular to the problem of critical collapse.

  18. Quantum vacuum interaction between two cosmic strings revisited

    NASA Astrophysics Data System (ADS)

    Muñoz-Castañeda, J. M.; Bordag, M.

    2014-03-01

    We reconsider the quantum vacuum interaction energy between two straight parallel cosmic strings. This problem was discussed several times in an approach treating both strings perturbatively and treating only one perturbatively. Here we point out that a simplifying assumption made by Bordag [Ann. Phys. (Berlin) 47, 93 (1990).] can be justified and show that, despite the global character of the background, the perturbative approach delivers a correct result. We consider the applicability of the scattering methods, developed in the past decade for the Casimir effect, for the cosmic string and find it not applicable. We calculate the scattering T-operator on one string. Finally, we consider the vacuum interaction of two strings when each carries a two-dimensional delta function potential.

  19. Trends in concentrations of polychlorinated biphenyls in fish tissue from selected sites in the Delaware River basin in New Jersey, New York, and Pennsylvania, 1969-98

    USGS Publications Warehouse

    Riva-Murray, Karen; Brightbill, Robin A.; Bilger, Michael D.

    2003-01-01

    Trends in concentrations of polychlorinated biphenyls in fish tissue from selected sites in the Delaware River basin in New Jersey, New York, and Pennsylvania, 1969-98 by Karen Riva-Murray, Robin A. Brightbill, and Michael D. Bilger U.S. GEOLOGICAL SURVEY Water-Resources Investigations Report 01-4066 ABSTRACT Polychlorinated biphenyl (PCB) concentrations in fish tissue collected during the 1990's from selected sites in the Delaware River Basin were compared with concentrations in fish tissue collected during 1969-88. Data collected by State and Federal agencies on concentrations in whole-body common carp (Cyprinus carpio) and white sucker (Catostomus commersoni), and edible portions of American eel (Anguilla rostrata), smallmouth bass (Micropterus dolomieu), and channel catfish (Ictalurus punctatus) during 1969-98 were compiled to define temporal trends in concentrations of PCBs in fish tissue from selected segments of the Delaware River, Lehigh River, Schuylkill River, and Brandywine Creek. The Delaware River in the vicinity of Trenton, New Jersey and Yardley, Pennsylvania (above the tidal influence) had the largest long-term data set among the sites considered for this study and was the only site with sufficient data for statistical analysis. A general pattern of decline in PCB concentrations during 1969-98 was apparent for this river segment. PCB concentrations in whole-body white sucker from this lower Delaware River segment declined during 1969-98 from a highest concentration of 7 micrograms per gram (?g/g, wet weight) in a sample collected during 1972 to 0.26 ?g/g (wet weight) in a sample collected during 1998. PCB concentration was negatively correlated with year (Spearman rank correlation -0.46, p < 0.08, n = 15); especially after removal of a sample from 1977 with an unusually low concentration (Spearman rank correlation -0.53, p = 0.05, n = 14). PCB concentrations in edible flesh of American eel declined during 1975-95, from a highest concentration of 3.8 ?g/g (wet weight) in a sample collected during 1976 to less than the reporting limit of 0.26 ?g/g (wet weight) in samples collected during 1993 and 1995. PCB concentrations in most samples (for species considered in this study) collected from the lower Delaware River exceeded the National Academy of Sciences and National Academy of Engineering (NAS/NAE) wildlife guideline level of 0.5 ?g/g during the 1970's and 1980's, and decreased to below this level during the 1990's. No samples of edible portions of game fish exceeded the U.S. Food and Drug Administration (FDA) tolerance level by the mid 1980's. However, the PCB concentration in a smallmouth bass fillet sample that was collected during 1998 (0.37 ?g/g) exceeded the Pennsylvania fish-consumption advisory level of 0.06 ?g/g, and the concentrations in whole-body common carp and white sucker collected during 1998 (1.10 ?g/g and 0.26 ?g/g, respectively) exceeded the New York State Department of Environmental Conservation wildlife criterion concentration of 0.11 ?g/g. (The concentration in carp also exceeded the 1973 NAS/NAE wildlife guideline concentration of 0.5 ?g/g.) Graphical analysis of PCB concentrations in whole white sucker and (or) edible portions of American eel from the upper Delaware River, lower Delaware River, middle Schuylkill River, and Brandywine Creek indicate a decline from the 1970's and (or) 1980's to the middle to late 1990's. Temporal trends in PCB concentrations in white sucker samples from the lower Lehigh and Schuylkill Rivers during 1979-98 are less clear; the PCB concentration (wet-weight basis) from a sample collected in 1998 from the lower Lehigh River was similar to that from a sample collected in 1979, and concentrations actually increased during 1982-98. Similarly, PCB concentrations in samples of white sucker and American eel from the lower Schuylkill River were highly variable over time. A decrease in lipid-adjusted PCB concentrations at both sites (for several whi

  20. Racquet string tension directly affects force experienced at the elbow: implications for the development of lateral epicondylitis in tennis players

    PubMed Central

    Mohandhas, Badri R; Makaram, Navnit; Drew, Tim S; Wang, Weijie; Arnold, Graham P

    2016-01-01

    Background Lateral epicondylitis (LE) occurs in almost half of all tennis players. Racket-string tension is considered to be an important factor influencing the development of LE. No literature yet exists that substantiates how string-tension affects force transmission to the elbow, as implicated in LE development. We establish a quantitative relationship between string-tension and elbow loading, analyzing tennis strokes using rackets with varying string-tensions. Methods Twenty recreational tennis players simulated backhand tennis strokes using three rackets strung at tensions of 200 N, 222 N and 245 N. Accelerometers recorded accelerations at the elbow, wrist and racket handle. Average peak acceleration was determined to correlate string-tension with elbow loading. Results Statistically significant differences (p < 0.05) were observed when average peak acceleration at the elbow at 200 N string-tension (acceleration of 5.58 m/s2) was compared with that at 222 N tension (acceleration of 6.83 m/s2) and 245 N tension (acceleration of 7.45 m/s2). The 200 N racket induced the least acceleration at the elbow. Conclusions Although parameters determining force transmission to the elbow during a tennis stroke are complex, the present study was able to control these parameters, isolating the effect of string-tension. Lower string-tensions transmit less force to the elbow in backhand strokes. Reducing string-tension should be considered favourably with respect to reducing the risk of developing LE. PMID:27583017

  1. Integrated hydraulic booster/tool string technology for unfreezing of stuck downhole strings in horizontal wells

    NASA Astrophysics Data System (ADS)

    Tian, Q. Z.

    2017-12-01

    It is common to use a jarring tool to unfreeze stuck downhole string. However, in a horizontal well, influenced by the friction caused by the deviated section, jarring effect is poor; on the other hand, the forcing point can be located in the horizontal section by a hydraulic booster and the friction can be reduced, but it is time-consuming and easy to break downhole string using a large-tonnage and constant pull force. A hydraulic booster - jar tool string has been developed for unfreezing operation in horizontal wells. The technical solution involves three elements: a two-stage parallel spring cylinder structure for increasing the energy storage capacity of spring accelerators; multiple groups of spring accelerators connected in series to increase the working stroke; a hydraulic booster intensifying jarring force. The integrated unfreezing tool string based on these three elements can effectively overcome the friction caused by a deviated borehole, and thus unfreeze a stuck string with the interaction of the hydraulic booster and the mechanical jar which form an alternatively dynamic load. Experimental results show that the jarring performance parameters of the hydraulic booster-jar unfreezing tool string for the horizontal wells are in accordance with original design requirements. Then field technical parameters were developed based on numerical simulation and experimental data. Field application shows that the hydraulic booster-jar unfreezing tool string is effective to free stuck downhole tools in a horizontal well, and it reduces hook load by 80% and lessens the requirement of workover equipment. This provides a new technology to unfreeze stuck downhole string in a horizontal well.

  2. Effect of changing of the parameters of the cable network of monitoring systems of high-rise buildings on the basis of string converters on their operability

    NASA Astrophysics Data System (ADS)

    Gusev, Nikolay; Svatovskaya, Larisa; Kucherenko, Alexandr

    2018-03-01

    The article is devoted to the problem of improving the reliability of monitoring systems for the technical conditions of high-rise buildings. The improvement is based on string sensors with an impulsed excitation method ensuring the maximum signal-to-noise ratio at their output. The influence of the parameters of the monitoring system on the shape of the excitation impulses of the string, and, consequently, on the amplitude of the string vibration of the string converter is also considered in the article. It has been experimentally proved that the parameters of the excitation impulses of the string converters. The article presents the results of the experiments showing the effect of the fronts duration of the excitation impulses on the amplitude of the oscillations of the strings. The influence of the fronts duration of the excitation impulse with the frontal lengths up to 0.5 ms is studied at the excitation impulse duration not exceeding 0.5 times the duration of natural oscillation periods of the string. The experimental data are compared with the theoretical ones and hypotheses explaining their difference are advanced. The article suggests some methods of reducing the influence of the cable-switching equipment system parameters on the amplitude of string oscillations. The possibilities of improving the reliability of the systems developed on the basis of string sensors with an impulsed excitation method and used for monitoring the technical conditions of the high-rise buildings are proposed.

  3. Whiteheadian Actual Entitities and String Theory

    NASA Astrophysics Data System (ADS)

    Bracken, Joseph A.

    2012-06-01

    In the philosophy of Alfred North Whitehead, the ultimate units of reality are actual entities, momentary self-constituting subjects of experience which are too small to be sensibly perceived. Their combination into "societies" with a "common element of form" produces the organisms and inanimate things of ordinary sense experience. According to the proponents of string theory, tiny vibrating strings are the ultimate constituents of physical reality which in harmonious combination yield perceptible entities at the macroscopic level of physical reality. Given that the number of Whiteheadian actual entities and of individual strings within string theory are beyond reckoning at any given moment, could they be two ways to describe the same non-verifiable foundational reality? For example, if one could establish that the "superject" or objective pattern of self- constitution of an actual entity vibrates at a specific frequency, its affinity with the individual strings of string theory would be striking. Likewise, if one were to claim that the size and complexity of Whiteheadian 'societies" require different space-time parameters for the dynamic interrelationship of constituent actual entities, would that at least partially account for the assumption of 10 or even 26 instead of just 3 dimensions within string theory? The overall conclusion of this article is that, if a suitably revised understanding of Whiteheadian metaphysics were seen as compatible with the philosophical implications of string theory, their combination into a single world view would strengthen the plausibility of both schemes taken separately. Key words: actual entities, subject/superjects, vibrating strings, structured fields of activity, multi-dimensional physical reality.

  4. Detection of low tension cosmic superstrings

    NASA Astrophysics Data System (ADS)

    Chernoff, David F.; Tye, S.-H. Henry

    2018-05-01

    Cosmic superstrings of string theory differ from conventional cosmic strings of field theory. We review how the physical and cosmological properties of the macroscopic string loops influence experimental searches for these relics from the epoch of inflation. The universe's average density of cosmic superstrings can easily exceed that of conventional cosmic strings having the same tension by two or more orders of magnitude. The cosmological behavior of the remnant superstring loops is qualitatively distinct because the string tension is exponentially smaller than the string scale in flux compactifications in string theory. Low tension superstring loops live longer, experience less recoil (rocket effect from the emission of gravitational radiation) and tend to cluster like dark matter in galaxies. Clustering enhances the string loop density with respect to the cosmological average in collapsed structures in the universe. The enhancement at the Sun's position is ~ 105. We develop a model encapsulating the leading order string theory effects, the current understanding of the string network loop production and the influence of cosmological structure formation suitable for forecasting the detection of superstring loops via optical microlensing, gravitational wave bursts and fast radio bursts. We evaluate the detection rate of bursts from cusps and kinks by LIGO- and LISA-like experiments. Clustering dominates rates for G μ < 10‑11.9 (LIGO cusp), G μ<10‑11.2 (LISA cusp), G μ < 10‑10.6 (LISA kink); we forecast experimentally accessible gravitational wave bursts for G μ>10‑14.2 (LIGO cusp), G μ>10‑15 (LISA cusp) and G μ>10‑ 14.1 (LISA kink).

  5. Constraints on cosmic strings using data from the first Advanced LIGO observing run

    NASA Astrophysics Data System (ADS)

    Abbott, B. P.; Abbott, R.; Abbott, T. D.; Acernese, F.; Ackley, K.; Adams, C.; Adams, T.; Addesso, P.; Adhikari, R. X.; Adya, V. B.; Affeldt, C.; Afrough, M.; Agarwal, B.; Agathos, M.; Agatsuma, K.; Aggarwal, N.; Aguiar, O. D.; Aiello, L.; Ain, A.; Ajith, P.; Allen, B.; Allen, G.; Allocca, A.; Altin, P. A.; Amato, A.; Ananyeva, A.; Anderson, S. B.; Anderson, W. G.; Antier, S.; Appert, S.; Arai, K.; Araya, M. C.; Areeda, J. S.; Arnaud, N.; Arun, K. G.; Ascenzi, S.; Ashton, G.; Ast, M.; Aston, S. M.; Astone, P.; Aufmuth, P.; Aulbert, C.; AultONeal, K.; Avila-Alvarez, A.; Babak, S.; Bacon, P.; Bader, M. K. M.; Bae, S.; Baker, P. T.; Baldaccini, F.; Ballardin, G.; Ballmer, S. W.; Banagiri, S.; Barayoga, J. C.; Barclay, S. E.; Barish, B. C.; Barker, D.; Barone, F.; Barr, B.; Barsotti, L.; Barsuglia, M.; Barta, D.; Bartlett, J.; Bartos, I.; Bassiri, R.; Basti, A.; Batch, J. C.; Baune, C.; Bawaj, M.; Bazzan, M.; Bécsy, B.; Beer, C.; Bejger, M.; Belahcene, I.; Bell, A. S.; Berger, B. K.; Bergmann, G.; Berry, C. P. L.; Bersanetti, D.; Bertolini, A.; Betzwieser, J.; Bhagwat, S.; Bhandare, R.; Bilenko, I. A.; Billingsley, G.; Billman, C. R.; Birch, J.; Birney, R.; Birnholtz, O.; Biscans, S.; Bisht, A.; Bitossi, M.; Biwer, C.; Bizouard, M. A.; Blackburn, J. K.; Blackman, J.; Blair, C. D.; Blair, D. G.; Blair, R. M.; Bloemen, S.; Bock, O.; Bode, N.; Boer, M.; Bogaert, G.; Bohe, A.; Bondu, F.; Bonnand, R.; Boom, B. A.; Bork, R.; Boschi, V.; Bose, S.; Bouffanais, Y.; Bozzi, A.; Bradaschia, C.; Brady, P. R.; Braginsky, V. B.; Branchesi, M.; Brau, J. E.; Briant, T.; Brillet, A.; Brinkmann, M.; Brisson, V.; Brockill, P.; Broida, J. E.; Brooks, A. F.; Brown, D. A.; Brown, D. D.; Brown, N. M.; Brunett, S.; Buchanan, C. C.; Buikema, A.; Bulik, T.; Bulten, H. J.; Buonanno, A.; Buskulic, D.; Buy, C.; Byer, R. L.; Cabero, M.; Cadonati, L.; Cagnoli, G.; Cahillane, C.; Calderón Bustillo, J.; Callister, T. A.; Calloni, E.; Camp, J. B.; Canepa, M.; Canizares, P.; Cannon, K. C.; Cao, H.; Cao, J.; Capano, C. D.; Capocasa, E.; Carbognani, F.; Caride, S.; Carney, M. F.; Casanueva Diaz, J.; Casentini, C.; Caudill, S.; Cavaglià, M.; Cavalier, F.; Cavalieri, R.; Cella, G.; Cepeda, C. B.; Cerboni Baiardi, L.; Cerretani, G.; Cesarini, E.; Chamberlin, S. J.; Chan, M.; Chao, S.; Charlton, P.; Chassande-Mottin, E.; Chatterjee, D.; Cheeseboro, B. D.; Chen, H. Y.; Chen, Y.; Cheng, H.-P.; Chincarini, A.; Chiummo, A.; Chmiel, T.; Cho, H. S.; Cho, M.; Chow, J. H.; Christensen, N.; Chu, Q.; Chua, A. J. K.; Chua, S.; Chung, A. K. W.; Chung, S.; Ciani, G.; Ciolfi, R.; Cirelli, C. E.; Cirone, A.; Clara, F.; Clark, J. A.; Cleva, F.; Cocchieri, C.; Coccia, E.; Cohadon, P.-F.; Colla, A.; Collette, C. G.; Cominsky, L. R.; Constancio, M.; Conti, L.; Cooper, S. J.; Corban, P.; Corbitt, T. R.; Corley, K. R.; Cornish, N.; Corsi, A.; Cortese, S.; Costa, C. A.; Coughlin, M. W.; Coughlin, S. B.; Coulon, J.-P.; Countryman, S. T.; Couvares, P.; Covas, P. B.; Cowan, E. E.; Coward, D. M.; Cowart, M. J.; Coyne, D. C.; Coyne, R.; Creighton, J. D. E.; Creighton, T. D.; Cripe, J.; Crowder, S. G.; Cullen, T. J.; Cumming, A.; Cunningham, L.; Cuoco, E.; Dal Canton, T.; Danilishin, S. L.; D'Antonio, S.; Danzmann, K.; Dasgupta, A.; Da Silva Costa, C. F.; Dattilo, V.; Dave, I.; Davier, M.; Davis, D.; Daw, E. J.; Day, B.; De, S.; DeBra, D.; Degallaix, J.; De Laurentis, M.; Deléglise, S.; Del Pozzo, W.; Denker, T.; Dent, T.; Dergachev, V.; De Rosa, R.; DeRosa, R. T.; DeSalvo, R.; Devenson, J.; Devine, R. C.; Dhurandhar, S.; Díaz, M. C.; Di Fiore, L.; Di Giovanni, M.; Di Girolamo, T.; Di Lieto, A.; Di Pace, S.; Di Palma, I.; Di Renzo, F.; Doctor, Z.; Dolique, V.; Donovan, F.; Dooley, K. L.; Doravari, S.; Dorrington, I.; Douglas, R.; Dovale Álvarez, M.; Downes, T. P.; Drago, M.; Drever, R. W. P.; Driggers, J. C.; Du, Z.; Ducrot, M.; Duncan, J.; Dwyer, S. E.; Edo, T. B.; Edwards, M. C.; Effler, A.; Eggenstein, H.-B.; Ehrens, P.; Eichholz, J.; Eikenberry, S. S.; Eisenstein, R. A.; Essick, R. C.; Etienne, Z. B.; Etzel, T.; Evans, M.; Evans, T. M.; Factourovich, M.; Fafone, V.; Fair, H.; Fairhurst, S.; Fan, X.; Farinon, S.; Farr, B.; Farr, W. M.; Fauchon-Jones, E. J.; Favata, M.; Fays, M.; Fehrmann, H.; Feicht, J.; Fejer, M. M.; Fernandez-Galiana, A.; Ferrante, I.; Ferreira, E. C.; Ferrini, F.; Fidecaro, F.; Fiori, I.; Fiorucci, D.; Fisher, R. P.; Fitz-Axen, M.; Flaminio, R.; Fletcher, M.; Fong, H.; Forsyth, P. W. F.; Forsyth, S. S.; Fournier, J.-D.; Frasca, S.; Frasconi, F.; Frei, Z.; Freise, A.; Frey, R.; Frey, V.; Fries, E. M.; Fritschel, P.; Frolov, V. V.; Fulda, P.; Fyffe, M.; Gabbard, H.; Gabel, M.; Gadre, B. U.; Gaebel, S. M.; Gair, J. R.; Gammaitoni, L.; Ganija, M. R.; Gaonkar, S. G.; Garufi, F.; Gaudio, S.; Gaur, G.; Gayathri, V.; Gehrels, N.; Gemme, G.; Genin, E.; Gennai, A.; George, D.; George, J.; Gergely, L.; Germain, V.; Ghonge, S.; Ghosh, Abhirup; Ghosh, Archisman; Ghosh, S.; Giaime, J. A.; Giardina, K. D.; Giazotto, A.; Gill, K.; Glover, L.; Goetz, E.; Goetz, R.; Gomes, S.; González, G.; Gonzalez Castro, J. M.; Gopakumar, A.; Gorodetsky, M. L.; Gossan, S. E.; Gosselin, M.; Gouaty, R.; Grado, A.; Graef, C.; Granata, M.; Grant, A.; Gras, S.; Gray, C.; Greco, G.; Green, A. C.; Groot, P.; Grote, H.; Grunewald, S.; Gruning, P.; Guidi, G. M.; Guo, X.; Gupta, A.; Gupta, M. K.; Gushwa, K. E.; Gustafson, E. K.; Gustafson, R.; Hall, B. R.; Hall, E. D.; Hammond, G.; Haney, M.; Hanke, M. M.; Hanks, J.; Hanna, C.; Hannam, M. D.; Hannuksela, O. A.; Hanson, J.; Hardwick, T.; Harms, J.; Harry, G. M.; Harry, I. W.; Hart, M. J.; Haster, C.-J.; Haughian, K.; Healy, J.; Heidmann, A.; Heintze, M. C.; Heitmann, H.; Hello, P.; Hemming, G.; Hendry, M.; Heng, I. S.; Hennig, J.; Henry, J.; Heptonstall, A. W.; Heurs, M.; Hild, S.; Hoak, D.; Hofman, D.; Holt, K.; Holz, D. E.; Hopkins, P.; Horst, C.; Hough, J.; Houston, E. A.; Howell, E. J.; Hu, Y. M.; Huerta, E. A.; Huet, D.; Hughey, B.; Husa, S.; Huttner, S. H.; Huynh-Dinh, T.; Indik, N.; Ingram, D. R.; Inta, R.; Intini, G.; Isa, H. N.; Isac, J.-M.; Isi, M.; Iyer, B. R.; Izumi, K.; Jacqmin, T.; Jani, K.; Jaranowski, P.; Jawahar, S.; Jiménez-Forteza, F.; Johnson, W. W.; Jones, D. I.; Jones, R.; Jonker, R. J. G.; Ju, L.; Junker, J.; Kalaghatgi, C. V.; Kalogera, V.; Kandhasamy, S.; Kang, G.; Kanner, J. B.; Karki, S.; Karvinen, K. S.; Kasprzack, M.; Katolik, M.; Katsavounidis, E.; Katzman, W.; Kaufer, S.; Kawabe, K.; Kéfélian, F.; Keitel, D.; Kemball, A. J.; Kennedy, R.; Kent, C.; Key, J. S.; Khalili, F. Y.; Khan, I.; Khan, S.; Khan, Z.; Khazanov, E. A.; Kijbunchoo, N.; Kim, Chunglee; Kim, J. C.; Kim, W.; Kim, W. S.; Kim, Y.-M.; Kimbrell, S. J.; King, E. J.; King, P. J.; Kirchhoff, R.; Kissel, J. S.; Kleybolte, L.; Klimenko, S.; Koch, P.; Koehlenbeck, S. M.; Koley, S.; Kondrashov, V.; Kontos, A.; Korobko, M.; Korth, W. Z.; Kowalska, I.; Kozak, D. B.; Krämer, C.; Kringel, V.; Krishnan, B.; Królak, A.; Kuehn, G.; Kumar, P.; Kumar, R.; Kumar, S.; Kuo, L.; Kutynia, A.; Kwang, S.; Lackey, B. D.; Lai, K. H.; Landry, M.; Lang, R. N.; Lange, J.; Lantz, B.; Lanza, R. K.; Lartaux-Vollard, A.; Lasky, P. D.; Laxen, M.; Lazzarini, A.; Lazzaro, C.; Leaci, P.; Leavey, S.; Lee, C. H.; Lee, H. K.; Lee, H. M.; Lee, H. W.; Lee, K.; Lehmann, J.; Lenon, A.; Leonardi, M.; Leroy, N.; Letendre, N.; Levin, Y.; Li, T. G. F.; Libson, A.; Littenberg, T. B.; Liu, J.; Lo, R. K. L.; Lockerbie, N. A.; London, L. T.; Lord, J. E.; Lorenzini, M.; Loriette, V.; Lormand, M.; Losurdo, G.; Lough, J. D.; Lousto, C. O.; Lovelace, G.; Lück, H.; Lumaca, D.; Lundgren, A. P.; Lynch, R.; Ma, Y.; Macfoy, S.; Machenschalk, B.; MacInnis, M.; Macleod, D. M.; Magaña Hernandez, I.; Magaña-Sandoval, F.; Magaña Zertuche, L.; Magee, R. M.; Majorana, E.; Maksimovic, I.; Man, N.; Mandic, V.; Mangano, V.; Mansell, G. L.; Manske, M.; Mantovani, M.; Marchesoni, F.; Marion, F.; Márka, S.; Márka, Z.; Markakis, C.; Markosyan, A. S.; Maros, E.; Martelli, F.; Martellini, L.; Martin, I. W.; Martynov, D. V.; Mason, K.; Masserot, A.; Massinger, T. J.; Masso-Reid, M.; Mastrogiovanni, S.; Matas, A.; Matichard, F.; Matone, L.; Mavalvala, N.; Mazumder, N.; McCarthy, R.; McClelland, D. E.; McCormick, S.; McCuller, L.; McGuire, S. C.; McIntyre, G.; McIver, J.; McManus, D. J.; McRae, T.; McWilliams, S. T.; Meacher, D.; Meadors, G. D.; Meidam, J.; Mejuto-Villa, E.; Melatos, A.; Mendell, G.; Mercer, R. A.; Merilh, E. L.; Merzougui, M.; Meshkov, S.; Messenger, C.; Messick, C.; Metzdorff, R.; Meyers, P. M.; Mezzani, F.; Miao, H.; Michel, C.; Middleton, H.; Mikhailov, E. E.; Milano, L.; Miller, A. L.; Miller, A.; Miller, B. B.; Miller, J.; Millhouse, M.; Minazzoli, O.; Minenkov, Y.; Ming, J.; Mishra, C.; Mitra, S.; Mitrofanov, V. P.; Mitselmakher, G.; Mittleman, R.; Moggi, A.; Mohan, M.; Mohapatra, S. R. P.; Montani, M.; Moore, B. C.; Moore, C. J.; Moraru, D.; Moreno, G.; Morriss, S. R.; Mours, B.; Mow-Lowry, C. M.; Mueller, G.; Muir, A. W.; Mukherjee, Arunava; Mukherjee, D.; Mukherjee, S.; Mukund, N.; Mullavey, A.; Munch, J.; Muniz, E. A. M.; Murray, P. G.; Napier, K.; Nardecchia, I.; Naticchioni, L.; Nayak, R. K.; Nelemans, G.; Nelson, T. J. N.; Neri, M.; Nery, M.; Neunzert, A.; Newport, J. M.; Newton, G.; Ng, K. K. Y.; Nguyen, T. T.; Nichols, D.; Nielsen, A. B.; Nissanke, S.; Nitz, A.; Noack, A.; Nocera, F.; Nolting, D.; Normandin, M. E. N.; Nuttall, L. K.; Oberling, J.; Ochsner, E.; Oelker, E.; Ogin, G. H.; Oh, J. J.; Oh, S. H.; Ohme, F.; Oliver, M.; Oppermann, P.; Oram, Richard J.; O'Reilly, B.; Ormiston, R.; Ortega, L. F.; O'Shaughnessy, R.; Ottaway, D. J.; Overmier, H.; Owen, B. J.; Pace, A. E.; Page, J.; Page, M. A.; Pai, A.; Pai, S. A.; Palamos, J. R.; Palashov, O.; Palomba, C.; Pal-Singh, A.; Pan, H.; Pang, B.; Pang, P. T. H.; Pankow, C.; Pannarale, F.; Pant, B. C.; Paoletti, F.; Paoli, A.; Papa, M. A.; Paris, H. R.; Parker, W.; Pascucci, D.; Pasqualetti, A.; Passaquieti, R.; Passuello, D.; Patricelli, B.; Pearlstone, B. L.; Pedraza, M.; Pedurand, R.; Pekowsky, L.; Pele, A.; Penn, S.; Perez, C. J.; Perreca, A.; Perri, L. M.; Pfeiffer, H. P.; Phelps, M.; Piccinni, O. J.; Pichot, M.; Piergiovanni, F.; Pierro, V.; Pillant, G.; Pinard, L.; Pinto, I. M.; Pitkin, M.; Poggiani, R.; Popolizio, P.; Porter, E. K.; Post, A.; Powell, J.; Prasad, J.; Pratt, J. W. W.; Predoi, V.; Prestegard, T.; Prijatelj, M.; Principe, M.; Privitera, S.; Prix, R.; Prodi, G. A.; Prokhorov, L. G.; Puncken, O.; Punturo, M.; Puppo, P.; Pürrer, M.; Qi, H.; Qin, J.; Qiu, S.; Quetschke, V.; Quintero, E. A.; Quitzow-James, R.; Raab, F. J.; Rabeling, D. S.; Radkins, H.; Raffai, P.; Raja, S.; Rajan, C.; Rakhmanov, M.; Ramirez, K. E.; Rapagnani, P.; Raymond, V.; Razzano, M.; Read, J.; Regimbau, T.; Rei, L.; Reid, S.; Reitze, D. H.; Rew, H.; Reyes, S. D.; Ricci, F.; Ricker, P. M.; Rieger, S.; Riles, K.; Rizzo, M.; Robertson, N. A.; Robie, R.; Robinet, F.; Rocchi, A.; Rolland, L.; Rollins, J. G.; Roma, V. J.; Romano, J. D.; Romano, R.; Romel, C. L.; Romie, J. H.; Rosińska, D.; Ross, M. P.; Rowan, S.; Rüdiger, A.; Ruggi, P.; Ryan, K.; Sachdev, S.; Sadecki, T.; Sadeghian, L.; Sakellariadou, M.; Salconi, L.; Saleem, M.; Salemi, F.; Samajdar, A.; Sammut, L.; Sampson, L. M.; Sanchez, E. J.; Sandberg, V.; Sandeen, B.; Sanders, J. R.; Sassolas, B.; Saulson, P. R.; Sauter, O.; Savage, R. L.; Sawadsky, A.; Schale, P.; Scheuer, J.; Schmidt, E.; Schmidt, J.; Schmidt, P.; Schnabel, R.; Schofield, R. M. S.; Schönbeck, A.; Schreiber, E.; Schuette, D.; Schulte, B. W.; Schutz, B. F.; Schwalbe, S. G.; Scott, J.; Scott, S. M.; Seidel, E.; Sellers, D.; Sengupta, A. S.; Sentenac, D.; Sequino, V.; Sergeev, A.; Shaddock, D. A.; Shaffer, T. J.; Shah, A. A.; Shahriar, M. S.; Shao, L.; Shapiro, B.; Shawhan, P.; Sheperd, A.; Shoemaker, D. H.; Shoemaker, D. M.; Siellez, K.; Siemens, X.; Sieniawska, M.; Sigg, D.; Silva, A. D.; Singer, A.; Singer, L. P.; Singh, A.; Singh, R.; Singhal, A.; Sintes, A. M.; Slagmolen, B. J. J.; Smith, B.; Smith, J. R.; Smith, R. J. E.; Son, E. J.; Sonnenberg, J. A.; Sorazu, B.; Sorrentino, F.; Souradeep, T.; Spencer, A. P.; Srivastava, A. K.; Staley, A.; Steer, D. A.; Steinke, M.; Steinlechner, J.; Steinlechner, S.; Steinmeyer, D.; Stephens, B. C.; Stone, R.; Strain, K. A.; Stratta, G.; Strigin, S. E.; Sturani, R.; Stuver, A. L.; Summerscales, T. Z.; Sun, L.; Sunil, S.; Sutton, P. J.; Swinkels, B. L.; Szczepańczyk, M. J.; Tacca, M.; Talukder, D.; Tanner, D. B.; Tápai, M.; Taracchini, A.; Taylor, J. A.; Taylor, R.; Theeg, T.; Thomas, E. G.; Thomas, M.; Thomas, P.; Thorne, K. A.; Thorne, K. S.; Thrane, E.; Tiwari, S.; Tiwari, V.; Tokmakov, K. V.; Toland, K.; Tonelli, M.; Tornasi, Z.; Torrie, C. I.; Töyrä, D.; Travasso, F.; Traylor, G.; Trifirò, D.; Trinastic, J.; Tringali, M. C.; Trozzo, L.; Tsang, K. W.; Tse, M.; Tso, R.; Tuyenbayev, D.; Ueno, K.; Ugolini, D.; Unnikrishnan, C. S.; Urban, A. L.; Usman, S. A.; Vahlbruch, H.; Vajente, G.; Valdes, G.; Vallisneri, M.; van Bakel, N.; van Beuzekom, M.; van den Brand, J. F. J.; Van Den Broeck, C.; Vander-Hyde, D. C.; van der Schaaf, L.; van Heijningen, J. V.; van Veggel, A. A.; Vardaro, M.; Varma, V.; Vass, S.; Vasúth, M.; Vecchio, A.; Vedovato, G.; Veitch, J.; Veitch, P. J.; Venkateswara, K.; Venugopalan, G.; Verkindt, D.; Vetrano, F.; Viceré, A.; Viets, A. D.; Vinciguerra, S.; Vine, D. J.; Vinet, J.-Y.; Vitale, S.; Vo, T.; Vocca, H.; Vorvick, C.; Voss, D. V.; Vousden, W. D.; Vyatchanin, S. P.; Wade, A. R.; Wade, L. E.; Wade, M.; Walet, R.; Walker, M.; Wallace, L.; Walsh, S.; Wang, G.; Wang, H.; Wang, J. Z.; Wang, M.; Wang, Y.-F.; Wang, Y.; Ward, R. L.; Warner, J.; Was, M.; Watchi, J.; Weaver, B.; Wei, L.-W.; Weinert, M.; Weinstein, A. J.; Weiss, R.; Wen, L.; Wessel, E. K.; Weßels, P.; Westphal, T.; Wette, K.; Whelan, J. T.; Whiting, B. F.; Whittle, C.; Williams, D.; Williams, R. D.; Williamson, A. R.; Willis, J. L.; Willke, B.; Wimmer, M. H.; Winkler, W.; Wipf, C. C.; Wittel, H.; Woan, G.; Woehler, J.; Wofford, J.; Wong, K. W. K.; Worden, J.; Wright, J. L.; Wu, D. S.; Wu, G.; Yam, W.; Yamamoto, H.; Yancey, C. C.; Yap, M. J.; Yu, Hang; Yu, Haocun; Yvert, M.; ZadroŻny, A.; Zanolin, M.; Zelenova, T.; Zendri, J.-P.; Zevin, M.; Zhang, L.; Zhang, M.; Zhang, T.; Zhang, Y.-H.; Zhao, C.; Zhou, M.; Zhou, Z.; Zhu, S. J.; Zhu, X. J.; Zucker, M. E.; Zweizig, J.; LIGO Scientific Collaboration; Virgo Collaboration

    2018-05-01

    Cosmic strings are topological defects which can be formed in grand unified theory scale phase transitions in the early universe. They are also predicted to form in the context of string theory. The main mechanism for a network of Nambu-Goto cosmic strings to lose energy is through the production of loops and the subsequent emission of gravitational waves, thus offering an experimental signature for the existence of cosmic strings. Here we report on the analysis conducted to specifically search for gravitational-wave bursts from cosmic string loops in the data of Advanced LIGO 2015-2016 observing run (O1). No evidence of such signals was found in the data, and as a result we set upper limits on the cosmic string parameters for three recent loop distribution models. In this paper, we initially derive constraints on the string tension G μ and the intercommutation probability, using not only the burst analysis performed on the O1 data set but also results from the previously published LIGO stochastic O1 analysis, pulsar timing arrays, cosmic microwave background and big-bang nucleosynthesis experiments. We show that these data sets are complementary in that they probe gravitational waves produced by cosmic string loops during very different epochs. Finally, we show that the data sets exclude large parts of the parameter space of the three loop distribution models we consider.

  6. Modeling the influence of string collective phenomena on the long range rapidity correlations between the transverse momentum and the multiplicities

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Andronov, E.; Vechernin, V.

    2016-01-22

    The long-range rapidity correlations between the multiplicities (n-n) and the transverse momentum and the multiplicity (pT-n) of charge particles are analyzed in the framework of the simple string inspired model with two types of sources. The sources of the first type correspond to the initial strings formed in a hadronic collision. The sources of the second type imitate the appearance of the emitters of a new kind resulting from interaction (fusion) of the initial strings. The model enabled to describe effectively the influence of the string fusion effects on the strength both the n-n and the pT-n correlations. It wasmore » found that in the region, where the process of string fusion comes into play, the calculation results predict the non-monotonic behaviour of the n-n and pT-n correlation coefficients with the growth of the mean number of initial strings, i.e. with the increase of the collision centrality. It was shown also that the increase of the event-by-event fluctuation in the number of primary strings leads to the change of the pT-n correlation sign from negative to positive. One can try to search these signatures of string collective phenomena in interactions of various nuclei at different energies varying the class of collision centrality and its width.« less

  7. On Atwood's Machine with a Nonzero Mass String

    NASA Astrophysics Data System (ADS)

    Tarnopolski, Mariusz

    2015-11-01

    Let us consider a classical high school exercise concerning two weights on a pulley and a string, illustrated in Fig. 1(a). A system like this is called an Atwood's machine and was invented by George Atwood in 1784 as a laboratory experiment to verify the mechanical laws of motion with constant acceleration. Nowadays, Atwood's machine is used for didactic purposes to demonstrate uniformly accelerated motion with acceleration arbitrarily smaller than the gravitational acceleration g. The simplest case is with a massless and frictionless pulley and a massless string. With little effort one can include the mass of the pulley in calculations. The mass of a string has been incorporated previously in some considerations and experiments. These include treatments focusing on friction, justifying the assumption of a massless string, incorporating variations in Earth's gravitational field, comparing the calculated value of g based on a simple experiment, taking the mass of the string into account in such a way that the resulting acceleration is constant, or in one exception solely focusing on a heavy string, but with a slightly different approach. Here we wish to provide i) a derivation of the acceleration and position dependence on the weights' masses based purely on basic dynamical reasoning similar to the conventional version of the exercise, and ii) focus on the influence of the string's linear density, or equivalently its mass, on the outcome compared to a massless string case.

  8. A group theoretic method for string loop diagram

    NASA Astrophysics Data System (ADS)

    Neveu, A.; West, P.

    1987-08-01

    The new approach to arbitrary string scattering proposed by the authors is used to compute the planar tadpole operator, including its measure, for the open bosonic string. The Virasoro gauge identities play a crucial role and are found to contain in general anomalous terms for open strings. Permanent address: Mathematics Department, King's College, London WC2R 2LS, UK.

  9. Self-Gravitating Fundamental Strings and Black Holes

    NASA Technical Reports Server (NTRS)

    Damour, T.; Veneziano, G.

    1999-01-01

    The configuration of typically highly excited M much greater than M(sub s) which is approximately equal to alpha(prime) to the 1/2 power string states is considered as the string coupling g is adiabatically increased. The size distribution of very massive single string states is studied and the mass shift, due to a long-range gravitational, dilatonic, and axionic attraction, is estimated.

  10. String junction as a baryonic constituent

    NASA Astrophysics Data System (ADS)

    Kalashnikova, Yu. S.; Nefediev, A. V.

    1996-02-01

    We extend the model for QCD string with quarks to consider the Mercedes Benz string configuration describing the three-quark baryon. Under the assumption of adiabatic separation of quark and string junction motion we formulate and solve the classical equation of motion for the junction. We dare to quantize the motion of the junction, and discuss the impact of these modes on the baryon spectra.

  11. Development and Trial of a Two Year Program of String Instruction. Final Report.

    ERIC Educational Resources Information Center

    Rolland, Paul; And Others

    A series of films focused on movement education and rhythm training in string playing with emphasis on the violin were developed. An introductory film deals with principles of movement in string playing. Fifteen additional titles offer guidance to the student and teacher in the various details of basic string instruction. A summary film presents a…

  12. The Physics of "String Passing through Ice"

    ERIC Educational Resources Information Center

    Mohazzabi, Pirooz

    2011-01-01

    One of the oldest yet interesting experiments related to heat and thermodynamics is placing a string on a block of ice and hanging two masses from the ends of the string. Sometime later, it is discovered that the string has passed through the ice without cutting it in half. A simple explanation of this effect is that the pressure caused by the…

  13. String Stability of a Linear Formation Flight Control System

    NASA Technical Reports Server (NTRS)

    Allen, Michael J.; Ryan, Jack; Hanson, Curtis E.; Parle, James F.

    2002-01-01

    String stability analysis of an autonomous formation flight system was performed using linear and nonlinear simulations. String stability is a measure of how position errors propagate from one vehicle to another in a cascaded system. In the formation flight system considered here, each i(sup th) aircraft uses information from itself and the preceding ((i-1)(sup th)) aircraft to track a commanded relative position. A possible solution for meeting performance requirements with such a system is to allow string instability. This paper explores two results of string instability and outlines analysis techniques for string unstable systems. The three analysis techniques presented here are: linear, nonlinear formation performance, and ride quality. The linear technique was developed from a worst-case scenario and could be applied to the design of a string unstable controller. The nonlinear formation performance and ride quality analysis techniques both use nonlinear formation simulation. Three of the four formation-controller gain-sets analyzed in this paper were limited more by ride quality than by performance. Formations of up to seven aircraft in a cascaded formation could be used in the presence of light gusts with this string unstable system.

  14. Self-force on an electric dipole in the spacetime of a cosmic string

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Muniz, C.R., E-mail: celiomuniz@yahoo.com; Bezerra, V.B., E-mail: valdir@ufpb.br

    2014-01-15

    We calculate the electrostatic self-force on an electric dipole in the spacetime generated by a static, thin, infinite and straight cosmic string. The electric dipole is held fixed in different configurations, namely, parallel, perpendicular to the cosmic string and oriented along the azimuthal direction around this topological defect, which is stretched along the z axis. We show that the self-force is equivalent to an interaction of the electric dipole with an effective dipole moment which depends on the linear mass density of the cosmic string and on the configuration. The plots of the self-forces as functions of the parameter whichmore » determines the angular deficit of the cosmic string are shown for those different configurations. -- Highlights: •Review of regularized Green’s function applied to the problem. •Self-force on an electric dipole in the string spacetime for some orientations. •Representation via graphs of the self-forces versus angular parameter of the cosmic string. •Self-force induced by the string seen as an interaction between two dipoles. •Discussion about the superposition principle in this non-trivial background.« less

  15. Chern-Simons improved Hamiltonians for strings in three space dimensions

    NASA Astrophysics Data System (ADS)

    Gordeli, Ivan; Melnikov, Dmitry; Niemi, Antti J.; Sedrakyan, Ara

    2016-07-01

    In the case of a structureless string the extrinsic curvature and torsion determine uniquely its shape in three-dimensional ambient space, by way of solution of the Frenet equation. In many physical scenarios there are in addition symmetries that constrain the functional form of the ensuing energy function. For example, the energy of a structureless string should be independent of the way the string is framed in the Frenet equation. Thus the energy should only involve the curvature and torsion as dynamical variables, in a manner that resembles the Hamiltonian of the Abelian Higgs model. Here we investigate the effect of symmetry principles in the construction of Hamiltonians for structureless strings. We deduce from the concept of frame independence that in addition to extrinsic curvature and torsion, the string can also engage a three-dimensional Abelian bulk gauge field as a dynamical variable. We find that the presence of a bulk gauge field gives rise to a long-range interaction between different strings. Moreover, when this gauge field is subject to Chern-Simons self-interaction, it becomes plausible that interacting strings are subject to fractional statistics in three space dimensions.

  16. Survival of pq -superstrings in field theory simulations

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Lizarraga, Joanes; Urrestilla, Jon, E-mail: joanes.lizarraga@ehu.eus, E-mail: jon.urrestilla@ehu.eus

    2016-04-01

    We perform large-scale field theoretical simulations in expanding universe to characterize a network of strings that can form composed bound states. The network consists of two copies of Abelian Higgs strings (which we label p and q , respectively) coupled via a potential term to give pq bound states. The simulations are performed using two different kinds of initial conditions: the first one with a network of p - and q -strings, and the second one with a network of q - and pq -strings. This way, we start from two opposite situations: one with no initial pq -strings, andmore » one with a large initial number of pq -strings. We find that in both cases the system scales, and in both cases the system prefers to have a low fraction of pq -strings. This is somewhat surprising in the case for the second type of conditions, showing that the unzipping mechanism is very efficient. We also find hints that both initial conditions tend to asymptote to a common configuration, though we would need a larger dynamical range to confirm it. The average velocities of the different types of strings in the network have also been explored for the first time.« less

  17. The reproductive output of sea lice Caligus rogercresseyi under controlled conditions.

    PubMed

    Bravo, Sandra

    2010-05-01

    Gravid females of Caligus rogercresseyi were collected from Atlantic salmon (Salmo salar) from a farm located at Chiloe Island (42 degrees 40'S73 degrees 15'W), Chile, to obtain information about the reproductive output of this parasite in vitro. The egg strings removed from the females were incubated under controlled conditions to obtain virgin adult females. One female which had mated only once produced eleven generations of eggs strings in a period of 74 days. The first egg strings of the females obtained in vitro were produced at 389 degree days ( degrees D) after egg incubation, while the next generations of eggs strings were produced with a periodicity between 4 and 6 days dependent on the water temperature. The average length of the egg string was 3.1mm and the mean number of eggs per string was 31. The values recorded in captivity for the egg string length and the number of eggs per string, were lower than the values recorded in gravid females from the field. One female survived for 79 days and males, maintained separately from the females, survived for 60 days. Copyright 2009 Elsevier Inc. All rights reserved.

  18. Searching for cosmic strings in CMB anisotropy maps using wavelets and curvelets

    NASA Astrophysics Data System (ADS)

    Hergt, Lukas; Amara, Adam; Brandenberger, Robert; Kacprzak, Tomasz; Réfrégier, Alexandre

    2017-06-01

    We use wavelet and curvelet transforms to extract signals of cosmic strings from simulated cosmic microwave background (CMB) temperature anisotropy maps, and to study the limits on the cosmic string tension which various ongoing CMB temperature anisotropy experiments will be able to achieve. We construct sky maps with size and angular resolution corresponding to various experiments. These maps contain the signals of a scaling solution of long string segments with a given string tension G μ, the contribution of the dominant Gaussian primordial cosmological fluctuations, and pixel by pixel white noise with an amplitude corresponding to the instrumental noise of the various experiments. In the case that we include white noise, we find that using curvelets we obtain lower bounds on the string tension than with wavelets. For maps with Planck specification, we obtain bounds comparable to what was obtained by the Planck collaboration [1]. Experiments with better angular resolution such as the South Pole Telescope third generation (SPT-3G) survey will be able to yield stronger limits. For maps with a specification of SPT-3G we find that string signals will be visible down to a string tension of G μ = 1.4 × 10-7.

  19. Segmented strings in AdS 3

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Callebaut, Nele; Gubser, Steven S.; Samberg, Andreas

    We study segmented strings in flat space and in AdS 3. In flat space, these well known classical motions describe strings which at any instant of time are piecewise linear. In AdS 3, the worldsheet is composed of faces each of which is a region bounded by null geodesics in an AdS 2 subspace of AdS 3. The time evolution can be described by specifying the null geodesic motion of kinks in the string at which two segments are joined. The outcome of collisions of kinks on the worldsheet can be worked out essentially using considerations of causality. We studymore » several examples of closed segmented strings in AdS 3 and find an unexpected quasi-periodic behavior. Here, we also work out a WKB analysis of quantum states of yo-yo strings in AdS 5 and find a logarithmic term reminiscent of the logarithmic twist of string states on the leading Regge trajectory.« less

  20. Segmented strings in AdS 3

    DOE PAGES

    Callebaut, Nele; Gubser, Steven S.; Samberg, Andreas; ...

    2015-11-17

    We study segmented strings in flat space and in AdS 3. In flat space, these well known classical motions describe strings which at any instant of time are piecewise linear. In AdS 3, the worldsheet is composed of faces each of which is a region bounded by null geodesics in an AdS 2 subspace of AdS 3. The time evolution can be described by specifying the null geodesic motion of kinks in the string at which two segments are joined. The outcome of collisions of kinks on the worldsheet can be worked out essentially using considerations of causality. We studymore » several examples of closed segmented strings in AdS 3 and find an unexpected quasi-periodic behavior. Here, we also work out a WKB analysis of quantum states of yo-yo strings in AdS 5 and find a logarithmic term reminiscent of the logarithmic twist of string states on the leading Regge trajectory.« less

  1. Self-gravitating strings in 2+1 dimensions

    NASA Astrophysics Data System (ADS)

    Ben-Menahem, Shahar

    1993-05-01

    We present a family of classical spacetimes in 2+1 dimensions. Such a spacetime is produced by a Nambu-Goto self-gravitating string. Because of the special properties of three-dimensional gravity, the metric is completely described as a Minkowski space with two identified world sheets. In the flat limit, the standard string is recovered. The formalism is developed for an open string with massive end points, but applies to other boundary conditions as well. We consider another limit, where the string tension vanishes in geometrical units but the end masses produce finite deficit angles. In this limit, our open string reduces to the free-masses solution of Gott, which possesses closed timelike curves when the relative motion of the two masses is sufficiently rapid. It is shown that the induced world sheet Liouville mode obeys (-classically)- a sinh- or cosh-Gordon differential equation, which reduces to the Liouville equation in the flat limit. A quadratic-action formulation of this system is presented. The possibility and significance of quantizing the self-gravitating string is discussed.

  2. Elliptic genus of E-strings

    NASA Astrophysics Data System (ADS)

    Kim, Joonho; Kim, Seok; Lee, Kimyeong; Park, Jaemo; Vafa, Cumrun

    2017-09-01

    We study a family of 2d N=(0, 4) gauge theories which describes at low energy the dynamics of E-strings, the M2-branes suspended between a pair of M5 and M9 branes. The gauge theory is engineered using a duality with type IIA theory, leading to the D2-branes suspended between an NS5-brane and 8 D8-branes on an O8-plane. We compute the elliptic genus of this family of theories, and find agreement with the known results for single and two E-strings. The partition function can in principle be computed for arbitrary number of E-strings, and we compute them explicitly for low numbers. We test our predictions against the partially known results from topological strings, as well as from the instanton calculus of 5d Sp(1) gauge theory. Given the relation to topological strings, our computation provides the all genus partition function of the refined topological strings on the canonical bundle over 1/2K3.

  3. Experimental observation of Bethe strings

    NASA Astrophysics Data System (ADS)

    Wang, Zhe; Wu, Jianda; Yang, Wang; Bera, Anup Kumar; Kamenskyi, Dmytro; Islam, A. T. M. Nazmul; Xu, Shenglong; Law, Joseph Matthew; Lake, Bella; Wu, Congjun; Loidl, Alois

    2018-02-01

    Almost a century ago, string states—complex bound states of magnetic excitations—were predicted to exist in one-dimensional quantum magnets. However, despite many theoretical studies, the experimental realization and identification of string states in a condensed-matter system have yet to be achieved. Here we use high-resolution terahertz spectroscopy to resolve string states in the antiferromagnetic Heisenberg-Ising chain SrCo2V2O8 in strong longitudinal magnetic fields. In the field-induced quantum-critical regime, we identify strings and fractional magnetic excitations that are accurately described by the Bethe ansatz. Close to quantum criticality, the string excitations govern the quantum spin dynamics, whereas the fractional excitations, which are dominant at low energies, reflect the antiferromagnetic quantum fluctuations. Today, Bethe’s result is important not only in the field of quantum magnetism but also more broadly, including in the study of cold atoms and in string theory; hence, we anticipate that our work will shed light on the study of complex many-body systems in general.

  4. The acoustics of the bagana

    NASA Astrophysics Data System (ADS)

    Weisser, Stephanie; Demolin, Didier

    2002-11-01

    The bagana is a big Ethiopian lyre with ten strings. The instrument is found in the area of the Amhara, the culturally dominant tribe of Ethiopia. It is an intimate instrument, played only with the voice, for prayer and meditation. It can be tuned in two pentatonic scales, and can be plucked with the fingers, string by string or with a plectrum, all strings together. The box of the bagana is made of wood covered with leather, and the strings are made of gut. They are very thick and between them and the bridge, there are ten buzzers. Therefore, the bagana produces a very deep and buzzing sound. The paper will analyze the acoustical function of several parts of this instrument, e.g., the sound box and its crosslike hole at the back. The influence of the buzzers on the fundamental frequency, the spectrum, the duration, and the intensity of the signal will be closely examined. The role of the five unused strings (the ''rest'' strings) will also be examined.

  5. Mechanical Properties of Nylon Harp Strings

    PubMed Central

    Lynch-Aird, Nicolas; Woodhouse, Jim

    2017-01-01

    Monofilament nylon strings with a range of diameters, commercially marketed as harp strings, have been tested to establish their long-term mechanical properties. Once a string had settled into a desired stress state, the Young’s modulus was measured by a variety of methods that probe different time-scales. The modulus was found to be a strong function of testing frequency and also a strong function of stress. Strings were also subjected to cyclical variations of temperature, allowing various thermal properties to be measured: the coefficient of linear thermal expansion and the thermal sensitivities of tuning, Young’s modulus and density. The results revealed that the particular strings tested are divided into two groups with very different properties: stress-strain behaviour differing by a factor of two and some parametric sensitivities even having the opposite sign. Within each group, correlation studies allowed simple functional fits to be found to the key properties, which have the potential to be used in automated tuning systems for harp strings. PMID:28772858

  6. Hydroball string sensing system

    DOEpatents

    Hurwitz, Michael J.; Ekeroth, Douglas E.; Squarer, David

    1991-01-01

    A hydroball string sensing system for a nuclear reactor that includes stainless tubes positioned to guide hydroball strings into and out of the nuclear reactor core. A sensor such as an ultrasonic transducer transmitter and receiver is positioned outside of the nuclear reactor core and adjacent to the tube. The presence of an object such a bullet member positioned at an end a hydroball string, or any one of the hydroballs interrupts the transmission of ultrasound from the transmitter to the receiver. Alternatively, if the bullet member and hydroballs include a ferritic material, either a Hall effect sensor or other magnetic field sensors such as a magnetic field rate of change sensor can be used to detect the location and position of a hydroball string. Placing two sensors along the tube with a known distance between the sensors enables the velocity of a hydroball string to be determined. This determined velocity can be used to control the flow rate of a fluid within the tube so as to control the velocity of the hydroball string.

  7. Open string with a background B field as the first order mechanics, noncommutativity, and soldering formalism

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Deriglazov, A. A.; Neves, C.; Oliveira, W.

    2007-09-15

    To study noncommutativity properties of the open string with constant B field, we construct a mechanical action that reproduces classical dynamics of the string sector under consideration. It allows one to apply the Dirac quantization procedure for constrained systems in a direct and unambiguous way. The mechanical action turns out to be the first order system without taking the strong field limit B{yields}{infinity}. In particular, it is true for the zero mode of the string coordinate, which means that the noncommutativity is an intrinsic property of this mechanical system. We describe the arbitrariness in the relation existing between the mechanicalmore » and the string variables and show that noncommutativity of the string variables on the boundary can be removed. This is in correspondence with the result of Seiberg and Witten on the relation among noncommutative and ordinary Yang-Mills theories. The recently developed soldering formalism helps us to establish a connection between the original open string action and the Polyakov action.« less

  8. Mechanical Properties of Nylon Harp Strings.

    PubMed

    Lynch-Aird, Nicolas; Woodhouse, Jim

    2017-05-04

    Monofilament nylon strings with a range of diameters, commercially marketed as harp strings, have been tested to establish their long-term mechanical properties. Once a string had settled into a desired stress state, the Young's modulus was measured by a variety of methods that probe different time-scales. The modulus was found to be a strong function of testing frequency and also a strong function of stress. Strings were also subjected to cyclical variations of temperature, allowing various thermal properties to be measured: the coefficient of linear thermal expansion and the thermal sensitivities of tuning, Young's modulus and density. The results revealed that the particular strings tested are divided into two groups with very different properties: stress-strain behaviour differing by a factor of two and some parametric sensitivities even having the opposite sign. Within each group, correlation studies allowed simple functional fits to be found to the key properties, which have the potential to be used in automated tuning systems for harp strings.

  9. Further Development of HS Field Theory

    NASA Astrophysics Data System (ADS)

    Abdurrahman, Abdulmajeed; Faridani, Jacqueline; Gassem, Mahmoud

    2006-04-01

    We present a systematic treatment of the HS Field theory of the open bosonic string and discuss its relationship to other full string field theories of the open bosonic string such as Witten's theory and the CVS theory. In the development of the HS field theory we encounter infinite dimensional matrices arising from the change of representation between the two theories, i.e., the HS field theory and the full string field theory. We give a general procedure of how to invert these gigantic matrices. The inversion of these matrices involves the computation of many infinite sums. We give the values of these sums and state their generalizations arising from considering higher order vertices (i.e., more than three strings) in string field theory. Moreover, we give a general procedure, on how to evaluate the generalized sums, that can be extended to many generic sums of similar properties. We also discuss the conformal operator connecting the HS field theory to that of the CVS string field theory.

  10. Plucked String on a Shoestring Budget

    NASA Astrophysics Data System (ADS)

    Gluck, Paul

    2009-01-01

    The physics of the plucked string has been treated in many articles and books.1-4 For our 12th-grade high school physics laboratory, we have built a cheap, simple sonometer apparatus for each pair of students on which they may investigate some interesting phenomena that arise when a string is plucked. Among these are the generation of harmonics (overtones) and the way their number depends on the length of a string and on where one plucks, the relation between the frequencies of the fundamental and those of the harmonics, and the way these are affected by changes in the length and the tension in the string. Such an experiment will help students appreciate the working of stringed musical instruments and, in particular, the contribution of overtones to the richness of sound produced.

  11. A Search for Cosmic String Loops Using GADGET-2 Cosmological N-Body Simulator

    NASA Astrophysics Data System (ADS)

    Braverman, William; Cousins, Bryce; Jia, Hewei

    2018-01-01

    Cosmic string loops are an extremely elusive hypothetical entity that have eluded the grasp of physicists and astronomers since their existence was postulated in the 1970’s. Finding evidence of their existence could be the first empirical evidence of string theory.Simulating their basic motion in a cold dark matter background using GADGET-2 allows us to predict where they may cluster during large scale structure formation (if they cluster at all). Here, we present our progress in placing cosmic strings into GADGET-2 with their basic equations of motion to lay a ground work for more complex simulations to find where these strings cluster. Ultimately, these simulations could lay a groundwork as to where future microlensing and gravitational wave observatories should look for cosmic strings.

  12. CMB temperature trispectrum of cosmic strings

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Hindmarsh, Mark; Ringeval, Christophe; Suyama, Teruaki

    2010-03-15

    We provide an analytical expression for the trispectrum of the cosmic microwave background (CMB) temperature anisotropies induced by cosmic strings. Our result is derived for the small angular scales under the assumption that the temperature anisotropy is induced by the Gott-Kaiser-Stebbins effect. The trispectrum is predicted to decay with a noninteger power-law exponent l{sup -{rho}}with 6<{rho}<7, depending on the string microstructure, and thus on the string model. For Nambu-Goto strings, this exponent is related to the string mean square velocity and the loop distribution function. We then explore two classes of wave number configuration in Fourier space, the kite andmore » trapezium quadrilaterals. The trispectrum can be of any sign and appears to be strongly enhanced for all squeezed quadrilaterals.« less

  13. Semianalytic calculation of cosmic microwave background anisotropies from wiggly and superconducting cosmic strings

    NASA Astrophysics Data System (ADS)

    Rybak, I. Yu.; Avgoustidis, A.; Martins, C. J. A. P.

    2017-11-01

    We study how the presence of world-sheet currents affects the evolution of cosmic string networks, and their impact on predictions for the cosmic microwave background (CMB) anisotropies generated by these networks. We provide a general description of string networks with currents and explicitly investigate in detail two physically motivated examples: wiggly and superconducting cosmic string networks. By using a modified version of the CMBact code, we show quantitatively how the relevant network parameters in both of these cases influence the predicted CMB signal. Our analysis suggests that previous studies have overestimated the amplitude of the anisotropies for wiggly strings. For superconducting strings the amplitude of the anisotropies depends on parameters which presently are not well known—but which can be measured in future high-resolution numerical simulations.

  14. Exploring the spectrum of regularized bosonic string theory

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Ambjørn, J., E-mail: ambjorn@nbi.dk; Makeenko, Y., E-mail: makeenko@nbi.dk

    2015-03-15

    We implement a UV regularization of the bosonic string by truncating its mode expansion and keeping the regularized theory “as diffeomorphism invariant as possible.” We compute the regularized determinant of the 2d Laplacian for the closed string winding around a compact dimension, obtaining the effective action in this way. The minimization of the effective action reliably determines the energy of the string ground state for a long string and/or for a large number of space-time dimensions. We discuss the possibility of a scaling limit when the cutoff is taken to infinity.

  15. Evolution of cosmic string networks

    NASA Technical Reports Server (NTRS)

    Albrecht, Andreas; Turok, Neil

    1989-01-01

    Results on cosmic strings are summarized including: (1) the application of non-equilibrium statistical mechanics to cosmic string evolution; (2) a simple one scale model for the long strings which has a great deal of predictive power; (3) results from large scale numerical simulations; and (4) a discussion of the observational consequences of our results. An upper bound on G mu of approximately 10(-7) emerges from the millisecond pulsar gravity wave bound. How numerical uncertainties affect this are discussed. Any changes which weaken the bound would probably also give the long strings the dominant role in producing observational consequences.

  16. Perturbations from cosmic strings in cold dark matter

    NASA Technical Reports Server (NTRS)

    Albrecht, Andreas; Stebbins, Albert

    1992-01-01

    A systematic linear analysis of the perturbations induced by cosmic strings in cold dark matter is presented. The power spectrum is calculated and it is found that the strings produce a great deal of power on small scales. It is shown that the perturbations on interesting scales are the result of many uncorrelated string motions, which indicates a much more Gaussian distribution than was previously supposed.

  17. Mean transverse momenta correlations in hadron-hadron collisions in MC toy model with repulsing strings

    DOE Office of Scientific and Technical Information (OSTI.GOV)

    Altsybeev, Igor

    2016-01-22

    In the present work, Monte-Carlo toy model with repulsing quark-gluon strings in hadron-hadron collisions is described. String repulsion creates transverse boosts for the string decay products, giving modifications of observables. As an example, long-range correlations between mean transverse momenta of particles in two observation windows are studied in MC toy simulation of the heavy-ion collisions.

  18. Perturbations from cosmic strings in cold dark matter

    NASA Technical Reports Server (NTRS)

    Albrecht, Andreas; Stebbins, Albert

    1991-01-01

    A systematic linear analysis of the perturbations induced by cosmic strings in cold dark matter is presented. The power spectrum is calculated and it is found that the strings produce a great deal of power on small scales. It is shown that the perturbations on interesting scales are the result of many uncorrelated string motions, which indicates a much more Gaussian distribution than was previously supposed.

  19. Relativistic strings - From soap films to a grand unified theory

    NASA Astrophysics Data System (ADS)

    Nesterenko, V. V.

    1986-11-01

    The concept of relativistic strings is considered in connection with the theory of minimal surfaces (e.g., soap films stretched onto closed wire contours). The role of relativistic strings in hadron physics is discussed. Attention is then given to the creation of a grand unified theory on the basis of the superstring concept. Finally, the role of relativistic strings in cosmology is examined.

  20. D1 string dynamics in curved backgrounds with fluxes

    NASA Astrophysics Data System (ADS)

    Banerjee, Aritra; Biswas, Sagar; Nayak, Rashmi R.

    2016-04-01

    We study various rotating and oscillating D-string configurations in some general backgrounds with fluxes. In particular, we look for solutions to the equations of motion of various rigidly rotating D-strings in AdS3 background with mixed flux, and in the intersecting D-brane geometries. We find out relations among various conserved charges corresponding to the breathing and rotating D-string configurations.

  1. Experimenting with string musical instruments

    NASA Astrophysics Data System (ADS)

    LoPresto, Michael C.

    2012-03-01

    What follows are several investigations involving string musical instruments developed for and used in a Science of Sound & Light course. The experiments make use of a guitar, orchestral string instruments and data collection and graphing software. They are designed to provide students with concrete examples of how mathematical formulae, when used in physics, represent reality that can actually be observed, in this case, the operation of string musical instruments.

  2. A Concept for Continuous Monitoring that Reduces Redundancy in Information Assurance Processes

    DTIC Science & Technology

    2011-09-01

    System.out.println(“Driver loaded”); String url=“jdbc:postgresql://localhost/IAcontrols”; String user = “ postgres ”; String pwd... postgres ”; Connection DB_mobile_conn = DriverManager.getConnection(url,user,pwd); System.out.println(“Database Connect ok...user = “ postgres ”; String pwd = “ postgres ”; Connection DB_mobile_conn = DriverManager.getConnection(url,user,pwd); System.out.println

  3. Energy and Momentum Transport in String Waves

    ERIC Educational Resources Information Center

    Juenker, D. W.

    1976-01-01

    Formulas are derived for the energy, momentum, and angular momentum transmitted by waves of arbitrary shape in an inextensible string by pure transverse waves in a string using Tait's procedure. (Author/CP)

  4. Separation of traveling and standing waves in a finite dispersive string with partial or continuous viscoelastic foundation

    NASA Astrophysics Data System (ADS)

    Cheng, Xiangle; Blanchard, Antoine; Tan, Chin An; Lu, Huancai; Bergman, Lawrence A.; McFarland, D. Michael; Vakakis, Alexander F.

    2017-12-01

    The free and forced vibrations of a linear string with a local spring-damper on a partial elastic foundation, as well as a linear string on a viscoelastic foundation conceptualized as a continuous distribution of springs and dampers, are studied in this paper. Exact, analytical results are obtained for the free and forced response to a harmonic excitation applied at one end of the string. Relations between mode complexity and energy confinement with the dispersion in the string system are examined for the steady-state forced vibration, and numerical methods are applied to simulate the transient evolution of energy propagation. Eigenvalue loci veering and normal mode localization are observed for weakly coupled subsystems, when the foundation stiffness is sufficiently large, for both the spatially symmetric and asymmetric systems. The forced vibration results show that nonproportional damping-induced mode complexity, for which there are co-existing regions of purely traveling waves and standing waves, is attainable for the dispersive string system. However, this wave transition phenomenon depends strongly on the location of the attached discrete spring-damper relative to the foundation and whether the excitation frequency Ω is above or below the cutoff frequency ωc. When Ω<ωc, the wave transition cannot be attained for a string on an elastic foundation, but is possible if the string is on a viscoelastic foundation. Although this study is primarily formulated for a harmonic boundary excitation at one end of the string, generalization of the mode complexity can be deduced for the steady-state forced response of the string-foundation system to synchronous end excitations and is confirmed numerically. This work represents a novel study to understand the wave transitions in a dispersive structural system and lays the groundwork for potentially effective passive vibration control strategies.

  5. Constraining the String Gauge Field by Galaxy Rotation Curves and Perihelion Precession of Planets

    NASA Astrophysics Data System (ADS)

    Cheung, Yeuk-Kwan E.; Xu, Feng

    2013-09-01

    We discuss a cosmological model in which the string gauge field coupled universally to matter gives rise to an extra centripetal force and will have observable signatures on cosmological and astronomical observations. Several tests are performed using data including galaxy rotation curves of 22 spiral galaxies of varied luminosities and sizes and perihelion precessions of planets in the solar system. The rotation curves of the same group of galaxies are independently fit using a dark matter model with the generalized Navarro-Frenk-White (NFW) profile and the string model. A remarkable fit of galaxy rotation curves is achieved using the one-parameter string model as compared to the three-parameter dark matter model with the NFW profile. The average χ2 value of the NFW fit is 9% better than that of the string model at a price of two more free parameters. Furthermore, from the string model, we can give a dynamical explanation for the phenomenological Tully-Fisher relation. We are able to derive a relation between field strength, galaxy size, and luminosity, which can be verified with data from the 22 galaxies. To further test the hypothesis of the universal existence of the string gauge field, we apply our string model to the solar system. Constraint on the magnitude of the string field in the solar system is deduced from the current ranges for any anomalous perihelion precession of planets allowed by the latest observations. The field distribution resembles a dipole field originating from the Sun. The string field strength deduced from the solar system observations is of a similar magnitude as the field strength needed to sustain the rotational speed of the Sun inside the Milky Way. This hypothesis can be tested further by future observations with higher precision.

  6. Incorporation of Predictive Population Modeling into the AOP Famework: A Case Study with White Suckers Exposed to Pulp Effluent

    EPA Science Inventory

    A need in ecological risk assessment is the ability to create linkages between chemically-induced alterations at molecular and biochemical levels of organization with adverse outcomes in whole organisms and populations. A predictive model was developed to translate changes in th...

  7. 75 FR 77961 - Endangered and Threatened Wildlife and Plants; Revised Critical Habitat for Santa Ana Sucker

    Federal Register 2010, 2011, 2012, 2013, 2014

    2010-12-14

    ... integrated water system that contains and provides the appropriate quantity of coarse substrates such as... reduces water temperature during summer and fall months. Therefore, a complex and integrated stream system... water velocities to support successful spawning. Swift (2001, p. 26) considered that only the Rialto...

  8. Decline of quaking aspen in the Interior West - examples from Utah

    Treesearch

    Dale L. Bartos; Robert B. Campbell

    1998-01-01

    Quaking aspen (Populus tremuloides) are unique because, in contrast to most western forest trees, they reproduce primarily by suckering from the parent root system. Generally disturbance or dieback is necessary to stimulate regeneration of aspen stands. These self-regenerating stands have existed for thousands of years. If they are lost from the...

  9. Repurposing an Old Game for an International World

    ERIC Educational Resources Information Center

    Hofstede, Gert Jan; Murff, Elizabeth J. Tipton

    2012-01-01

    The game SO LONG SUCKER was designed in the United States in 1964 with the aim of showing how potentially unethical behavior necessary for winning was inherent in the game's incentive structure. Sessions with East Asian participants, however, led to very different game dynamics in which collaborative rather than antagonistic behaviors occurred.…

  10. 78 FR 22556 - Endangered and Threatened Wildlife and Plants; Revised Recovery Plan for Lost River Sucker and...

    Federal Register 2010, 2011, 2012, 2013, 2014

    2013-04-16

    ... very similar in ecology. They both predominantly inhabit lake environments but also periodically... from entrainment in water management structures, were cited as causes for declines in populations prior... survival is also low in these populations. The last time a substantial group of juveniles joined the adult...

  11. Food habits of Bald Eagles breeding in the Arizona desert

    Treesearch

    Teryl G. Grubb

    1995-01-01

    Of 1814 foraging attempts, prey captures, or nest deliveries by Bald Eagles (Haliaeetus leucocephalus) in 14 Arizona breeding areas during 1983-1985, 1471 observations were identifiable to at least class: fish (76%), mammal (18%), bird (4%), and reptile/amphibian (2%). Forty-five species were recorded: catfish (Ictalurus punctatus, Pylodictis olivaris), suckers (...

  12. Progress estimating incidence rates of tumors and deformities in St. Louis River white sucker

    EPA Science Inventory

    The St. Louis River Area of Concern (AOC) was listed for the Beneficial Use Impairment (BUI) of Fish Tumors and Other Deformities without the benefit of histological information. Information on the fish tumor incidence rate is important for the future removal of the BUI. Two year...

  13. English Revisited: Sucker Punch, Moffatt, Avatars and Forensic Evidence

    ERIC Educational Resources Information Center

    Russell, Jennifer; Beavis, Catherine

    2012-01-01

    Contemporary curriculum guidelines, such as those provided in the incoming Australian curriculum, call for English to attend to multimodal forms of text and literacy as well as more traditional forms. Students are expected to become capable and critical readers, users and creators of texts and forms of literacy that span everything from newspapers…

  14. Communicating as an American Immigrant in Israel: The "Freier" Phenomenon and the Pursuit of an Alternative Value System.

    ERIC Educational Resources Information Center

    Bloch, Linda-Renee

    1998-01-01

    Examines a cultural phenomenon of "freier," or "sucker," in Israeli society, reflecting almost diametrically opposed values of American immigrants and native Israelis. In 47 interviews with American immigrants concerning communication interactions, insight was gained into the status and posture of the immigrants in the host…

  15. Rootstock performance in the 2009 NC-140 peach trial across 11 states

    USDA-ARS?s Scientific Manuscript database

    From 14 to 18 Prunus rootstocks budded with ‘Redhaven’ peach were planted at 16 locations in North America in 2009. Seven-year performance from 12 remaining locations in 11 states showed that significant differences among rootstocks and sites were found for survival, root suckers, growth, bloom dat...

  16. Bracing for Disaster

    ERIC Educational Resources Information Center

    Schaffhauser, Dian

    2011-01-01

    A freak blizzard, a mentally ill and armed student, a record-breaking flood. No matter how idyllic a campus may feel, no matter how cocooned the ivory tower, disaster can strike. If a campus is unprepared, it comes like a sucker punch, potentially turning a crisis into a tragedy of unimagined proportions--and causing reverberations that will be…

  17. Acoustic data transmission through a drill string

    DOEpatents

    Drumheller, D.S.

    1988-04-21

    Acoustical signals are transmitted through a drill string by canceling upward moving acoustical noise and by preconditioning the data in recognition of the comb filter impedance characteristics of the drill string. 5 figs.

  18. Computer Center: BASIC String Models of Genetic Information Transfer.

    ERIC Educational Resources Information Center

    Spain, James D., Ed.

    1984-01-01

    Discusses some of the major genetic information processes which may be modeled by computer program string manipulation, focusing on replication and transcription. Also discusses instructional applications of using string models. (JN)

  19. Effects of ordinary and superconducting cosmic strings on primordial nucleosynthesis

    NASA Technical Reports Server (NTRS)

    Hodges, Hardy M.; Turner, Michael S.

    1988-01-01

    A precise calculation is done of the primordial nucleosynthesis constraint on the energy per length of ordinary and superconducting cosmic strings. A general formula is provided for the constraint on the string tension for ordinary strings. Using the current values for the various parameters that describe the evolution of loops, the constraint for ordinary strings is G mu less than 2.2 x 10 to the minus 5 power. Our constraint is weaker than previously quoted limits by a factor of approximately 5. For superconducting loops, with currents generated by primordial magnetic fields, the constraint can be less or more stringent than this limit, depending on the strength of the magnetic field. It is also found in this case that there is a negligible amount of entropy production if the electromagnetic radiation from strings thermalizes with the radiation background.

  20. Evolution and End Point of the Black String Instability: Large D Solution.

    PubMed

    Emparan, Roberto; Suzuki, Ryotaku; Tanabe, Kentaro

    2015-08-28

    We derive a simple set of nonlinear, (1+1)-dimensional partial differential equations that describe the dynamical evolution of black strings and branes to leading order in the expansion in the inverse of the number of dimensions D. These equations are easily solved numerically. Their solution shows that thin enough black strings are unstable to developing inhomogeneities along their length, and at late times they asymptote to stable nonuniform black strings. This proves an earlier conjecture about the end point of the instability of black strings in a large enough number of dimensions. If the initial black string is very thin, the final configuration is highly nonuniform and resembles a periodic array of localized black holes joined by short necks. We also present the equations that describe the nonlinear dynamics of anti-de Sitter black branes at large D.

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